<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.90.20586" ID-PMC="PMC5784240" ID-Pensoft-Pub="1314-2003-90-1" ID-Pensoft-UUID="80067D29FFFB7D34FF80E95D553F4254" ID-PubMed="29391851" ID-Zenodo-Dep="1138341" ModsDocID="1314-2003-90-1" checkinTime="1510830429569" checkinUser="pensoft" docAuthor="Berry, Paul E., Kainulainen, Kent &amp; Ee, Benjamin W. van" docDate="2017" docId="FCFAA8DB887252E2B0A07BE2D7C0A45E" docLanguage="en" docName="PhytoKeys 90: 1-87" docOrigin="PhytoKeys 90" docPubDate="2017-11-15" docSource="http://dx.doi.org/10.3897/phytokeys.90.20586" docTitle="Croton nudatus Baill., Adansonia 1: 168. 1861, as ‘ nudatum’" docType="treatment" docVersion="3" id="80067D29FFFB7D34FF80E95D553F4254" lastPageNumber="54" masterDocId="80067D29FFFB7D34FF80E95D553F4254" masterDocTitle="A Nomenclator of Croton (Euphorbiaceae) in Madagascar, the Comoros Archipelago, and the Mascarene Islands" masterLastPageNumber="87" masterPageNumber="1" pageNumber="53" updateTime="1668142129654" updateUser="ExternalLinkService">
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<mods:title id="BE18E3B26115967B743154D07829F28A">A Nomenclator of Croton (Euphorbiaceae) in Madagascar, the Comoros Archipelago, and the Mascarene Islands</mods:title>
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<mods:namePart id="BFA390813D91E8A24377D42A380F83F0">Berry, Paul E.</mods:namePart>
<mods:affiliation id="4D1FD92D645C49A5EE31CDA360452B66">Herbarium, Department of Ecology and Evolutionary Biology, University of Michigan, 3600 Varsity Drive, Ann Arbor, Michigan 48108, U. S. A.</mods:affiliation>
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<mods:namePart id="DE37F9510A798A514B5D3AB52F20B204">Kainulainen, Kent</mods:namePart>
<mods:affiliation id="EF87E9AEC839F7A7BB4A4D8B13E78C6A">Herbarium, Department of Ecology and Evolutionary Biology, University of Michigan, 3600 Varsity Drive, Ann Arbor, Michigan 48108, U. S. A.</mods:affiliation>
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<mods:namePart id="88A85D20C1B7EED0EBFEB93C93DD1EE9">Ee, Benjamin W. van</mods:namePart>
<mods:affiliation id="0BC1359453E1983C4FBEC567C0C5FFD7">Department of Biology, Universidad de Puerto Rico, Recinto Universitario de Mayagueez, Mayagueez, PR 00680, Puerto Rico, U. S. A.</mods:affiliation>
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<paragraph id="29349286629490A6049A375DD8933A44" pageId="52" pageNumber="53">
102. 
<taxonomicName id="B0BD9A1B2C0EFB607AAF1BD4E0FFDDA0" LSID="FCFAA8DB-8872-52E2-B0A0-7BE2D7C0A45E" authority="Baill., Adansonia 1: 168. 1861, as ' nudatum'" authorityName="Baill., Adansonia 1: 168. 1861, as ‘ nudatum’" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Croton nudatus" order="Malpighiales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="nudatus">
Croton nudatus Baill., Adansonia 1: 168. 1861, as 
<normalizedToken id="FA993A5B191089B527239FDDC9349F69" originalValue="‘nudatum’">'nudatum'</normalizedToken>
</taxonomicName>
</paragraph>
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<subSubSection id="2A42AA94253DCA73E7A2F8590C08DD82" pageId="52" pageNumber="53" type="reference_group">
<paragraph id="5F453C3E801E25D99D4B55FC95A5C549" pageId="52" pageNumber="53">
<taxonomicName id="878FA96F3983CB1F7AD4944998E34487" authorityName="Kuntze" authorityYear="1891" baseAuthorityName="Baill." class="Magnoliopsida" family="Euphorbiaceae" genus="Oxydectes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Oxydectes nudata" order="Malpighiales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="nudata">Oxydectes nudata</taxonomicName>
(Baill.) Kuntze, Revis. Gen. Pl. 2: 612. 1891. Type. Based on 
<taxonomicName id="600A4B0E6EE49994755DCAC5DF0459AA" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Croton nudatus" order="Malpighiales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="nudatus">Croton nudatus</taxonomicName>
Baill.
