Revision and phylogenetic analysis of the spider genus Philisca Simon (Araneae: Anyphaenidae, Amaurobioidinae) 3443
Author
Soto, Eduardo M.
Author
Ramírez, Martín J.
text
Zootaxa
2012
2012-08-29
3443
1
1
65
https://biotaxa.org/Zootaxa/article/view/zootaxa.3443.1.1
journal article
10.11646/zootaxa.3443.1.1
11755334
5254614
Philisca tripunctata
(Nicolet)
Figs 1a
,
3
,
34b
,
38b
Clubiona tripunctata
Nicolet, 1849: 138
(
2 female
syntypes
from
Chile
, no specific locality, in
MHNP 4219
, not reexamined).
Liparotoma pardalis
Mello-Leitão, 1943: 405
(immature
holotype
from
Chile
,
Santiago
, J. Carbalho coll., in MNRJ, not reexamined). Synonymized by
Ramírez, 1993
.
Liparotoma tripunctatum
:
Ramírez, 1993: 198
;
2003: 189–191
.
Philisca tripunctata
:
Ramírez, 2003: 189
.
Diagnosis:
Distinguished from other
Philisca
by having only one tooth on the retromargin of the chelicerae and by the dark, rectangular ventral patch on the opistosoma (
Fig. 1a
) (
Ramírez 1993
: fig. 16). Male chelicerae not modified (
Fig. 34b
).
FIGURE 2.
Philisca hahni
Simon
(EMS-00122, EMS-00139, EMS-00123).
a–c
female (
a
dorsal habitus;
b
epigyne, ventral view;
c
cleared epigyne, dorsal view);
d–e
male left palp (
d
ventral view;
e
retrolateral view). (AB: accessory bulbs, CD: copulatory ducts, MF: median field, S: spermathecae).
FIGURE 3.
Philisca tripunctata
Nicolet
(MACN-Ar 27294 and MACN-Ar 26812).
a–c
female (
a
dorsal habitus;
b
epigyne, ventral view;
c
cleared epigyne, dorsal view);
d–e
male left palp (
d
ventral view;
e
retrolateral view). (AB: accessory bulbs, CD: copulatory ducts, MF: median field, S: spermathecae).
Description:
Male and female described in
Ramírez (1993
,
2003
).
Distribution:
Chile
, from Choapa province to Cabo de Hornos.
Argentina
, from
Neuquén
to
Tierra del Fuego
provinces (
Fig. 38b
).
Variability:
Spines in male and female:
IV
: femur p 0, r d1ap.
New records:
ARGENTINA
:
Neuquén
:
P.N. Nahuel Huapi, Península Quetrihué, bosque de arrayanes,
24.II.1996
, M. Ramírez,
1 female
(MACN-Ar 14869).
Río Negro
:
P.N. Nahuel Huapi, cerca de Puerto Alegre, Lago Frías,
S41.04394º
W71.79962º
(+-
20m
GPS), elev.
811m
(barómetro),
29.XII.2010
, M. Ramírez, V. Werenkraut, S. Aisen,
1 immature
(MACN-Ar 28031).
CHILE
:
Región IV (
Coquimbo
):
Choapa Prov.: Cuesta Cavilolén,
23.5km
(por aire) NW Los Vilos,
S31.76669º
W71.32727º
(GPS +-
300m
),
600m
elev. (barómetro), bosque relicto en quebrada, colecta general,
2.XI.2011
, M.J. Ramírez, A. Ojanguren, J. Pizarro et al. (MJR-loc-60),
1 female
(MACN-Ar 28297),
1 immature
(MACN-Ar 28288),
6 immatures
(MACN-Ar 28300).
Región V (
Valparaíso
):
Quillota Prov.: Cuesta El Melón,
17.7km
(por aire) S La Ligua,
S32.60880º
W71.23638º
(GPS +-
100m
),
578m
elev. (barómetro), bosque relicto en quebrada, colecta general,
3.XI.2011
, M.J. Ramírez, A. Ojanguren, J. Pizarro et al. (MJR-loc-63),
2 immatures
(MACN-Ar 28286). Reserva Parque El Boldo, Zapallar,
300m
NNE de ruta costera,
S35.54593º
W71.45179º
(GPS, trayecto +-
100m
), elev.
114m
(barómetro),
12.II.2011
, bosque en quebrada, M. Ramírez, E. Soto, J. Pizarro (MJR-loc-01),
1 female
(MACN-Ar 28030),
1 immature
(MACN-Ar 28029),
1 immature
(MACN-Ar 28028),
1 immature
(MACN-Ar 28027); Quebrada El Tigre,
2.5km
E Zapallar,
S32.55143º
W71.43278º
(GPS, trayecto +-
220m
), elev.
