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<mods:title id="13092566427A1B401404BE5F853BB048">A revision of Centrioncus Speiser (Diptera, Diopsidae, Centrioncinae) with descriptions of new species from Angola, Burundi, and Kenya</mods:title>
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<mods:namePart id="2B5BBAB6141636AFC35F283DDBC6BDA7">Feijen, Hans R.</mods:namePart>
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<mods:namePart id="B5D7226EB2F3972EBCF0F8759579056B">Feijen, Cobi</mods:namePart>
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<treatment id="F9B5D85A82435D5CA4D159144E3A9F01" ID-GBIF-Taxon="209152256" LSID="urn:lsid:plazi:treatment:F9B5D85A82435D5CA4D159144E3A9F01" httpUri="http://treatment.plazi.org/id/F9B5D85A82435D5CA4D159144E3A9F01" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Diopsidae" scope_order="Diptera" scope_subFamily="Centrioncinae">
<subSubSection id="002C556AFD2F76F0C10A3C28109D2580" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="7AF42877B504CF0FD61699F93E1A1FE2" pageId="0" pageNumber="1">
<taxonomicName id="20B65191C7A3134D71FFEF1C606C6F31" ID-CoL="S93G" LSID="F9B5D85A-8243-5D5C-A4D1-59144E3A9F01" authority="Feijen" authorityName="Feijen" authorityYear="1983" class="Insecta" family="Diopsidae" genus="Centrioncus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Centrioncus decellei" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="decellei">Centrioncus decellei Feijen</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="F33A47067382707A88EAC8B7DD40AAC4" pageId="0" pageNumber="1" type="description">
<paragraph id="4EF48E99289428C41E95005F93122B05" pageId="0" pageNumber="1">
<figureCitation id="1DCFA32B11449411702AEA431AAE25E1" captionStart="Figures 110, 111" captionStartId="F28" captionText="Figures 110, 111. Centrioncus decellei, ♀, holotype, Amanikro, Ivory Coast 110 wing, dorsal view 111 sternite 7, ventral view. Scale bars: 0.5 mm (110); 0.1 mm (111). Both drawings were processed, based on original pencil drawings from Feijen (1983: figs 12, 26)." figureDoi="10.3897/zookeys.1144.95619.figures110-111" httpUri="https://binary.pensoft.net/fig/803941" pageId="0" pageNumber="1">Figs 110</figureCitation>
<figureCitation id="B0284AE73A0965E14755712A43AD7249" captionStart="Figures 110, 111" captionStartId="F28" captionText="Figures 110, 111. Centrioncus decellei, ♀, holotype, Amanikro, Ivory Coast 110 wing, dorsal view 111 sternite 7, ventral view. Scale bars: 0.5 mm (110); 0.1 mm (111). Both drawings were processed, based on original pencil drawings from Feijen (1983: figs 12, 26)." figureDoi="10.3897/zookeys.1144.95619.figures110-111" httpUri="https://binary.pensoft.net/fig/803941" pageId="0" pageNumber="1">, 111</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="37C009CB6EEB69823F0139A340BDC192" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="35FD3260D83178A0F669BDC2E471FFA8" pageId="0" pageNumber="1">
<taxonomicName id="795E2835947696A320752ED2436734CE" authorityName="Feijen" authorityYear="1983" class="Insecta" family="Diopsidae" genus="Centrioncus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Centrioncus decellei" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="decellei">Centrioncus decellei</taxonomicName>
Feijen, 1983: 73.
