<document id="2ED112794EE349E842F38106259EB169" ID-CLB-Dataset="24614" ID-DOI="10.5281/zenodo.3576287" ID-GBIF-Dataset="80959f8f-aa73-499e-93ec-5c07458c7935" ID-ISSN="0024-0672" ID-Zenodo-Dep="3576287" IM.materialsCitations_requiresApprovalFor="plazi" IM.metadata_requiresApprovalFor="plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1576401033600" checkinUser="plazi" docAuthor="Schilthuizen, Menno &amp; Perreau, Michel" docDate="2008" docId="F97A87E2740DE66E66EFFC7FFBF8FE7D" docLanguage="en" docName="ZM.82.19.189-210.pdf" docOrigin="Zool. Med. Leiden 82 (19)" docStyle="DocumentStyle{}" docTitle="Ptomaphaginus burckhardti Schilthuizen &amp; Perreau 2008, spec. nov." docType="treatment" docVersion="8" lastPageNumber="203" masterDocId="0543FF9A7400E660674CFFA5FFD2FFE5" masterDocTitle="New species and new records of Ptomaphaginus Portevin from the Sunda region, Southeast Asia (Coleoptera: Leiodidae: Cholevinae)" masterLastPageNumber="210" masterPageNumber="189" pageNumber="202" updateTime="1732603390766" updateUser="ExternalLinkService">
<mods:mods id="6FAFD856511EA4B73DDA420A717FCBD5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8A42B869FE01671A0891D4F918436712">
<mods:title id="CA258B7EAEF7662C74BD0E59907412DF">New species and new records of Ptomaphaginus Portevin from the Sunda region, Southeast Asia (Coleoptera: Leiodidae: Cholevinae)</mods:title>
</mods:titleInfo>
<mods:name id="0D16130E8343C12FB06A442D630E9165" type="personal">
<mods:role id="22502DB692A4A5CA9C5D2A94FB312BF8">
<mods:roleTerm id="0ECD60BBECCBD6E8CF3546CCD819BEBF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="30A85D674B5149516947187059C6BC66">Schilthuizen, Menno</mods:namePart>
</mods:name>
<mods:name id="89138D0C8B51C727BE41810243F2AD34" type="personal">
<mods:role id="29A35B05DADEB1E2A2FC220F1E4CF72F">
<mods:roleTerm id="DEE5E80905EBA87ADF046285CC1F0F33">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B7800E9275680DC5DACA8C0910C38D0C">Perreau, Michel</mods:namePart>
</mods:name>
<mods:typeOfResource id="F44B2448429B2956247FF960399AFF0F">text</mods:typeOfResource>
<mods:relatedItem id="432BB09F380DE34FCF029F3DD167C3C0" type="host">
<mods:titleInfo id="E39E6518B882E9F2FF795106143EA26A">
<mods:title id="7934FFDC4BD865F137700363D67455D8">Zool. Med. Leiden</mods:title>
</mods:titleInfo>
<mods:part id="DD7165A500CB40BF0A8BF6A6B427BE81">
<mods:date id="37D52C1FE9EB6664F83C2362AA66FBCD">2008</mods:date>
<mods:detail id="FFFB0232BB515A9113007BC1AE073E61" type="pubDate">
<mods:number id="5312DFFAF5D9EF02F80E946A6FA6BF83">2008-01-01</mods:number>
</mods:detail>
<mods:detail id="3325AC431BEDE7615B8910093F70EC00" type="volume">
<mods:number id="DBEC11D02330DD391690B8D3EEAB7B39">82</mods:number>
</mods:detail>
<mods:detail id="562492DAB584C4B735CA8B83A013DD34" type="issue">
<mods:number id="D7A4856EDB44092D004117C1B39524D3">19</mods:number>
</mods:detail>
<mods:extent id="4BA987B057BD56BA4FB281E75CEE5596" unit="page">
<mods:start id="94172726581CCC71DC9A22890038C68A">189</mods:start>
<mods:end id="B1A5EA79147880E251AE384C22751E42">210</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="062CA975A9A3F8F1138960DB97984603">journal article</mods:classification>
<mods:identifier id="519F2320C1ED6F4BAAAF96149AA79481" type="CLB-Dataset">24614</mods:identifier>
<mods:identifier id="DF520E279A0390F6A7413CB4A7E15AC9" type="DOI">10.5281/zenodo.3576287</mods:identifier>
<mods:identifier id="96A274158E89A1B4A6FE6A1E441FF050" type="GBIF-Dataset">80959f8f-aa73-499e-93ec-5c07458c7935</mods:identifier>
<mods:identifier id="471D03FEA637D155DF5EE70B57C85526" type="ISSN">0024-0672</mods:identifier>
<mods:identifier id="95BE2D967B76E9182EF19D36671383D3" type="Zenodo-Dep">3576287</mods:identifier>
</mods:mods>
<treatment id="F97A87E2740DE66E66EFFC7FFBF8FE7D" ID-DOI="http://doi.org/10.5281/zenodo.6232016" ID-GBIF-Taxon="160743173" ID-Zenodo-Dep="6232016" LSID="urn:lsid:plazi:treatment:F97A87E2740DE66E66EFFC7FFBF8FE7D" httpUri="http://treatment.plazi.org/id/F97A87E2740DE66E66EFFC7FFBF8FE7D" lastPageId="14" lastPageNumber="203" pageId="13" pageNumber="202" scope_family="Leiodidae" scope_order="Coleoptera" scope_subFamily="Cholevinae">
<subSubSection id="39C9657F740DE66D66EFFC7FFC86FC17" box="[419,852,986,1010]" pageId="13" pageNumber="202" type="nomenclature">