</paragraph>
<paragraph id="386CEE8B90C5081FEA88EBE361F28E08" pageId="52" pageNumber="53">
<taxonomicName id="E04DF7A8CF2DEC991632CC645997F087" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Croton boivinanus subsp. var. var. brevifolius" order="Malpighiales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="variety" species="boivinanus" subSpecies="var." variety="brevifolius">Croton boivinanus var. brevifolius</taxonomicName>
Radcl.-Sm., Gen. Croton Madag. Comoro 12. 2016. Type. Madagascar. Prov. Antsiranana: Fivonandra Antsiranana II, 79 km au Sud 
<normalizedToken id="4F9370CFE9850838226C4BAA905DD1B2" originalValue="d’Antsiranana">d'Antsiranana</normalizedToken>
par route RN6, et 15 km 
<normalizedToken id="7771C3F8D94084DB1951CC38CBDAAD3A" originalValue="à">a</normalizedToken>
<normalizedToken id="4565D660BE54BF4C6C77C1367183A535" originalValue="l’Est">l'Est</normalizedToken>
de 
<normalizedToken id="D0601CC5102117EDB02BB5808FDFC0FC" originalValue="l’ancien">l'ancien</normalizedToken>
chantier du Colas 
<normalizedToken id="24880B41C2577E40F02172C15E886B61" originalValue="à">a</normalizedToken>
Marotaolana, Campement 
<normalizedToken id="F7992470085A04ECF3A6A639943D9C99" originalValue="à">a</normalizedToken>
<normalizedToken id="21417D9A984ACE8F203F50498C1BE95E" originalValue="l’Est">l'Est</normalizedToken>
du village 
<normalizedToken id="0484E237EB2A8F9A891F43A6CD8ADF56" originalValue="d’Ampantsona">d'Ampantsona</normalizedToken>
, 
<geoCoordinate id="236109BCA7DF50F20D10463F9BFEDF1C" degrees="12" direction="south" minutes="51" orientation="latitude" precision="15" seconds="20" value="-12.855556">12°51'20&quot;S</geoCoordinate>
, 
<geoCoordinate id="A66059D823260B230ED5741843CDEC33" degrees="49" direction="east" minutes="18" orientation="longitude" precision="15" seconds="10" value="49.302776">49°18'10&quot;E</geoCoordinate>
, 394-551 m, 3-6 Jun 1997, 
<emphasis id="04F9313A62928B711C844884F3B135AF" italics="true" pageId="52" pageNumber="53">O. Andrianantoanina &amp; R. Bezara 1068</emphasis>
(holotype: K!; isotypes: G!, MO!, P [P00433267]!).
</paragraph>
<paragraph id="CCF225F37A65C8701C6E84A9B97AA4D8" pageId="52" pageNumber="53">
<taxonomicName id="A465E2662677E7846E485835203F7A5C" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Croton hirsutissimus" order="Malpighiales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="hirsutissimus">Croton hirsutissimus</taxonomicName>
Radcl.-Sm., Gen. Croton Madag. Comoro 99. 2016. Type. Madagascar. Prov. Antsiranana: versant Est du Massif de 
<normalizedToken id="CBC2004965EA52E471FBEC749CC45BE5" originalValue="l’Ankerana">l'Ankerana</normalizedToken>
(partie S du Massif de Mafokovo, au N de 
<normalizedToken id="54EDEE614D5C87C90C0CB297542437FE" originalValue="Vohémar">Vohemar</normalizedToken>
), 50-400 m, 17 Dec 1966, 
<emphasis id="331980E470BB63B415CFCAB4CE41F326" italics="true" pageId="52" pageNumber="53">Service Forestier 27363</emphasis>
- 
<emphasis id="68D5ED9AA536C970EBA996691273C321" italics="true" pageId="52" pageNumber="53">SF</emphasis>
(holotype: P [P00154439]!).