</paragraph>
</subSubSection>
<subSubSection id="4E194D2DB5200AEA715262129212D506" pageId="0" pageNumber="1" type="type material">
<paragraph id="895ABF328B1F400E57939C6382A70D62" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="68741E22BF22DBB24E8756D0C7F03FAD" pageId="0" pageNumber="1">
Ivory Coast, 
<emphasis id="8852685A7439BA2B580C1E3F78FC1CB9" bold="true" italics="true" pageId="0" pageNumber="1">holotype</emphasis>
♀, Amanikro, 50 km nw of Abengourou [
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, 
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, 52 km NW of Abengourou, 194 m], ix. 1961, J. Decelle (MRAC) (not examined, but notes and pencil drawings for 
<bibRefCitation id="45A85F3D9B4A2E51270965B11F2BB827" author="Feijen, HR" journalOrPublisher="Zoologische Verhandelingen Leiden" pageId="0" pageNumber="1" pagination="1 - 137" publicationUrl="http://www.repository.naturalis.nl/document/149145" refId="B12" refString="Feijen, HR, 1983. Systematics and phylogeny of Centrioncidae, a new Afromontane family of Diptera (Schizophora). Zoologische Verhandelingen Leiden 202: 1 - 137, http://www.repository.naturalis.nl/document/149145" title="Systematics and phylogeny of Centrioncidae, a new Afromontane family of Diptera (Schizophora)." url="http://www.repository.naturalis.nl/document/149145" volume="202" year="1983">Feijen (1983)</bibRefCitation>
were used).
</paragraph>
</subSubSection>
<subSubSection id="C1EC38E8CFF8A03E3B748915FC696480" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="1F2F9F8CCD4FD4FA23AF0140ACEC6659" pageId="0" pageNumber="1">Diagnosis</paragraph>
<paragraph id="45A9A31F721EB5682E35CF8DD8E0A832" pageId="0" pageNumber="1">
<emphasis id="D5F768BD2B31ED536E977017ABD1543D" bold="true" pageId="0" pageNumber="1">
(after 
<bibRefCitation id="54845057E898844ACEC853CFE7F5542B" author="Feijen, HR" journalOrPublisher="Zoologische Verhandelingen Leiden" pageId="0" pageNumber="1" pagination="1 - 137" publicationUrl="http://www.repository.naturalis.nl/document/149145" refId="B12" refString="Feijen, HR, 1983. Systematics and phylogeny of Centrioncidae, a new Afromontane family of Diptera (Schizophora). Zoologische Verhandelingen Leiden 202: 1 - 137, http://www.repository.naturalis.nl/document/149145" title="Systematics and phylogeny of Centrioncidae, a new Afromontane family of Diptera (Schizophora)." url="http://www.repository.naturalis.nl/document/149145" volume="202" year="1983">Feijen 1983</bibRefCitation>
).
</emphasis>
<taxonomicName id="BA5E3045850C299E838F34B16C14465F" authorityName="Feijen" authorityYear="1983" class="Insecta" family="Diopsidae" genus="Centrioncus" kingdom="Animalia" lsidName="Centrioncus decellei" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="decellei">
<emphasis id="BFC483A6850482BBD2714D23DBF4DC7D" italics="true" pageId="0" pageNumber="1">Centrioncus decellei</emphasis>
</taxonomicName>
can be recognised by the flat, uniformly pruinose frons; pruinose blackish brown scutum, brownish humeral calli, chestnut-brown lateral edges; scutellum blackish brown with brown edges and lateral sides, scutellar spines yellowish; blackish brown pleura, brown anterior part of anepisternum and posterior part of anepimeron, katepisternum with brown spot anteriorly; scutellum projecting at angle of 45° from body axis; pale brown fore femur on distal half of inner side with dark brown, stripe-like spot, with ~ 38 tubercles; small, elongate, very vague, brownish central spot around crossvein r-m (Fig. 
<figureCitation id="45EF36F58087AE6A4A7D68480DEE7890" captionStart="Figures 110, 111" captionStartId="F28" captionText="Figures 110, 111. Centrioncus decellei, ♀, holotype, Amanikro, Ivory Coast 110 wing, dorsal view 111 sternite 7, ventral view. Scale bars: 0.5 mm (110); 0.1 mm (111). Both drawings were processed, based on original pencil drawings from Feijen (1983: figs 12, 26)." figureDoi="10.3897/zookeys.1144.95619.figures110-111" httpUri="https://binary.pensoft.net/fig/803941" pageId="0" pageNumber="1">110</figureCitation>
), in tip of cell br, mainly in posterior part of basal quarter of cell r4+5, slightly extending into cell bm+dm; vein M4 distally of crossvein dm-m curving downward towards wing margin; tergites blackish brown, apical edges paler; female 7th spiracle in tergite (Fig. 
