Sticholotis
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publication ID |
https://doi.org/ 10.11646/zootaxa.4326.1.1 |
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publication LSID |
lsid:zoobank.org:pub:A2A72998-86A8-4E77-A2E3-64615Edb2D5F |
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DOI |
https://doi.org/10.5281/zenodo.5309686 |
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persistent identifier |
https://treatment.plazi.org/id/03A987ED-FFAA-FFE8-BDB6-EECA5EFAF8D6 |
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treatment provided by |
Plazi |
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scientific name |
Sticholotis |
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Key to the species of Sticholotis View in CoL View at ENA from China
1. Disc of elytra unicolored and without spots ( Figs 11–28 View FIGURES 11 – 19. 11 – 13 View FIGURES 20 – 28. 20 – 22 )...................................................... 2
- Disc of elytra bicolored or unicolored with black or red spots ( Figs 29–118 View FIGURES 29 – 37. 29 – 31 View FIGURES 38 – 46. 38 – 40 View FIGURES 47 – 55. 47 – 49 View FIGURES 56 – 64. 56 – 58 View FIGURES 65 – 73. 65 – 67 View FIGURES 74 – 82. 74 – 76 View FIGURES 83 – 91. 83 – 85 View FIGURES 92 – 100. 92 – 94 View FIGURES 101 – 109. 101 – 103 View FIGURES 110 – 118. 110 – 112 )....................................... 7
2. Elytra without dark lateral margins ( Figs 11–19 View FIGURES 11 – 19. 11 – 13 )............................................................. 3
- Elytra with dark lateral margins ( Figs 20–28 View FIGURES 20 – 28. 20 – 22 ) ................................................................ 5
3. Elytron covered by uniform fine punctures ( Figs 14–19 View FIGURES 11 – 19. 11 – 13 )....................................................... 4
- Elytron with row of large punctures along sutural and lateral margins ( Figs 11–12 View FIGURES 11 – 19. 11 – 13 ).................. S. vietnamica Hoàng
4. Body uniformly yellow ( Fig 14 View FIGURES 11 – 19. 11 – 13 ).............................................. S. simplifulva Wang & Ren sp. nov.
- Body uniformly brown ( Fig 17 View FIGURES 11 – 19. 11 – 13 )........................................................... S. hirashimai Sasaji
5. Elytron without dark sutural margin ( Figs 23–28 View FIGURES 20 – 28. 20 – 22 )............................................................ 6
- Elytron with dark sutural margin ( Fig. 20 View FIGURES 20 – 28. 20 – 22 ).................................................. S. carinica (Gorham)
6. Head and pronotum black. Elytron with disc yellow to red and lateral margin black ( Figs 23–25 View FIGURES 20 – 28. 20 – 22 )..... S. hoabinhensis Hoàng
- Dorsum uniformly reddish brown ( Figs 26–28 View FIGURES 20 – 28. 20 – 22 )................................ S. denticuligera Wang & Ren sp. nov.
7. Elytron brown to black, with yellow to red spots ( Figs 29–49 View FIGURES 29 – 37. 29 – 31 View FIGURES 38 – 46. 38 – 40 View FIGURES 47 – 55. 47 – 49 ).................................................. 8
- Elytron yellow to brown, with black spots ( Figs 50–118 View FIGURES 47 – 55. 47 – 49 View FIGURES 56 – 64. 56 – 58 View FIGURES 65 – 73. 65 – 67 View FIGURES 74 – 82. 74 – 76 View FIGURES 83 – 91. 83 – 85 View FIGURES 92 – 100. 92 – 94 View FIGURES 101 – 109. 101 – 103 View FIGURES 110 – 118. 110 – 112 )..................................................... 17
8. Elytron with an irregularly shaped macula ( Fig. 29 View FIGURES 29 – 37. 29 – 31 ).......................................................... 9
- Elytron with two rounded spots ( Figs 32–49 View FIGURES 29 – 37. 29 – 31 View FIGURES 38 – 46. 38 – 40 View FIGURES 47 – 55. 47 – 49 )............................................................... 10
9. Elytra with spots elongate-oval. TL: 2.0– 2.4 mm ............................................... S. bilineata Weise
- Elytra with irregular red patterns ( Fig. 29 View FIGURES 29 – 37. 29 – 31 ). TL: 2.27 mm ........................... S. flammula Wang & Ren sp. nov.
