<document id="4CB359AD449536BB157ACD16BE059BB4" ID-CLB-Dataset="24614" ID-DOI="10.5281/zenodo.3576287" ID-GBIF-Dataset="80959f8f-aa73-499e-93ec-5c07458c7935" ID-ISSN="0024-0672" ID-Zenodo-Dep="3576287" IM.materialsCitations_requiresApprovalFor="plazi" IM.metadata_requiresApprovalFor="plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1576401033600" checkinUser="plazi" docAuthor="Schilthuizen, Menno &amp; Perreau, Michel" docDate="2008" docId="F97A87E27403E66466D1FF65FC98FEDD" docLanguage="en" docName="ZM.82.19.189-210.pdf" docOrigin="Zool. Med. Leiden 82 (19)" docStyle="DocumentStyle{}" docTitle="Ptomaphaginus bryantioides Schilthuizen &amp; Perreau 2008, spec. nov." docType="treatment" docVersion="8" lastPageNumber="193" masterDocId="0543FF9A7400E660674CFFA5FFD2FFE5" masterDocTitle="New species and new records of Ptomaphaginus Portevin from the Sunda region, Southeast Asia (Coleoptera: Leiodidae: Cholevinae)" masterLastPageNumber="210" masterPageNumber="189" pageNumber="192" updateTime="1732603390766" updateUser="ExternalLinkService">
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<mods:title id="71F1716E7535D9B814D5A952DEDF30BF">New species and new records of Ptomaphaginus Portevin from the Sunda region, Southeast Asia (Coleoptera: Leiodidae: Cholevinae)</mods:title>
</mods:titleInfo>
<mods:name id="960B9CC6B3258DCC0F33D7D016DF7C4E" type="personal">
<mods:role id="633F0E1AC9666EFFE3123F5B489FC5EA">
<mods:roleTerm id="334022996AAB000B9FAFA87B7EFE0601">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0CF265957A372309DD589DE94D0B064E">Schilthuizen, Menno</mods:namePart>
</mods:name>
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<mods:namePart id="A98B20F829796188BFD404E400A78C00">Perreau, Michel</mods:namePart>
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<mods:typeOfResource id="867FB0E9D5617B8A4018143030C28DC6">text</mods:typeOfResource>
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<mods:titleInfo id="47306C54D1DD7C64E6ECF751E66E9966">
<mods:title id="5044C10AFA7740CB21A841E25810BFD3">Zool. Med. Leiden</mods:title>
</mods:titleInfo>
<mods:part id="CFF3234F4399C94155824AAB8C9EA631">
<mods:date id="9E2FE808772CA412377C6E7D289E476C">2008</mods:date>
<mods:detail id="C7C9F747CDC6FFA2FD67ADDD41F45F0B" type="pubDate">
<mods:number id="1C520D4D4DD93AED385AFEB60E07B9FC">2008-01-01</mods:number>
</mods:detail>
<mods:detail id="FEAD223791F9CCBEC6D08F06E178035C" type="volume">
<mods:number id="5C8F4100D60046A78C51DA5753D8E04F">82</mods:number>
</mods:detail>
<mods:detail id="3E6FBACF9A1816FCBDAA1C107CF9B05B" type="issue">
<mods:number id="56918FC241FD134B9A2D18EF8CDFD1D2">19</mods:number>
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<mods:start id="7CF889447261D8F4FEA0B5EBDFDCDD12">189</mods:start>
<mods:end id="58606F6301B445EE4B01255BC0F14471">210</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="957ECE5C947FDA4923C2C0731EA4DBAA">journal article</mods:classification>
<mods:identifier id="9765400C09B473D2C2EE763383A2CF4C" type="CLB-Dataset">24614</mods:identifier>
<mods:identifier id="41458F9CC12E5118F5E0800E2CB5928E" type="DOI">10.5281/zenodo.3576287</mods:identifier>
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<treatment id="F97A87E27403E66466D1FF65FC98FEDD" ID-DOI="http://doi.org/10.5281/zenodo.6231982" ID-GBIF-Taxon="160743190" ID-Zenodo-Dep="6231982" LSID="urn:lsid:plazi:treatment:F97A87E27403E66466D1FF65FC98FEDD" httpUri="http://treatment.plazi.org/id/F97A87E27403E66466D1FF65FC98FEDD" lastPageId="4" lastPageNumber="193" pageId="3" pageNumber="192" scope_family="Leiodidae" scope_order="Coleoptera" scope_subFamily="Cholevinae">
<subSubSection id="39C9657F7403E66366D1FF65FC8BFF3D" box="[413,857,192,216]" pageId="3" pageNumber="192" type="nomenclature">
<paragraph id="716C36F47403E66366D1FF65FC8BFF3D" blockId="3.[413,857,192,248]" box="[413,857,192,216]" pageId="3" pageNumber="192">
<heading id="2A2481987403E66366D1FF65FC8BFF3D" box="[413,857,192,216]" centered="true" fontSize="10" level="3" pageId="3" pageNumber="192" reason="1">
<taxonomicName id="B6D34D777403E66366D1FF65FD34FF3D" ID-CoL="4Q988" authority="Schilthuizen &amp; Perreau, 2008" authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[413,742,192,216]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="bryantioides" status="sp. nov.">
<emphasis id="43A7EAE67403E66366D1FF65FD34FF3D" bold="true" box="[413,742,192,216]" italics="true" pageId="3" pageNumber="192">Ptomaphaginus bryantioides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5894579D7403E66365A0FF64FC8BFF3D" box="[748,857,193,216]" pageId="3" pageNumber="192" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="39C9657F7403E6636577FF45FD6EFF1D" box="[571,700,224,248]" pageId="3" pageNumber="192" type="description">