<figureCitation id="51C6390053C5438C7D2ECA5C9FA525A9" captionStart="Figures 110, 111" captionStartId="F28" captionText="Figures 110, 111. Centrioncus decellei, ♀, holotype, Amanikro, Ivory Coast 110 wing, dorsal view 111 sternite 7, ventral view. Scale bars: 0.5 mm (110); 0.1 mm (111). Both drawings were processed, based on original pencil drawings from Feijen (1983: figs 12, 26)." figureDoi="10.3897/zookeys.1144.95619.figures110-111" httpUri="https://binary.pensoft.net/fig/803941" pageId="0" pageNumber="1">111</figureCitation>
); female tergite 7 with small mesal gap at posterior edge; female tergite 10 with 3 pairs of setulae; trapezoidal anterior sclerite of female sternite 7 narrow (w/l ratio: ~ 2.7); posterior sclerite of female sternite 7 rounded U-shaped, relatively short lateral arms with tri-lobed posterior apices (Fig. 
<figureCitation id="29B3FF55288E9C7246B6A9483B1DABB7" captionStart="Figures 110, 111" captionStartId="F28" captionText="Figures 110, 111. Centrioncus decellei, ♀, holotype, Amanikro, Ivory Coast 110 wing, dorsal view 111 sternite 7, ventral view. Scale bars: 0.5 mm (110); 0.1 mm (111). Both drawings were processed, based on original pencil drawings from Feijen (1983: figs 12, 26)." figureDoi="10.3897/zookeys.1144.95619.figures110-111" httpUri="https://binary.pensoft.net/fig/803941" pageId="0" pageNumber="1">111</figureCitation>
); female cercus elongate, l/w ratio: 4.1; subanal plate triangular, with convex lateral sides; spermathecae wrinkled, with a dimple and some tiny tubercles.
</paragraph>
<caption id="16B820E1877051D3063D9BFEB12C3474" doi="10.3897/zookeys.1144.95619.figures110-111" httpUri="https://binary.pensoft.net/fig/803941" pageId="0" pageNumber="1" start="Figures 110, 111" startId="F28">
<paragraph id="9CCB76D9371E9C02AD0AB406FC74FE94" pageId="0" pageNumber="1">
<emphasis id="EB13CF0EAF59039D8D98851E2CB8F76C" bold="true" pageId="0" pageNumber="1">Figures 110, 111.</emphasis>
<taxonomicName id="582F59AC7429DE6C78DE10D1F87B0E20" authorityName="Feijen" authorityYear="1983" class="Insecta" family="Diopsidae" genus="Centrioncus" kingdom="Animalia" lsidName="Centrioncus decellei" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="decellei">
<emphasis id="F6F37335D028F544775F7B8A3027F722" italics="true" pageId="0" pageNumber="1">Centrioncus decellei</emphasis>
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, ♀, holotype, Amanikro, Ivory Coast 
<emphasis id="B8FBACBEA77024B7E18E87389D47584C" bold="true" pageId="0" pageNumber="1">110</emphasis>
wing, dorsal view 
<emphasis id="E27E1D1654859B9A0529BD8FB92FEFB9" bold="true" pageId="0" pageNumber="1">111</emphasis>
sternite 7, ventral view. Scale bars: 0.5 mm (
<emphasis id="B0DC53052AC64CD1DF5052A5D115F32F" bold="true" pageId="0" pageNumber="1">110</emphasis>
); 0.1 mm (
<emphasis id="E6BA1460A59518F731F0B90019F86D7E" bold="true" pageId="0" pageNumber="1">111</emphasis>
). Both drawings were processed, based on original pencil drawings from 
<bibRefCitation id="90EFD8FD5365547AA2545D021034BC6F" author="Feijen, HR" journalOrPublisher="Zoologische Verhandelingen Leiden" pageId="0" pageNumber="1" pagination="1 - 137" publicationUrl="http://www.repository.naturalis.nl/document/149145" refId="B12" refString="Feijen, HR, 1983. Systematics and phylogeny of Centrioncidae, a new Afromontane family of Diptera (Schizophora). Zoologische Verhandelingen Leiden 202: 1 - 137, http://www.repository.naturalis.nl/document/149145" title="Systematics and phylogeny of Centrioncidae, a new Afromontane family of Diptera (Schizophora)." url="http://www.repository.naturalis.nl/document/149145" volume="202" year="1983">Feijen (1983</bibRefCitation>
: figs 12, 26).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="979EE2E9ABAB18088C4E0BAC44CAC8E9" pageId="0" pageNumber="1" type="description">
<paragraph id="81DCB26BCAC8670121B1DEF9D788E8E0" pageId="0" pageNumber="1">Supplementary description.</paragraph>
<paragraph id="E1D4E0B28ED223426A1A4E47B4062F10" pageId="0" pageNumber="1">
<emphasis id="DDBD2D47582F89065984968BB3A7A3D7" bold="true" pageId="0" pageNumber="1">
<emphasis id="D00E33A54173D8BBDE3FAB4A1B0D41B3" italics="true" pageId="0" pageNumber="1">Wing</emphasis>
.