<paragraph id="716C36F4740DE66D66EFFC7FFC86FC17" blockId="13.[419,852,986,1042]" box="[419,852,986,1010]" pageId="13" pageNumber="202">
<heading id="2A248198740DE66D66EFFC7FFC86FC17" box="[419,852,986,1010]" centered="true" fontSize="10" level="3" pageId="13" pageNumber="202" reason="1">
<taxonomicName id="B6D34D77740DE66D66EFFC7FFD33FC17" ID-CoL="6WLRC" authority="Schilthuizen &amp; Perreau, 2008" authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[419,737,986,1010]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="202" phylum="Arthropoda" rank="species" species="burckhardti" status="sp. nov.">
<emphasis id="43A7EAE6740DE66D66EFFC7FFD33FC17" bold="true" box="[419,737,986,1010]" italics="true" pageId="13" pageNumber="202">Ptomaphaginus burckhardti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5894579D740DE66D65ABFC7EFC86FC17" box="[743,852,987,1010]" pageId="13" pageNumber="202" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="39C9657F740DE66D650BFC5FFD7DFBF7" box="[583,687,1018,1042]" pageId="13" pageNumber="202" type="description">
<paragraph id="716C36F4740DE66D650BFC5FFD7DFBF7" blockId="13.[419,852,986,1042]" box="[583,687,1018,1042]" pageId="13" pageNumber="202">
<heading id="2A248198740DE66D650BFC5FFD7DFBF7" box="[583,687,1018,1042]" centered="true" fontSize="10" level="3" pageId="13" pageNumber="202" reason="1">
(
<figureCitation id="E9E82A71740DE66D6503FC5FFD70FBF7" box="[591,674,1018,1042]" captionStart="Figs 7-12" captionStartId="17.[151,190,1413,1432]" captionTargetId="figure@18.[493,888,185,983]" captionTargetPageId="18" captionText="Figs 7-12. Fig. 7, Ptomaphaginus burckhardti spec. nov., holotype; aedeagus, dorsal view. Fig. 8, P. burckhardti spec. nov., holotype; aedeagus, lateral view. Fig. 9, P. kurbatovi spec. nov., paratype (Cibodas); aedeagus, dorsal view. Fig. 10, P. kurbatovi spec. nov., paratype (Cibodas); aedeagus, lateral view. Fig. 11, P. sabahensis spec. nov., holotype; aedeagus, dorsal view. Fig. 12, P. sabahensis spec. nov., holotype; aedeagus, lateral view. All scale lines 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3576291" httpUri="https://zenodo.org/record/3576291/files/figure.png" pageId="13" pageNumber="202">figs 7-8</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="39C9657F740DE66D67DBFB93FCD5FB80" pageId="13" pageNumber="202" type="materials_examined">
<paragraph id="716C36F4740DE66D67DBFB93FCD5FB80" blockId="13.[151,1118,1077,1125]" pageId="13" pageNumber="202">
Type material.— 
<materialsCitation id="C1BB3CA9740DE66D6677FB93FE9BFB80" ID-GBIF-Occurrence="2512737951" collectingDate="1987-05-01" collectionCode="MHNG" collectorName="Burckhardt &amp; Lobl" country="Malaysia" elevation="2600" location="Mount Kinabalu" pageId="13" pageNumber="202" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="AE688856740DE66D6677FB93FE46FBAC" box="[315,404,1078,1097]" pageId="13" pageNumber="202" type="holotype">Holotype</typeStatus>
<specimenCount id="67D5FD7D740DE66D66D7FB90FE7FFBAC" box="[411,429,1077,1097]" pageId="13" pageNumber="202" type="male">♂</specimenCount>
: 
<collectingCountry id="09C47664740DE66D66F5FB93FDC3FBAC" box="[441,529,1078,1097]" name="Malaysia" pageId="13" pageNumber="202">Malaysia</collectingCountry>
: 
<collectingRegion id="B317F816740DE66D6556FB93FD87FBAC" box="[538,597,1078,1097]" country="Malaysia" name="Sabah" pageId="13" pageNumber="202">Sabah</collectingRegion>
: 
<location id="740C602F740DE66D6513FB93FD2CFBAC" LSID="urn:lsid:plazi:treatment:F97A87E2740DE66E66EFFC7FFBF8FE7D:740C602F740DE66D6513FB93FD2CFBAC" box="[607,766,1078,1097]" country="Malaysia" name="Mount Kinabalu" pageId="13" pageNumber="202" stateProvince="Sabah">Mount Kinabalu</location>
, 
<quantity id="B62B9B11740DE66D6444FB93FC9EFBAC" box="[776,844,1078,1097]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="13" pageNumber="202" unit="m" value="2600.0">
<elevation id="FAFED1C7740DE66D6444FB93FC9EFBAC" box="[776,844,1078,1097]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="13" pageNumber="202" unit="m" value="2600.0">2600 m</elevation>
</quantity>
alt., 
<date id="056D1034740DE66D6430FB93FC17FBAC" box="[892,965,1078,1097]" pageId="13" pageNumber="202" value="1987-05-01">
<collectingDate id="1529E9DC740DE66D6430FB93FC17FBAC" box="[892,965,1078,1097]" pageId="13" pageNumber="202" value="1987-05-01">1.v.1987</collectingDate>
</date>
leg. 