<paragraph id="716C36F47403E6636577FF45FD6EFF1D" blockId="3.[413,857,192,248]" box="[571,700,224,248]" pageId="3" pageNumber="192">
<heading id="2A2481987403E6636577FF45FD6EFF1D" box="[571,700,224,248]" centered="true" fontSize="10" level="3" pageId="3" pageNumber="192" reason="1">
(
<figureCitation id="E9E82A717403E663650EFF45FD62FF1D" box="[578,688,224,248]" captionStart="Figs 16-25. 16" captionStartId="20.[151,190,1437,1456]" captionTargetBox="[151,1119,192,1408]" captionTargetPageId="20" captionText="Figs 16-25. 16, Ptomaphaginus anas spec. nov. holotype; aedeagus, dorsal view. 17, Ptomaphaginus anas spec. nov., holotype; aedeagus, lateral view. 18, Ptomaphaginus similipes spec. nov., holotype; aedeagus, dorsal view. 19, Ptomaphaginus similipes spec. nov., holotype; aedeagus, lateral view. 20, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, dorsal view. 21, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, lateral view (basal part partly obscured by the genital segment). 22, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, ventral view. 23, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, lateral view. 24, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, dorsal view. 25, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576295" httpUri="https://zenodo.org/record/3576295/files/figure.png" pageId="3" pageNumber="192">figs 20-21</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="39C9657F7403E66367DBFE85FF0EFD32" pageId="3" pageNumber="192" type="materials_examined">
<paragraph id="716C36F47403E66367DBFE85FE78FDAE" blockId="3.[151,1120,287,727]" pageId="3" pageNumber="192">
Type material.— 
<materialsCitation id="C1BB3CA97403E6636673FE85FCABFEAA" ID-GBIF-Occurrence="2512737995" collectingDate="2000-05-15" collectionCode="BOR" collectorName="J. de Roode" country="Malaysia" county="Danum Valley Field Centre" latitude="5.0" location="Valley Field Centre" longLatPrecision="1305" longitude="117.833336" municipality="Danum Valley" pageId="3" pageNumber="192" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="AE6888567403E6636673FE85FE4AFED6" box="[319,408,288,307]" pageId="3" pageNumber="192" type="holotype">Holotype</typeStatus>
<specimenCount id="67D5FD7D7403E66366EDFEBAFE61FED6" box="[417,435,287,307]" pageId="3" pageNumber="192" type="male">♂</specimenCount>
: 
<collectingCountry id="09C476647403E663668CFE85FDCAFED6" box="[448,536,288,307]" name="Malaysia" pageId="3" pageNumber="192">Malaysia</collectingCountry>
: 
<collectingRegion id="B317F8167403E6636568FE85FD8DFED6" box="[548,607,288,307]" country="Malaysia" name="Sabah" pageId="3" pageNumber="192">Sabah</collectingRegion>
: 
<collectingCounty id="980D4E787403E6636526FE85FCA7FED6" box="[618,885,288,307]" pageId="3" pageNumber="192">
<collectingMunicipality id="9108AC8E7403E6636526FE85FD27FED6" box="[618,757,288,307]" pageId="3" pageNumber="192">
Danum 
<location id="740C602F7403E66365F6FE85FD27FED6" LSID="urn:lsid:plazi:treatment:F97A87E27403E66466D1FF65FC98FEDD:740C602F7403E66365F6FE85FD27FED6" box="[698,757,288,307]" country="Malaysia" county="Danum Valley Field Centre" latitude="5.0" longLatPrecision="1305" longitude="117.833336" municipality="Danum Valley" name="Valley Field Centre" pageId="3" pageNumber="192" stateProvince="Sabah">Valley</location>
</collectingMunicipality>
Field Centre
</collectingCounty>
(
<locationDeviation id="60DE2FD67403E66364CFFE85FF1AFEAA" location="Lahad Datu" pageId="3" pageNumber="192">
<quantity id="B62B9B117403E66364CFFE85FC13FED6" box="[899,961,288,307]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="192" unit="km" value="60.0">60 km</quantity>
WSW of Lahad Datu
</locationDeviation>
), 
<geoCoordinate id="14E750337403E6636795FE99FECFFEAA" box="[217,285,316,335]" degrees="5" direction="north" minutes="00" orientation="latitude" pageId="3" pageNumber="192" precision="925" value="5.0">5 º00’N</geoCoordinate>
<geoCoordinate id="14E750337403E6636668FE99FEA6FEAA" box="[292,372,316,335]" degrees="117" direction="east" minutes="50" orientation="longitude" pageId="3" pageNumber="192" precision="925" value="117.833336">117º50’E</geoCoordinate>
, 
<date id="056D10347403E6636632FE99FE06FEAA" box="[382,468,316,335]" pageId="3" pageNumber="192" value="2000-05-15">
<collectingDate id="1529E9DC7403E6636632FE99FE06FEAA" box="[382,468,316,335]" pageId="3" pageNumber="192" value="2000-05-15">15.v.2000</collectingDate>
</date>
, 
<collectingMethod id="A8924EE37403E6636692FE99FDC4FEAA" box="[478,534,316,335]" pageId="3" pageNumber="192">pitfall</collectingMethod>
with chicken, leg. 