357m
(barómetro),
12.II.2011
, bosque de boldo y belloto en quebrada, M. Ramírez, E. Soto, J. Pizarro (MJR-loc-02),
1 immature
(MACN-Ar 28024),
1 immature
(MACN-Ar 28023); sendero a Quebrada El Tigre,
2.3km
E Zapallar,
S32.55368º
W71.43493º
(GPS, trayecto +-
80m
), elev.
348–357m
(barómetro),
12.II.2011
, bosque de boldo, M. Ramírez, E. Soto, J. Pizarro (MJR-loc-02),
1 immature
(MACN-Ar 28025),
1 immature
(MACN-Ar 28026). Marga Marga Prov.: Sector Granizo, P.N. La Campana,
5.2km
(por aire) ENE Olmué,
S32.98124º
W71.13016º
(GPS, +-
200m
),
456m
(barómetro),
4.XI.2011
, bosque higrófilo en quebrada, colecta general, M.J. Ramírez, A. Ojanguren, J. Pizarro et al. (MJR-loc-65),
1 female
(MACN-Ar 28363).
Región VI (Bernardo
O'Higgins
):
Curicó Prov.:
20km
E of Potrero Grande, El Relvo,
S35º11’9”
W70º56’14.9”
, fogging
Nothofagus dombeyi
,
Laurelia
,
Colletia hystrix
,
Podocarpus saligna
,
16.I.2004
, J. Barriga T.,
1 female
(MACN-Ar 27387);
15km
E of Potrero Grande, Puente Los Morongos,
S36º12.09’
W70º58.06’
, fogging
Nothofagus alpina
,
N. dombeyi
,
Laurelia
sp.
,
30.III.2004
, J. Barriga T. (MACN-Ar 27385).
Región VIII (
Bío-Bío
)
:
Arauco Prov.
: P.N. Nahuelbuta,
S37º47’
W73º00’
,
1200m
,
12.II.2005
, fogging
Araucaria araucana
, J. Barriga T.
,
1 male
(MACN-Ar 27379). Concepción Prov.: Cerro Caracol, Concepción,
200m
,
S36º51’
W73º02’
,
17.XI.1993
, N. Platnick, K. Catley, M. Ramírez, T. Allen,
1 female
,
1 male
and
1 immature
(MACN-Ar 14859).
Región IX (
Araucanía
):
Malleco Prov.: P.N. Nahuelbuta,
S37º47’
W73º00’
,
1200m
,
12.II.2005
, fogging
A. araucana
, J. Barriga
,
2 females
and
7 males
(MACN-Ar 27378); P.N. Nahuelbuta,
12.II.2005
, fogging
N. dombeyi
, J. Barriga T.
,
4 females
and
9 males
(MACN-Ar 27381),
1 female
(MACN-Ar 26813),
1 female
(MACN-Ar 26759),
1 female
(MACN-Ar 27294),
1 female
(MACN-Ar 28114),
2 males
(MACN-Ar 27382),
1 male
(MACN-Ar 26760),
1 male
(MACN-Ar 26812),
1 male
(MACN-Ar 28115); Malalcahuello Way Lonquimay, fogging,
1221m
, 29°C,
S38°26.388’
W71°30.613’
,
N. dombeyi
,
50m
, 6:55PM,
18.I.2007
, Arias et al. Berkeley,
1 female
(CASENT 9039009).
Región X (
Los Lagos
)
:
Valdivia Prov.: P. Oncol, Quitaqui Way, fogging,
519m
, 13ºC,
S39º42.071’
W73º18.547’
,
Saxegothaea conspicua
,
45m
, 3:32PM,
10.I.2007
, Arias et al. Berkeley,
1 female
(CASENT 9039113),
1 male
(CASENT 9039117),
1 female
and
9 immatures
(CASENT 9039106); Chaihuin Res. Costera Valdiviana, Las Garzas Way, fogging,
357m
, 17ºC, 3:27PM
13.I.2007
,
S39º59.669’
W73º34.721’
,
N. dombeyi
,
35m
, Arias et al. Berkeley,
2 females
and
3 immatures
(CASENT 9039085). Osorno Prov.: P.N. Puyehue, forest behind cabañas de la Fundación de las Raíces, between Rte 215 and Río Gol Gol, ca.