</paragraph>
<paragraph id="CDBF66B5BB33C1D2336EE6C29D669B0D" pageId="52" pageNumber="53">
<taxonomicName id="F567812C6BFEB6987D2E0642558081E0" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Croton menabeensis subsp. var. var. furfuraceus" order="Malpighiales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="variety" species="menabeensis" subSpecies="var." variety="furfuraceus">Croton menabeensis var. furfuraceus</taxonomicName>
Radcl.-Sm., Gen. Croton Madag. Comoro 101. 2016. Type. Madagascar. Prov. Mahajanga: Sofia Region, Antsohihy District, Antsatrana, Bora, 17 Jul 1970, 
<emphasis id="D705CA1EF9C27F444B5F217B8B4AF8C8" italics="true" pageId="52" pageNumber="53">Service Forestier 30011</emphasis>
(holotype: P [P00154438]!).
</paragraph>
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<paragraph id="2E41BED9AAAD700F6806587713FECF96" pageId="52" pageNumber="53">
<typeStatus id="8096505B56FF403227123B758CFAF5B5">Type</typeStatus>
.
</paragraph>
<paragraph id="0F9DD3970A1647C60D626DAA9E23DEA4" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<materialsCitation id="5F4D38DDAA0C3AF038E3B75CE137986E" collectingDate="1848-12" collectorName="de Diego-Suares, L. H. Boivin" country="Madagascar" location="Baie de Diego-Suares" municipality="Diana Region" specimenCount="1" stateProvince="Antsiranana" typeStatus="holotype">
<collectingCountry id="957A02CC5C46F3371F9D59593C88EA29" name="Madagascar">Madagascar</collectingCountry>
. Prov. 
<collectingRegion id="E0ECA3B80E77261CF4E90981CD0D56E2" country="Madagascar" name="Antsiranana">Antsiranana</collectingRegion>
: 
<collectingMunicipality id="29178F5740EF10CFD61D7D60B1C81754">Diana Region</collectingMunicipality>
, 
<location id="160F15241C0040619D3665ACE6496D34" LSID="urn:lsid:plazi:treatment:FCFAA8DB887252E2B0A07BE2D7C0A45E:160F15241C0040619D3665ACE6496D34" country="Madagascar" municipality="Diana Region" name="Baie de Diego-Suares" stateProvince="Antsiranana">
Baie 
<collectorName id="A061389D8253DA12D1A62A6A26D5593D">
de 
<normalizedToken id="EFFCEB53E553A1ED7C538DF96894B88E" originalValue="Diego-Suarès">Diego-Suares</normalizedToken>
</collectorName>
</location>
, 
<collectingDate id="E00512A95C7D7500B34B090863DE25B1" value="1848-12">Dec 1848</collectingDate>
, 
<emphasis id="D1B8AF654BEFF6D2BCD51C672BD94817" italics="true" pageId="52" pageNumber="53">
<collectorName id="5607773175B5410D77E5B8A7A4A17FCE">L.H. Boivin</collectorName>
2659
</emphasis>
(
<typeStatus id="A2BB534A4B9E56F8F62678477B845E47">holotype</typeStatus>
: P [P00389498]!)
</materialsCitation>
. 
<materialsCitation id="9EBE1E368AB7F899C9D2483B80C3F936" collectingDate="2009-10-25" collectorName="B. van Ee, P. E. Berry, K. J. Wurdack, E. A. Haber, H. Razafindraibe" country="Madagascar" county="Diana Region" elevation="50" latitude="-12.235556" location="Orangea" longLatPrecision="21" longitude="49.36111" municipality="Orangea" specimenCount="1" stateProvince="Antsiranana" typeStatus="holotype">
<collectingCountry id="09D367FEE2899674FE8B8213EADDEEE4" name="Madagascar">Madagascar</collectingCountry>
. Prov. 