<collectorName id="DC2653227403E6636585FE99FCE4FEAA" box="[713,822,316,335]" pageId="3" pageNumber="192">J. de Roode</collectorName>
(
<collectionCode id="17C2AE317403E663640FFE99FCA1FEAA" box="[835,883,316,335]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="3" pageNumber="192" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="C1BB3CA97403E66364CEFE99FDFCFE8E" ID-GBIF-Occurrence="2512737998" collectingDate="2000-05-15" collectionCode="BOR" collectorName="J. de Roode" country="Malaysia" county="Danum Valley Field Centre" latitude="5.0" location="Valley Field Centre" longLatPrecision="1305" longitude="117.833336" municipality="Danum Valley" pageId="3" pageNumber="192" specimenCount="11" specimenCount-female="9" specimenCount-male="2" stateProvince="Sabah" typeStatus="paratype">
<typeStatus id="AE6888567403E66364CEFE99FC33FEAA" box="[898,993,316,335]" pageId="3" pageNumber="192" type="paratype">Paratypes</typeStatus>
, same collection data as 
<typeStatus id="AE6888567403E6636644FEFDFE8CFE8E" box="[264,350,344,363]" pageId="3" pageNumber="192" type="holotype">holotype</typeStatus>
: 
<specimenCount id="67D5FD7D7403E663662AFEFDFE4BFE8E" box="[358,409,343,363]" pageId="3" pageNumber="192" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="67D5FD7D7403E66366EFFEFDFE04FE8E" box="[419,470,343,363]" pageId="3" pageNumber="192" type="female">9 ♀♀</specimenCount>
(in 
<collectionCode id="17C2AE317403E66366B5FEFDFDFAFE8E" box="[505,552,344,363]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="3" pageNumber="192" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="C1BB3CA97403E663657AFEFDFDEAFE62" ID-GBIF-Occurrence="2512737972" collectingDate="2000-09-22" collectingDateMax="2000-09-27" collectingDateMin="2000-09-22" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" county="Danum Valley Field Centre" latitude="5.0" location="Valley Field Centre" longLatPrecision="1305" longitude="117.833336" municipality="Danum Valley" pageId="3" pageNumber="192" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Sabah" typeStatus="holotype">
same locality as 
<typeStatus id="AE6888567403E6636582FEFDFCF6FE8E" box="[718,804,344,363]" pageId="3" pageNumber="192" type="holotype">holotype</typeStatus>
, 
<date id="056D10347403E6636460FEFDFC74FE8E" box="[812,934,344,363]" pageId="3" pageNumber="192" value="2000-09-22" valueMax="2000-09-27" valueMin="2000-09-22">
<collectingDate id="1529E9DC7403E6636460FEFDFC74FE8E" box="[812,934,344,363]" pageId="3" pageNumber="192" value="2000-09-22" valueMax="2000-09-27" valueMin="2000-09-22">22-27.ix.2000</collectingDate>
</date>
, 
<collectingMethod id="A8924EE37403E66364E2FEFDFC34FE8E" box="[942,998,344,363]" pageId="3" pageNumber="192">pitfall</collectingMethod>
with human dung, leg. 
<collectorName id="DC2653227403E66367B0FED1FE41FE62" box="[252,403,372,391]" pageId="3" pageNumber="192">M. Schilthuizen</collectorName>
: 
<specimenCount id="67D5FD7D7403E66366D7FED1FE1DFE62" box="[411,463,371,391]" pageId="3" pageNumber="192" type="male">2 ♂♂</specimenCount>
<specimenCount id="67D5FD7D7403E6636699FED1FE25FE62" box="[469,503,371,391]" pageId="3" pageNumber="192" type="female">1 ♀</specimenCount>
(
<collectionCode id="17C2AE317403E663654EFED1FDE0FE62" box="[514,562,372,391]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="3" pageNumber="192" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="C1BB3CA97403E663650EFED1FCE7FE46" ID-GBIF-Occurrence="2512737987" collectingDate="1997-02-27" collectionCode="BOR" country="Malaysia" location="Kiansom" pageId="3" pageNumber="192" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Sabah" typeStatus="paratype">
<collectingCountry id="09C476647403E663650EFED1FD48FE62" box="[578,666,372,391]" name="Malaysia" pageId="3" pageNumber="192">Malaysia</collectingCountry>
: 
<collectingRegion id="B317F8167403E66365EEFED1FD0FFE62" box="[674,733,372,391]" country="Malaysia" name="Sabah" pageId="3" pageNumber="192">Sabah</collectingRegion>
: 
<location id="740C602F7403E66365AAFED1FCE8FE62" LSID="urn:lsid:plazi:treatment:F97A87E27403E66466D1FF65FC98FEDD:740C602F7403E66365AAFED1FCE8FE62" box="[742,826,372,391]" country="Malaysia" name="Kiansom" pageId="3" pageNumber="192" stateProvince="Sabah">Kiansom</location>
(