30km
E
Entre Lagos
,
S40º39'59"
W72º10'19"
, beating vegetation,
22–24.I.2003
, S. Lew 090,
2 females
(CASENT 9016666). Chiloé, Chinquen,
S42°37.092’
W74°06.073’
,
15m
, fogging 150cc/l
Nothofagus nitida
, 5:38PM,
1.III.2008
, Arias et al. UCB,
2 females
(CASENT 9034018),
1 female
(CASENT 9034019),
1 male
(CASENT 9034013); Chiloé, Chinquen,
S42°37.098’
W74°06.025’
,
31m
, fogging 150cc/l
N. nitida
, 6:50PM,
1.III.2008
, Arias et al. UCB,
1 female
and
1 immature
(CASENT 9034037).
Región XII (
Magallanes y la Antártica chilena
):
Tierra del Fuego Prov.: Isla Grande de Tierra del Fuego, Lago Blanco,
125km
SE Porvenir,
S54º03.012’
W68º57.472’
,
120m
elev, collecting under debris along shore of lake,
14.XII.1999
, C. Griswold CL134,
1 female
(CASENT 9035282).
Región XIV (de
los Ríos
):
Valdivia Prov.: Chaihuin Res. Costera Valdiviana, Las Garzas way, fogging
357m
17ºC,
S39º59.669’
W73º34.721’
,
N. dombeyi
35m
, 3:27PM,
13.I.2007
, Arias et al. Berkeley,
1 female
(CASENT 9039088); Res. Costera Valdiviana, Chaihuin, Las Garzas,
360m
,
S39°59.687’
W73°35.227
, 17°C, fogging 150cc/ l
N. nitida
, 7:30PM,
26.II.2008
, Arias et al. UCB,
1 male
,
1 female
and
1 immature
(CASENT 9039047),
1 male
(CASENT 9039048); Chaihuin, Reserva Costera Valdiviana, Las Garzas,
S39°59.640’
W73°35.197
, 360m, 20.9°C, fogging 150cc/l
N. nitida
11:00AM,
27.II.2008
, Arias et al. UCB,
1 female
(CASENT 9039033).
Material re-examined:
ARGENTINA
:
Neuquén
:
Lago Espejo
, rivera SE,
20.I.1985
,
M. Ramírez
,
7 females
and
1 male
(
MACN-Ar 14851
);
1 female
and
4 immatures
(
MACN-Ar 14865
);
P.N. Lanín
, Lago Tromen,
I.1991
,
V. Fiorani
,
1 male
(
MACN-Ar 14867
); Hua Hum,
I.1985
,
M. Ramírez
,
1 female
(
MACN-Ar 14852
); Guillén,
II.1968
,
E. Maury
and
Muller
,
1 female
and
1 immature
(
MACN-Ar 14831
)
.
Chubut
:
Río Menéndez
,
I.1990
, M.
Ramírez
,
1 female
(
MACN-Ar 15280
),
Lago Futalaufquen
,
Playa Blanca
,
I.1990
,
M. Ramírez
,
1 female
(MACN-
Ar
14868)
and
1 immature
(
MACN-Ar 14866
);
Río Arroyares
,
I.1990
, M.
Ramírez
,
1 female
(
MACN-Ar 14860
);
Lago Verde
,
I.1990
,
M. Ramírez
,
1 immature
(
MACN-Ar 14853
)
.
Río Negro
:
Bariloche
,
7.III.1942
,
Birabén
,
1 female
(
MACN-Ar 14864
)
.
Tierra del Fuego
:
Bahía Buen Suceso
,
16–31.I.1986
,
E. Maury
,
1 female
(
MACN-Ar 14856
); Isla de los Estados,
Bahía Crosby
,
18.X.1941
,
2 immatures
(
MACN-Ar 14854
); Lago Fagnano,
28.II.1957
,
Vellard
,
1 immature
(
MACN-Ar 14829
)
.
CHILE
:
Región
VIII (
Bío-Bío
)
:
Concepción Prov.
:
Hualpén
,
11.I.1989
, M.
Ramírez
,
1 female
(
MACN-Ar 14848
).
Región IX
(
Araucanía
)
:
Malleco Prov.
:
Fdo. Marta Ester
,
15 km
W of Victoria
,
14.I.1989
,
M. Ramírez
,
1 female
(
MACN-Ar 14834
),
1 male
(
MACN-Ar 14828
); Mon. Nat. Contulmo,
12.I.1989
,
M. Ramírez
,
2 females
(
MACN-Ar 14827
).
Región X
(
Los Lagos
)
:
Llanquihue Prov.
:
Caleta La Arena
,
30.I.1991
, M.
Ramírez
,
1 female
(
MACN-Ar 14832
)
.