</emphasis>
Small, elongate, very vague, brownish central spot around crossvein r-m (Fig. 
<figureCitation id="07CACF81FA00E7F97927469285B7972B" captionStart="Figures 110, 111" captionStartId="F28" captionText="Figures 110, 111. Centrioncus decellei, ♀, holotype, Amanikro, Ivory Coast 110 wing, dorsal view 111 sternite 7, ventral view. Scale bars: 0.5 mm (110); 0.1 mm (111). Both drawings were processed, based on original pencil drawings from Feijen (1983: figs 12, 26)." figureDoi="10.3897/zookeys.1144.95619.figures110-111" httpUri="https://binary.pensoft.net/fig/803941" pageId="0" pageNumber="1">110</figureCitation>
), in tip of cell br, mainly in posterior part of basal quarter of cell r4+5 to three-quarters distance to crossvein dm-m, slightly extending into cell bm+dm; vein CuA+CuP from vein CuP onward extending under angle of 25° to wing margin in straight line; vein M4, especially distal of crossvein dm-m, curving downward to wing margin (Fig. 
<figureCitation id="47D8AFAA8F388E03AD4B7B6636BD9F37" captionStart="Figures 110, 111" captionStartId="F28" captionText="Figures 110, 111. Centrioncus decellei, ♀, holotype, Amanikro, Ivory Coast 110 wing, dorsal view 111 sternite 7, ventral view. Scale bars: 0.5 mm (110); 0.1 mm (111). Both drawings were processed, based on original pencil drawings from Feijen (1983: figs 12, 26)." figureDoi="10.3897/zookeys.1144.95619.figures110-111" httpUri="https://binary.pensoft.net/fig/803941" pageId="0" pageNumber="1">110</figureCitation>
).
</paragraph>
<paragraph id="C2A6F89C586B43865654635FF1CCB868" pageId="0" pageNumber="1">
<emphasis id="F5A3210DE9AFD86CBF5F1D80F3F1A25B" bold="true" pageId="0" pageNumber="1">
<emphasis id="F64F64276A56D602262459635FC51042" italics="true" pageId="0" pageNumber="1">Female postabdomen</emphasis>
.
</emphasis>
Anterior sclerite of sternite 7 trapezoidal with w/l ratio: ~ 2.7 (Fig. 
<figureCitation id="78E03E37EBB9FECD308517517505AE95" captionStart="Figures 110, 111" captionStartId="F28" captionText="Figures 110, 111. Centrioncus decellei, ♀, holotype, Amanikro, Ivory Coast 110 wing, dorsal view 111 sternite 7, ventral view. Scale bars: 0.5 mm (110); 0.1 mm (111). Both drawings were processed, based on original pencil drawings from Feijen (1983: figs 12, 26)." figureDoi="10.3897/zookeys.1144.95619.figures110-111" httpUri="https://binary.pensoft.net/fig/803941" pageId="0" pageNumber="1">111</figureCitation>
, Table 
<tableCitation id="16BA69F8658360BFBCFA1AF237B77128" captionStart="Table 8" captionStartId="T8" captionText="Table 8. Ratio width / length for the anterior sclerite of ♀ sternite 7, ratio length / width for ♀ cercus and ratio length / greatest width for ♂ cercus. Species are arranged according to three species-groups recognised." httpUri="http://table.plazi.org/id/BA654C5FE474EF511E0FDD042BFC63BE" pageId="0" pageNumber="1" tableUuid="BA654C5FE474EF511E0FDD042BFC63BE">8</tableCitation>
); posterior sclerite of sternite 7 rounded U-shaped, with relatively short lateral arms, with 3-4-lobed posterior apices (Fig. 