<locationDeviation id="60DE2FD67403E663640AFED1FB87FE62" box="[838,1109,372,391]" location="Kota Kinabalu" pageId="3" pageNumber="192">
<quantity id="B62B9B117403E663640AFED1FC53FE62" box="[838,897,372,391]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="192" unit="km" value="20.0">20 km</quantity>
ESE of Kota Kinabalu
</locationDeviation>
), 
<date id="056D10347403E66367DBFE35FF22FE46" box="[151,240,400,419]" pageId="3" pageNumber="192" value="1997-02-27">
<collectingDate id="1529E9DC7403E66367DBFE35FF22FE46" box="[151,240,400,419]" pageId="3" pageNumber="192" value="1997-02-27">27.ii.1997</collectingDate>
</date>
, 
<collectingMethod id="A8924EE37403E66367B5FE35FE03FE46" box="[249,465,400,419]" pageId="3" pageNumber="192">flight interception trap</collectingMethod>
, leg. UMS students: 
<specimenCount id="67D5FD7D7403E66365DEFE35FD15FE46" box="[658,711,399,419]" pageId="3" pageNumber="192" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="67D5FD7D7403E663659DFE35FD26FE46" box="[721,756,399,419]" pageId="3" pageNumber="192" type="female">1 ♀</specimenCount>
(
<collectionCode id="17C2AE317403E66365B3FE35FCFDFE46" box="[767,815,400,419]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="3" pageNumber="192" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="C1BB3CA97403E6636472FE35FD46FE5A" ID-GBIF-Occurrence="2512737946" collectingDate="2000-04-11" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" location="Kiansom" pageId="3" pageNumber="192" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
same locality, 
<date id="056D10347403E663648EFE35FBCFFE46" box="[962,1053,400,419]" pageId="3" pageNumber="192" value="2000-04-11">
<collectingDate id="1529E9DC7403E663648EFE35FBCFFE46" box="[962,1053,400,419]" pageId="3" pageNumber="192" value="2000-04-11">11.iv.2000</collectingDate>
</date>
, 
<collectingMethod id="A8924EE37403E663636BFE35FB8DFE46" box="[1063,1119,400,419]" pageId="3" pageNumber="192">pitfall</collectingMethod>
with chicken, leg. 
<collectorName id="DC2653227403E6636604FE09FE32FE5A" box="[328,480,428,447]" pageId="3" pageNumber="192">M. Schilthuizen</collectorName>
, 
<specimenCount id="67D5FD7D7403E66366A6FE09FDF3FE5A" box="[490,545,427,447]" pageId="3" pageNumber="192" type="male">1 ♂♂</specimenCount>
, 
<specimenCount id="67D5FD7D7403E6636561FE09FD83FE5A" box="[557,593,427,447]" pageId="3" pageNumber="192" type="female">1 ♀</specimenCount>
(
<collectionCode id="17C2AE317403E6636512FE09FD5CFE5A" box="[606,654,428,447]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="3" pageNumber="192" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="C1BB3CA97403E66365D3FE09FF01FE12" ID-GBIF-Occurrence="2512737996" collectingDate="2000-04-10" collectingDateMax="2000-04-12" collectingDateMin="2000-04-10" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" location="Kota Kinabalu" municipality="Tun Fuad Stephen Municipal Park" pageId="3" pageNumber="192" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Sabah" typeStatus="paratype">
<collectingCountry id="09C476647403E66365D3FE09FD25FE5A" box="[671,759,428,447]" name="Malaysia" pageId="3" pageNumber="192">Malaysia</collectingCountry>
: 
<collectingRegion id="B317F8167403E663644DFE09FCEEFE5A" box="[769,828,428,447]" country="Malaysia" name="Sabah" pageId="3" pageNumber="192">Sabah</collectingRegion>
: 
<collectingMunicipality id="9108AC8E7403E663640BFE09FF11FE3E" pageId="3" pageNumber="192">Tun Fuad Stephen Municipal Park</collectingMunicipality>
(outskirts of 
<location id="740C602F7403E6636605FE6DFE05FE3E" LSID="urn:lsid:plazi:treatment:F97A87E27403E66466D1FF65FC98FEDD:740C602F7403E6636605FE6DFE05FE3E" box="[329,471,456,475]" country="Malaysia" municipality="Tun Fuad Stephen Municipal Park" name="Kota Kinabalu" pageId="3" pageNumber="192" stateProvince="Sabah">Kota Kinabalu</location>
), 
<date id="056D10347403E66366A5FE6DFDB3FE3E" box="[489,609,456,475]" pageId="3" pageNumber="192" value="2000-04-10" valueMax="2000-04-12" valueMin="2000-04-10">
<collectingDate id="1529E9DC7403E66366A5FE6DFDB3FE3E" box="[489,609,456,475]" pageId="3" pageNumber="192" value="2000-04-10" valueMax="2000-04-12" valueMin="2000-04-10">10-12.iv.2000</collectingDate>
</date>
, 
<collectingMethod id="A8924EE37403E6636521FE6DFD77FE3E" box="[621,677,456,475]" pageId="3" pageNumber="192">pitfall</collectingMethod>
with lamb, leg. 