<collectorName id="DC265322740DE66D64B8FB93FB8CFBAC" box="[1012,1118,1078,1097]" pageId="13" pageNumber="202">Burckhardt</collectorName>
and 
<collectorName id="DC265322740DE66D678DFBF7FF3EFB80" box="[193,236,1106,1125]" pageId="13" pageNumber="202">Löbl</collectorName>
(
<collectionCode id="17C2AE31740DE66D67BBFBF7FE91FB80" box="[247,323,1106,1125]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="13" pageNumber="202" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="C1BB3CA9740DE66D661DFBF7FCD1FB80" ID-GBIF-Occurrence="2512737968" box="[337,771,1105,1125]" collectingDate="1987-05-01" collectionCode="CMPR" collectorName="Burckhardt &amp; Lobl" country="Malaysia" elevation="2600" location="Mount Kinabalu" pageId="13" pageNumber="202" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<typeStatus id="AE688856740DE66D661DFBF7FE7AFB80" box="[337,424,1106,1125]" pageId="13" pageNumber="202" type="paratype">Paratype</typeStatus>
: 
<specimenCount id="67D5FD7D740DE66D66FDFBF7FE01FB80" box="[433,467,1105,1125]" pageId="13" pageNumber="202" type="male">1 ♂</specimenCount>
: same data as 
<typeStatus id="AE688856740DE66D6517FBF7FD7DFB80" box="[603,687,1106,1125]" pageId="13" pageNumber="202" type="holotype">holotype</typeStatus>
(
<collectionCode id="17C2AE31740DE66D65F6FBF7FD2FFB80" box="[698,765,1106,1125]" country="India" name="Centre for Medicinal Plants Research" pageId="13" pageNumber="202" type="Herbarium">CMPR</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="39C9657F740DE66D678DFB28FE7EFAC7" pageId="13" pageNumber="202" type="diagnosis">
<paragraph id="716C36F4740DE66D678DFB28FE7EFAC7" blockId="13.[151,1119,1165,1659]" pageId="13" pageNumber="202">
Diagnosis.— Habitus slender, ovoid. Uniformly light brown. Eyes reduced. Elytra laterally not curved, narrowed caudad in an approximately straight line. Male with a large and deep semicircular depression extended on the 5 
<superScript id="86A69BBC740DE66D6466FB6EFCE5FB3C" attach="left" box="[810,823,1227,1241]" fontSize="6" pageId="13" pageNumber="202">th</superScript>
and 6 
<superScript id="86A69BBC740DE66D6433FB6EFC5EFB3C" attach="left" box="[895,908,1227,1241]" fontSize="6" pageId="13" pageNumber="202">th</superScript>
visible abdominal sternites, bordered on the front half with long and dense hairs, and a central notch on the apical edge of the 6 
<superScript id="86A69BBC740DE66D66D4FAACFE77FAF2" attach="none" box="[408,421,1289,1303]" fontSize="6" pageId="13" pageNumber="202">th</superScript>
.