<figureCitation id="214DBDC1CCB499B36067CB8CBF5C9E34" captionStart="Figures 110, 111" captionStartId="F28" captionText="Figures 110, 111. Centrioncus decellei, ♀, holotype, Amanikro, Ivory Coast 110 wing, dorsal view 111 sternite 7, ventral view. Scale bars: 0.5 mm (110); 0.1 mm (111). Both drawings were processed, based on original pencil drawings from Feijen (1983: figs 12, 26)." figureDoi="10.3897/zookeys.1144.95619.figures110-111" httpUri="https://binary.pensoft.net/fig/803941" pageId="0" pageNumber="1">111</figureCitation>
); female cercus elongate with l/w ratio: 4.1 (Table 
<tableCitation id="9C06194C5307415AD64CE21B42D35221" captionStart="Table 8" captionStartId="T8" captionText="Table 8. Ratio width / length for the anterior sclerite of ♀ sternite 7, ratio length / width for ♀ cercus and ratio length / greatest width for ♂ cercus. Species are arranged according to three species-groups recognised." httpUri="http://table.plazi.org/id/BA654C5FE474EF511E0FDD042BFC63BE" pageId="0" pageNumber="1" tableUuid="BA654C5FE474EF511E0FDD042BFC63BE">8</tableCitation>
); subanal plate triangular with convex lateral sides.
</paragraph>
</subSubSection>
<subSubSection id="CE604DD23C74E707A26E52A319EC2721" pageId="0" pageNumber="1" type="distribution">
<paragraph id="3D5E8BB0E0FAB4FD2554539259D342CB" pageId="0" pageNumber="1">Distribution and habitat.</paragraph>
<paragraph id="C1D769DA60F1962E6F44431565CE4B68" pageId="0" pageNumber="1">
There are several Amanikro locations in Ivory Coast, three of which are north-west of Abengourou. However, given the label distance of 50 km from Abengourou, the correct collecting location must be Amanikro at 
<geoCoordinate id="371B02C29CEF2CFCBC7ADC88E59E2CBE" degrees="7" direction="north" minutes="1" orientation="latitude" precision="15" seconds="0" value="7.016667">7°1'0&quot;N</geoCoordinate>
, 
<geoCoordinate id="9E546657D53F844EDF7795094A69BA72" degrees="3" direction="west" minutes="52" orientation="longitude" precision="15" seconds="0" value="-3.8666668">3°52'0&quot;W</geoCoordinate>
, 52 km NW of Abengourou, 194 m. The holotype was collected in lowland forest. This species is the only 
<taxonomicName id="67CBF3D06EC4E4FB3287FE6A1957DD3B" authorityName="Speiser" authorityYear="1910" class="Insecta" family="Diopsidae" genus="Centrioncus" kingdom="Animalia" lsidName="Centrioncus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="766C4B9AEEC9119438B6765BF9D32A6B" italics="true" pageId="0" pageNumber="1">Centrioncus</emphasis>
</taxonomicName>
known to occur at a low altitude, while the single specimen is also the only 
<taxonomicName id="17F38C6C38704D7E16728792E4A69CEC" authorityName="Speiser" authorityYear="1910" class="Insecta" family="Diopsidae" genus="Centrioncus" kingdom="Animalia" lsidName="Centrioncus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="E5364C2870F6FE10949163B314197A4E" italics="true" pageId="0" pageNumber="1">Centrioncus</emphasis>
</taxonomicName>
known from West Africa.
</paragraph>
</subSubSection>
</treatment>
</document>