<collectorName id="DC2653227403E6636404FE6DFC33FE3E" box="[840,993,456,475]" pageId="3" pageNumber="192">M. Schilthuizen</collectorName>
, 
<specimenCount id="67D5FD7D7403E66364A1FE6DFBF1FE3E" box="[1005,1059,455,475]" pageId="3" pageNumber="192" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="67D5FD7D7403E663637DFE6DFB8DFE3E" box="[1073,1119,455,475]" pageId="3" pageNumber="192" type="female">5♀♀</specimenCount>
(
<collectionCode id="17C2AE317403E66367D1FE41FF1FFE12" box="[157,205,484,503]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="3" pageNumber="192" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="C1BB3CA97403E6636797FE41FE97FE12" ID-GBIF-Occurrence="2512737969" box="[219,325,483,503]" collectingDate="2000-04-10" collectingDateMax="2000-04-12" collectingDateMin="2000-04-10" collectionCode="FRCS" collectorName="M. Schilthuizen" country="Malaysia" isTailEnumeration="true" location="Kota Kinabalu" municipality="Tun Fuad Stephen Municipal Park" pageId="3" pageNumber="192" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<specimenCount id="67D5FD7D7403E6636797FE41FF2EFE12" box="[219,252,483,503]" pageId="3" pageNumber="192" type="male">1 ♂</specimenCount>
(
<collectionCode id="17C2AE317403E663664BFE41FEEDFE12" box="[263,319,484,503]" country="Malaysia" httpUri="http://grbio.org/cool/sxta-ddia" name="Forest Research Centre" pageId="3" pageNumber="192">FRCS</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="C1BB3CA97403E6636601FE41FF01FDF6" ID-GBIF-Occurrence="2512737993" collectingDate="2000-11-24" collectingDateMax="2000-11-27" collectingDateMin="2000-11-24" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" isTailEnumeration="true" location="Kota Kinabalu" municipality="Tun Fuad Stephen Municipal Park" pageId="3" pageNumber="192" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
same locality, 
<date id="056D10347403E6636683FE41FD98FE12" box="[463,586,484,503]" pageId="3" pageNumber="192" value="2000-11-24" valueMax="2000-11-27" valueMin="2000-11-24">
<collectingDate id="1529E9DC7403E6636683FE41FD98FE12" box="[463,586,484,503]" pageId="3" pageNumber="192" value="2000-11-24" valueMax="2000-11-27" valueMin="2000-11-24">24-27.xi.2000</collectingDate>
</date>
, 
<collectingMethod id="A8924EE37403E663651DFE41FD5BFE12" box="[593,649,484,503]" pageId="3" pageNumber="192">pitfall</collectingMethod>
with dung or beef, leg. 
<collectorName id="DC2653227403E6636428FE41FC2BFE12" box="[868,1017,484,503]" pageId="3" pageNumber="192">M. Schilthuizen</collectorName>
, 
<specimenCount id="67D5FD7D7403E663634DFE41FBF0FE12" box="[1025,1058,483,503]" pageId="3" pageNumber="192" type="male">1 ♂</specimenCount>
, 
<specimenCount id="67D5FD7D7403E6636360FE41FB8DFE12" box="[1068,1119,483,503]" pageId="3" pageNumber="192" type="female">3 ♀♀</specimenCount>
(
<collectionCode id="17C2AE317403E66367D1FDA5FF1FFDF6" box="[157,205,512,531]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="3" pageNumber="192" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="C1BB3CA97403E6636790FDA5FEAAFDCA" ID-GBIF-Occurrence="2512737948" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" elevation="200" location="Sugud Forest Reserve" pageId="3" pageNumber="192" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="paratype">
<location id="740C602F7403E6636790FDA5FE79FDF6" LSID="urn:lsid:plazi:treatment:F97A87E27403E66466D1FF65FC98FEDD:740C602F7403E6636790FDA5FE79FDF6" box="[220,427,512,531]" country="Malaysia" name="Sugud Forest Reserve" pageId="3" pageNumber="192" stateProvince="Sabah">Sugud Forest Reserve</location>
, 
<locationDeviation id="60DE2FD67403E66366F8FDA5FD76FDF6" box="[436,676,512,531]" location="Kota Kinabalu" pageId="3" pageNumber="192">
<quantity id="B62B9B117403E66366F8FDA5FE3CFDF6" box="[436,494,512,531]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="192" unit="km" value="15.0">15 km</quantity>
S of Kota Kinabalu
</locationDeviation>
(
<collectingRegion id="B317F8167403E66365FCFDA5FD3EFDF6" box="[688,748,512,531]" country="Malaysia" name="Sabah" pageId="3" pageNumber="192">Sabah</collectingRegion>
), 
<quantity id="B62B9B117403E66365B0FDA5FCE6FDF6" box="[764,820,512,531]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="192" unit="m" value="200.0">
<elevation id="FAFED1C77403E66365B0FDA5FCE6FDF6" box="[764,820,512,531]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="192" unit="m" value="200.0">200 m</elevation>
</quantity>
alt., 
<collectingMethod id="A8924EE37403E663642EFDA5FC48FDF6" box="[866,922,512,531]" pageId="3" pageNumber="192">pitfall</collectingMethod>
with carrion, leg. 