</paragraph>
</subSubSection>
<subSubSection id="39C9657F740DE66D678DFA8FFDC0F99F" pageId="13" pageNumber="202" type="description">
<paragraph id="716C36F4740DE66D678DFA8FFDC0F99F" blockId="13.[151,1119,1165,1659]" pageId="13" pageNumber="202">
Description.— Habitus slender, ovoid, uniformly light brown. Length: 
<quantity id="B62B9B11740DE66D6482FA8CFBF0FAA4" box="[974,1058,1321,1345]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="202" unit="mm" value="2.5">2.5 mm</quantity>
. Eyes reduced, each only 0.1 times as wide as the width of the head. Antennae slender, 1.25 times longer than the width of the head. Profemur as wide as the protibia. Pronotum 1.7 times as wide as long, with caudal corners slightly drawn out. Elytra slender, 1.25 times as long (measured from the caudal tip of the scutellum) as their combined width, the widest part at the base, the sides very slightly curved, rectilinearly narrowed from base to apex. Apterous. Male with a large and deep semicircular depression extended on the 5 
<superScript id="86A69BBC740DE66D67E8F9A1FF63F9F7" attach="left" box="[164,177,1540,1554]" fontSize="6" pageId="13" pageNumber="202">th</superScript>
and 6 
<superScript id="86A69BBC740DE66D67B4F9A1FED7F9F7" attach="left" box="[248,261,1540,1554]" fontSize="6" pageId="13" pageNumber="202">th</superScript>
visible abdominal sternites, bordered on the front half with long and dense hairs, and a central notch on the apical edge of the 6 
<superScript id="86A69BBC740DE66D6593F986FD3EF9D4" attach="none" box="[735,748,1571,1585]" fontSize="6" pageId="13" pageNumber="202">th</superScript>
. Aedeagus (
<figureCitation id="E9E82A71740DE66D643AF981FC15F9D9" box="[886,967,1572,1596]" captionStart="Figs 7-12" captionStartId="17.[151,190,1413,1432]" captionTargetId="figure@18.[493,888,185,983]" captionTargetPageId="18" captionText="Figs 7-12. Fig. 7, Ptomaphaginus burckhardti spec. nov., holotype; aedeagus, dorsal view. Fig. 8, P. burckhardti spec. nov., holotype; aedeagus, lateral view. Fig. 9, P. kurbatovi spec. nov., paratype (Cibodas); aedeagus, dorsal view. Fig. 10, P. kurbatovi spec. nov., paratype (Cibodas); aedeagus, lateral view. Fig. 11, P. sabahensis spec. nov., holotype; aedeagus, dorsal view. Fig. 12, P. sabahensis spec. nov., holotype; aedeagus, lateral view. All scale lines 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3576291" httpUri="https://zenodo.org/record/3576291/files/figure.png" pageId="13" pageNumber="202">figs 7-8</figureCitation>
) slender. The apex is tapered terminally and ends in a flattened, duck-bill-shaped processus. It carries several long, curved, lateral setae.
</paragraph>
</subSubSection>
<subSubSection id="39C9657F740EE66E678DFF65FED7FF1D" pageId="14" pageNumber="203" type="distribution">
<paragraph id="716C36F4740EE66E678DFF65FED7FF1D" blockId="14.[151,1119,192,408]" pageId="14" pageNumber="203">Distribution.— The single known specimen was found at high altitude on Mount Kinabalu.</paragraph>
</subSubSection>
<subSubSection id="39C9657F740EE66E678DFEA5FC1AFE9D" pageId="14" pageNumber="203" type="discussion">
<paragraph id="716C36F4740EE66E678DFEA5FC1AFE9D" blockId="14.[151,1119,192,408]" pageId="14" pageNumber="203">
Remarks.— At first glance, the species is similar to a 
<taxonomicName id="B6D34D77740EE66E6441FEA5FC66FEFD" authorityName="Perreau" authorityYear="2000" box="[781,948,256,280]" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="43A7EAE6740EE66E6441FEA5FC66FEFD" box="[781,948,256,280]" italics="true" pageId="14" pageNumber="203">Ptomaphaminus</emphasis>
</taxonomicName>
, because of the small size, the lack of pigmentation and reduced eyes, which suggest an adaptation to a deep montane litter biotope (as does the absence of wings). However, since it lacks the diagnostic thoracal characters of this genus, it is placed in 
<taxonomicName id="B6D34D77740EE66E646DFEC5FC13FE9D" authorityName="Bryanti Jeannel" authorityYear="1936" box="[801,961,352,376]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="43A7EAE6740EE66E646DFEC5FC13FE9D" box="[801,961,352,376]" italics="true" pageId="14" pageNumber="203">Ptomaphaginus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="39C9657F740EE66E678DFE25FBF8FE7D" box="[193,1066,384,408]" pageId="14" pageNumber="203" type="etymology">
<paragraph id="716C36F4740EE66E678DFE25FBF8FE7D" blockId="14.[151,1119,192,408]" box="[193,1066,384,408]" pageId="14" pageNumber="203">Etymology.— Dedicated to Daniel Burckhardt, who first collected this species.</paragraph>
</subSubSection>
</treatment>
</document>