<collectingRegion id="A9AD0B1A88C27A72E7EC0133FFF35549" country="Madagascar" name="Antsiranana">Antsiranana</collectingRegion>
: 
<collectingCounty id="30E47479F0AE1B769443143D4B8B8B72">
<pageBreakToken id="12CD033EA1B5C5CAAA6AFB5B1A5151F5" pageId="53" pageNumber="54" start="start">Diana</pageBreakToken>
Region
</collectingCounty>
, 
<collectingRegion id="35157605BF9ABA3621B6D2A5EDFF9F4F" country="Madagascar" name="Antsiranana">Antsiranana</collectingRegion>
II, 
<collectingMunicipality id="B506FA3BFCF29E062583727B7E608F92">Orangea</collectingMunicipality>
, road going uphill from military checkpoint at entrance to 
<location id="B223EDE106A9129747B64EADA9B6A192" LSID="urn:lsid:plazi:treatment:FCFAA8DB887252E2B0A07BE2D7C0A45E:B223EDE106A9129747B64EADA9B6A192" country="Madagascar" county="Diana Region" latitude="-12.235556" longLatPrecision="21" longitude="49.36111" municipality="Orangea" name="Orangea" stateProvince="Antsiranana">Orangea</location>
, 
<geoCoordinate id="EBD3ABEF54A53DCBB0DEBBCF3B68F4E3" degrees="12" direction="south" minutes="14" orientation="latitude" precision="15" seconds="08" value="-12.235556">12°14'08&quot;S</geoCoordinate>
, 
<geoCoordinate id="95D45313021B0E9EC504E52A7A0808D8" degrees="49" direction="east" minutes="21" orientation="longitude" precision="15" seconds="40" value="49.36111">49°21'40&quot;E</geoCoordinate>
, 
<quantity id="E6238F0B75BFDB2F6B412B8354DD4510" metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">
<elevation id="128DD56839472247B5652D73236BBA04" metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">50 m</elevation>
</quantity>
, 
<collectingDate id="02E54EAFFD5C085FA07DA459BE2A7576" value="2009-10-25">25 Oct 2009</collectingDate>
, 
<emphasis id="C28818D7C8034ABD654B4D55740D107F" italics="true" pageId="53" pageNumber="54">
<collectorName id="EE62F760426BFE75CD96A3D85DDADF1E">B. van Ee</collectorName>
, 
<collectorName id="FAF6912361ED4E29F6B9019507CE0D8A">P.E. Berry</collectorName>
, 
<collectorName id="BE494A4DFB8CB66FC7B9E1F51C9D044A">K.J. Wurdack</collectorName>
, 
<collectorName id="9E0845241E7269A0D4B6CBF66B171F7C">E.A. Haber</collectorName>
&amp; 
<collectorName id="768F977CF8AA081AA14338C30F50F2C4">H. Razafindraibe</collectorName>
1081
</emphasis>
(epitype, designated by 
<bibRefCitation id="93C3D85D3FCC422EA2688831DA104F29" DOI="https://doi.org/10.15553/c2017v722a15" author="Kainulainen, K" journalOrPublisher="Candollea" pageId="72" pageNumber="73" pagination="371 - 402" refId="B31" refString="Kainulainen, K, Van Ee, B, Razafindraibe, H, Berry, PE, 2017b. A revision of the Adenophorus Group and other glandular-leaved species of Croton (Euphorbiaceae) from northern Madagascar and Mayotte, including three new species. Candollea 72: 371 - 402, DOI: https://doi.org/10.15553/c2017v722a15" title="A revision of the Adenophorus Group and other glandular-leaved species of Croton (Euphorbiaceae) from northern Madagascar and Mayotte, including three new species." url="https://doi.org/10.15553/c2017v722a15" volume="72" year="2017 b">Kainulainen et al. 2017b</bibRefCitation>
, pg. 374: MICH [MICH1517189]!; additional duplicate: TAN)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0AC10BE5C07A0D6EFEAA49B4485C7497" pageId="53" pageNumber="54" type="distribution">
<paragraph id="3960A74CC665844148DDC671F624F317" pageId="53" pageNumber="54">Habit and distribution.</paragraph>
<paragraph id="0BDECB3A46C080BB6D63A6A66EBE4129" pageId="53" pageNumber="54">Shrubs; northern and western Madagascar (Antsiranana, Mahajanga).</paragraph>
</subSubSection>
</treatment>
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