<collectorName id="DC2653227403E6636309FDA5FEDCFDCA" pageId="3" pageNumber="192">M. Schilthuizen</collectorName>
, 
<specimenCount id="67D5FD7D7403E663665AFDB9FEEAFDCA" box="[278,312,539,559]" pageId="3" pageNumber="192" type="female">1 ♀</specimenCount>
(
<collectionCode id="17C2AE317403E663660EFDB9FEA0FDCA" box="[322,370,540,559]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="3" pageNumber="192" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="C1BB3CA97403E66366CCFDB9FE77FDAE" ID-GBIF-Occurrence="2512737956" collectingDate="1996-06-24" collectingDateMax="1996-07-01" collectingDateMin="1996-06-24" collectionCode="CJRZ, CMPR" country="Malaysia" location="Batu Punggul Resort" pageId="3" pageNumber="192" specimenCount="20" specimenCount-female="7" specimenCount-male="13" stateProvince="Sabah" typeStatus="paratype">
<collectingCountry id="09C476647403E66366CCFDB9FE0AFDCA" box="[384,472,540,559]" name="Malaysia" pageId="3" pageNumber="192">Malaysia</collectingCountry>
: 
<collectingRegion id="B317F8167403E66366ACFDB9FDC9FDCA" box="[480,539,540,559]" country="Malaysia" name="Sabah" pageId="3" pageNumber="192">Sabah</collectingRegion>
: 
<location id="740C602F7403E663656FFDB9FD35FDCA" LSID="urn:lsid:plazi:treatment:F97A87E27403E66466D1FF65FC98FEDD:740C602F7403E663656FFDB9FD35FDCA" box="[547,743,540,559]" country="Malaysia" name="Batu Punggul Resort" pageId="3" pageNumber="192" stateProvince="Sabah">Batu Punggul Resort</location>
, 
<date id="056D10347403E66365A3FDB9FCAFFDCA" box="[751,893,540,559]" pageId="3" pageNumber="192" value="1996-06-24" valueMax="1996-07-01" valueMin="1996-06-24">
<collectingDate id="1529E9DC7403E66365A3FDB9FCAFFDCA" box="[751,893,540,559]" pageId="3" pageNumber="192" value="1996-06-24" valueMax="1996-07-01" valueMin="1996-06-24">24.vi-1.vii.1996</collectingDate>
</date>
, 
<collectingMethod id="A8924EE37403E66364C9FDB9FB8EFDCA" box="[901,1116,540,559]" pageId="3" pageNumber="192">flight interception trap</collectingMethod>
, 
<specimenCount id="67D5FD7D7403E66367DBFD9DFF04FDAE" box="[151,214,567,587]" pageId="3" pageNumber="192" type="male">13 ♂♂</specimenCount>
, 
<specimenCount id="67D5FD7D7403E66367ACFD9DFEC6FDAE" box="[224,276,567,587]" pageId="3" pageNumber="192" type="female">7 ♀♀</specimenCount>
(
<collectionCode id="17C2AE317403E6636653FD9DFE87FDAE" box="[287,341,568,587]" pageId="3" pageNumber="192">CJRZ</collectionCode>
; 
<collectionCode id="17C2AE317403E6636611FD9DFE4DFDAE" box="[349,415,568,587]" country="India" name="Centre for Medicinal Plants Research" pageId="3" pageNumber="192" type="Herbarium">CMPR</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="716C36F47403E66367DBFDF1FF0EFD32" blockId="3.[151,1120,287,727]" pageId="3" pageNumber="192">
<materialsCitation id="C1BB3CA97403E66367DBFDF1FF0EFD32" ID-GBIF-Occurrence="2512737963" collectingDate="1997-02-27" collectingDateMax="2000-12-29" collectingDateMin="1997-02-27" location="Valley Field" municipality="Danum Valley" pageId="3" pageNumber="192" specimenCount="25" specimenCount-female="9" specimenCount-male="4" stateProvince="Centre">
Additional material (not included in the type series).— Tun Fuad Stephen Municipal Park, 
<date id="056D10347403E66364ADFDF1FB89FD82" box="[993,1115,596,615]" pageId="3" pageNumber="192" value="2000-09-24" valueMax="2000-09-27" valueMin="2000-09-24">
<collectingDate id="1529E9DC7403E66364ADFDF1FB89FD82" box="[993,1115,596,615]" pageId="3" pageNumber="192" value="2000-09-24" valueMax="2000-09-27" valueMin="2000-09-24">24-27.ix.2000</collectingDate>
</date>
, 
<specimenCount id="67D5FD7D7403E66367DBFDD5FF1EFD66" box="[151,204,623,643]" pageId="3" pageNumber="192" type="male">4 ♂♂</specimenCount>
, 
<specimenCount id="67D5FD7D7403E6636794FDD5FEDFFD66" box="[216,269,623,643]" pageId="3" pageNumber="192" type="female">9 ♀♀</specimenCount>
in 70% ethanol; Kiansom, 
<date id="056D10347403E6636542FDD5FD5CFD66" box="[526,654,624,643]" pageId="3" pageNumber="192" value="2000-12-25" valueMax="2000-12-29" valueMin="2000-12-25">
<collectingDate id="1529E9DC7403E6636542FDD5FD5CFD66" box="[526,654,624,643]" pageId="3" pageNumber="192" value="2000-12-25" valueMax="2000-12-29" valueMin="2000-12-25">25-29.xii.2000</collectingDate>
</date>
, 
<specimenCount id="67D5FD7D7403E66365D4FDD5FCD9FD66" box="[664,779,624,643]" pageId="3" pageNumber="192" type="generic">8 specimens</specimenCount>
of unidentified sex in 70% ethanol; Kiansom, 
<date id="056D10347403E66367B6FD29FE81FD7A" box="[250,339,652,671]" pageId="3" pageNumber="192" value="1997-02-27">
<collectingDate id="1529E9DC7403E66367B6FD29FE81FD7A" box="[250,339,652,671]" pageId="3" pageNumber="192" value="1997-02-27">27.ii.1997</collectingDate>
</date>
, 
<specimenCount id="67D5FD7D7403E663662CFD29FE04FD7A" box="[352,470,652,671]" pageId="3" pageNumber="192" type="generic">2 specimens</specimenCount>
of unidentified sex in 100% ethanol; 
<collectingMunicipality id="9108AC8E7403E6636401FD29FC0BFD7A" box="[845,985,652,671]" pageId="3" pageNumber="192">
Danum 
<location id="740C602F7403E66364D2FD29FC0BFD7A" LSID="urn:lsid:plazi:treatment:F97A87E27403E66466D1FF65FC98FEDD:740C602F7403E66364D2FD29FC0BFD7A" box="[926,985,652,671]" municipality="Danum Valley" name="Valley Field" pageId="3" pageNumber="192" stateProvince="Centre">Valley</location>
</collectingMunicipality>
Field 
<collectingRegion id="B317F8167403E6636356FD29FB89FD7A" box="[1050,1115,652,671]" country="France" name="Centre" pageId="3" pageNumber="192">Centre</collectingRegion>
, 
<date id="056D10347403E66367DBFD0DFF3CFD5E" box="[151,238,680,699]" pageId="3" pageNumber="192" value="2000-05-15">
<collectingDate id="1529E9DC7403E66367DBFD0DFF3CFD5E" box="[151,238,680,699]" pageId="3" pageNumber="192" value="2000-05-15">15.v.2000</collectingDate>
</date>
, 
<specimenCount id="67D5FD7D7403E66367BAFD0DFEBAFD5E" box="[246,360,680,699]" pageId="3" pageNumber="192" type="generic">2 specimens</specimenCount>
of unidentified sex in 100% ethanol. (For full collection details, see Type material above.)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="39C9657F7403E663678DFCA5FB99FCBD" pageId="3" pageNumber="192" type="diagnosis">
<paragraph id="716C36F47403E663678DFCA5FB99FCBD" blockId="3.[151,1119,768,1657]" pageId="3" pageNumber="192">
Diagnosis.— Habitus broad, rectangular, flat. Thorax on average 1.8 times as wide as long. Aedeagus (
<figureCitation id="E9E82A717403E663663CFC85FE0EFCDD" box="[368,476,800,824]" captionStart="Figs 16-25. 16" captionStartId="20.[151,190,1437,1456]" captionTargetBox="[151,1119,192,1408]" captionTargetPageId="20" captionText="Figs 16-25. 16, Ptomaphaginus anas spec. nov. holotype; aedeagus, dorsal view. 17, Ptomaphaginus anas spec. nov., holotype; aedeagus, lateral view. 18, Ptomaphaginus similipes spec. nov., holotype; aedeagus, dorsal view. 19, Ptomaphaginus similipes spec. nov., holotype; aedeagus, lateral view. 20, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, dorsal view. 21, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, lateral view (basal part partly obscured by the genital segment). 22, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, ventral view. 23, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, lateral view. 24, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, dorsal view. 25, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576295" httpUri="https://zenodo.org/record/3576295/files/figure.png" pageId="3" pageNumber="192">figs 20-21</figureCitation>
) with two apical lateral ‘wings’ and a long terminal processus. Male forelegs with long hairs on the ventral side of the profemur and protibia.
</paragraph>
</subSubSection>
<subSubSection id="39C9657F7403E663678DFCC5FEFEFA5D" pageId="3" pageNumber="192" type="description">
<paragraph id="716C36F47403E663678DFCC5FEFEFA5D" blockId="3.[151,1119,768,1657]" pageId="3" pageNumber="192">
Description.— Habitus broad and rectangular, relatively flat. Moderately pigment- ed: mostly chestnut brown; only the tarsi and the edges of thorax and elytra lighter brown. Length 
<quantity id="B62B9B117403E663660FFC05FE6DFC5D" box="[323,447,928,952]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.9" metricValueMin="2.1" pageId="3" pageNumber="192" unit="mm" value="2.5" valueMax="2.9" valueMin="2.1">2.1-2.9 mm</quantity>
(n = 26). Antennae slender and relatively long, 1.1-1.2 times as long as the width of the head (n = 5). Articles 6, 9, and 10 square. Male protarsi moderately dilated: the protibia at its largest width (excluding the lateral spines) is 1.5 times broader than the first article of the protarsus (n = 1). Female protarsi undilated. In the male, the ventral side of profemur and protibia carry long hairs, some of which are as long as the tarsus. Thorax 1.76-1.84 times as wide as long (n = 5), relatively flat, with the caudal corners distinctly drawn out. The mesosternal carina is strongly developed, its edge somewhat thickened. Elytra short, as long (measured from the caudal tip of the scutellum) as their combined width. Elytral apices distinctly truncate, identical in both sexes. The 3 
<superScript id="86A69BBC7403E6636651FB1AFEE6FB3D" attach="none" box="[285,308,1215,1240]" fontSize="6" pageId="3" pageNumber="192">rd-</superScript>
5 
<superScript id="86A69BBC7403E663660DFB1AFE9CFB28" attach="left" box="[321,334,1215,1229]" fontSize="6" pageId="3" pageNumber="192">th</superScript>
visible abdominal sternite in the male each have a broad and very indistinct central notch and are slightly depressed in the vicinity of these notches. Aedeagus (
<figureCitation id="E9E82A717403E6636780FAA5FEE4FAFD" box="[204,310,1280,1304]" captionStart="Figs 16-25. 16" captionStartId="20.[151,190,1437,1456]" captionTargetBox="[151,1119,192,1408]" captionTargetPageId="20" captionText="Figs 16-25. 16, Ptomaphaginus anas spec. nov. holotype; aedeagus, dorsal view. 17, Ptomaphaginus anas spec. nov., holotype; aedeagus, lateral view. 18, Ptomaphaginus similipes spec. nov., holotype; aedeagus, dorsal view. 19, Ptomaphaginus similipes spec. nov., holotype; aedeagus, lateral view. 20, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, dorsal view. 21, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, lateral view (basal part partly obscured by the genital segment). 22, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, ventral view. 23, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, lateral view. 24, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, dorsal view. 25, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576295" httpUri="https://zenodo.org/record/3576295/files/figure.png" pageId="3" pageNumber="192">figs 20-21</figureCitation>
) in dorsal view abruptly narrowing towards the apex, and there adorned with two lateral ‘wings’. The tip is composed of a long, fragile and slightly asymmetric processus. In lateral view, the aedeagus is moderately curved, the terminal processus lies in the plane of the rest of the aedeagus, whereas the two lateral wings point ventrad. Male genital segment relatively broad, at the apex slightly triangular, 2.5 times as long as wide.
</paragraph>
</subSubSection>
<subSubSection id="39C9657F7403E663678DFA65FE38F9DD" pageId="3" pageNumber="192" type="distribution">
<paragraph id="716C36F47403E663678DFA65FE38F9DD" blockId="3.[151,1119,768,1657]" pageId="3" pageNumber="192">
Distribution.— This species is widely distributed in the lowlands of 
<collectingRegion id="B317F8167403E6636486FA65FBDFFA3D" box="[970,1037,1472,1496]" country="Malaysia" name="Sabah" pageId="3" pageNumber="192">Sabah</collectingRegion>
(all localities lie below 
<quantity id="B62B9B117403E663661FFA45FE46FA1D" box="[339,404,1504,1528]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="192" unit="m" value="300.0">300 m</quantity>
altitude), and apparently has a broad tolerance for habitat 
<typeStatus id="AE6888567403E6636350FA44FB88FA1C" box="[1052,1114,1505,1529]" pageId="3" pageNumber="192">types</typeStatus>
, having been found both in primary (Danum Valley) and in secondary forests (Tun Fuad Stephen Park, Kota Kinabalu).
</paragraph>
</subSubSection>
<subSubSection id="39C9657F7403E664678DF9E5FE22FF1D" lastPageId="4" lastPageNumber="193" pageId="3" pageNumber="192" type="discussion">
<paragraph id="716C36F47403E664678DF9E5FE22FF1D" blockId="3.[151,1119,768,1657]" lastBlockId="4.[151,1119,192,312]" lastPageId="4" lastPageNumber="193" pageId="3" pageNumber="192">
Remarks.— Based on aedeagal shape, 
<taxonomicName id="B6D34D777403E663653FF9E4FCD9F9BD" authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[627,779,1600,1624]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="bryantioides">
<emphasis id="43A7EAE67403E663653FF9E4FCD9F9BD" box="[627,779,1600,1624]" italics="true" pageId="3" pageNumber="192">P. bryantioides</emphasis>
</taxonomicName>
is closely related to 
<taxonomicName id="B6D34D777403E66364BFF9E4FB8AF9BD" authorityName="Jeannel" authorityYear="1936" box="[1011,1112,1600,1624]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="43A7EAE67403E66364BFF9E4FB8AF9BD" box="[1011,1112,1600,1624]" italics="true" pageId="3" pageNumber="192">P. bryanti</emphasis>
</taxonomicName>
, from which it differs in the habitus, which is very broad and stocky in 
<taxonomicName id="B6D34D777403E663648CF9C4FB8BF99D" authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[960,1113,1632,1656]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="bryantioides">
<emphasis id="43A7EAE67403E663648CF9C4FB8BF99D" box="[960,1113,1632,1656]" italics="true" pageId="3" pageNumber="192">P. bryantioides</emphasis>
</taxonomicName>
. For example, the thoracal index for the new species is 1.76-1.84, whereas for the 
<typeStatus id="AE6888567404E6646369FF65FF1BFF1C" pageId="4" pageNumber="193" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="B6D34D777404E66467A0FF44FE82FF1D" authorityName="Jeannel" authorityYear="1936" box="[236,336,224,248]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="43A7EAE67404E66467A0FF44FE82FF1D" box="[236,336,224,248]" italics="true" pageId="4" pageNumber="193">P. bryanti</emphasis>
</taxonomicName>
it is only 1.56.
</paragraph>
</subSubSection>
<subSubSection id="39C9657F7404E664678DFEA5FC98FEDD" pageId="4" pageNumber="193" type="etymology">
<paragraph id="716C36F47404E664678DFEA5FC98FEDD" blockId="4.[151,1119,192,312]" pageId="4" pageNumber="193">
Etymology.— 
<taxonomicName id="B6D34D777404E6646613FEA5FE0FFEFD" authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[351,477,256,280]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="bryantioides">
<emphasis id="43A7EAE67404E6646613FEA5FE0FFEFD" box="[351,477,256,280]" italics="true" pageId="4" pageNumber="193">bryantioides</emphasis>
</taxonomicName>
, a Latin adjective, meaning ‘like 
<taxonomicName id="B6D34D777404E6646400FEA5FC45FEFD" authorityName="Jeannel" authorityYear="1936" box="[844,919,256,280]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="43A7EAE67404E6646400FEA5FC45FEFD" box="[844,919,256,280]" italics="true" pageId="4" pageNumber="193">bryanti</emphasis>
</taxonomicName>
’. The name refers to the similarity in aedeagal shape between these two species.
</paragraph>
</subSubSection>
</treatment>
</document>