10. Elytra with anterior spot separated from anterior margin ( Figs 32–37 View FIGURES 29 – 37. 29 – 31 )........................................... 11
- Elytra with anterior spot meeting anterior margin ( Figs 38–47 View FIGURES 38 – 46. 38 – 40 View FIGURES 47 – 55. 47 – 49 )................................................. 14
11. Elytron with six longitudinal rows of large punctures......................................... S. confucii (Mulsant)
- Elytron with one longitudinal row of large punctures along the suture or without punctures ( Figs 32, 35 View FIGURES 29 – 37. 29 – 31 )............... 12
12. Elytra with red or brown external margins ( Figs 32, 35 View FIGURES 29 – 37. 29 – 31 )...................................................... 13
- Elytra without red or brown external margins................................................ S. morimotoi Kamiya
13. Pronotum red, elytron dark brown to black with two red spots ( Figs 32–33 View FIGURES 29 – 37. 29 – 31 ). Penis long and slender ( Fig. 155 View FIGURES 150 – 158. 150 – 153 ). TL: 2.24–2.87 mm, TW: 2.08–2.57 mm, TH: 1.05–1.15 mm ................................................. S. formosana Weise
- Pronotum black, elytron black with two dark red spots ( Figs 35–36 View FIGURES 29 – 37. 29 – 31 ). Penis short and stout ( Figs 160–161 View FIGURES 159 – 168. 159 – 163 ). TL: 2.27 mm, TW: 2.01 mm, TH: 1.09 mm .............................................................. S. taiwanensis Miyatake
14. Spots on elytra regularly shaped. Parameres slightly shorter than penis guide ( Figs 41–49 View FIGURES 38 – 46. 38 – 40 View FIGURES 47 – 55. 47 – 49 )........................... 15
- Spots on elytra irregularly shaped ( Figs 38–40 View FIGURES 38 – 46. 38 – 40 ). Parameres distinctly longer than penis guide ( Figs 166–167 View FIGURES 159 – 168. 159 – 163 ). TL: 2.18–2.27 mm, TW: 1.91–2.11 mm, TH: 1.01–1.05 mm ................................... S. wenchangia Wang & Ren sp. nov.
15. Penis relatively longer and slender ( Figs 175 View FIGURES 169 – 177. 169 – 173 , 179 View FIGURES 178 – 186. 178 – 182 ). Additional dorsal strut-like projection on tegminal basal piece large, almost same length to strut ( Figs 176 View FIGURES 169 – 177. 169 – 173 , 180 View FIGURES 178 – 186. 178 – 182 )...................................................................... 16
- Penis relatively shorter and stouter ( Fig. 171 View FIGURES 169 – 177. 169 – 173 ). Additional dorsal strut-like projection on tegminal basal piece small, half length of strut ( Figs 172 View FIGURES 169 – 177. 169 – 173 ). TL: 2.32–2.44 mm, TW: 1.98–2.11 mm, TH: 1.05–1.19 mm ..................... S. petila Yu & Pang
16. Elytral spots medium-sized, the distance between them larger than spot diameters ( Figs 44–46 View FIGURES 38 – 46. 38 – 40 ). TL: 1.6–2.1mm, TW: 1.3–1.8 mm, TH: 0.9–1.2 mm ..................................................................... S. ruficeps Weise
- Elytral spots very large, the distance between them about 1/3 of one diameter ( Figs 47–49 View FIGURES 47 – 55. 47 – 49 ). TL: 2.70–2.90 mm, TW: 2.51–2.67 mm, TH: 1.38–1.48 mm ................................................. S. dadugangensis Wang & Ren sp. nov.
17. Elytra with a dark band along anterior and lateral margins ( Figs 50–94 View FIGURES 47 – 55. 47 – 49 View FIGURES 56 – 64. 56 – 58 View FIGURES 65 – 73. 65 – 67 View FIGURES 74 – 82. 74 – 76 View FIGURES 83 – 91. 83 – 85 View FIGURES 92 – 100. 92 – 94 ).......................................... 18
- Elytra without continuous dark band along margins ( Figs 95–118 View FIGURES 92 – 100. 92 – 94 View FIGURES 101 – 109. 101 – 103 View FIGURES 110 – 118. 110 – 112 ).............................................. 33
18. Elytra with large central sutural spot ( Figs 50–55 View FIGURES 47 – 55. 47 – 49 )........................................................... 19
- Elytra with large central sutural spot and in addition a pair of rounded spots ( Figs 56–94 View FIGURES 56 – 64. 56 – 58 View FIGURES 65 – 73. 65 – 67 View FIGURES 74 – 82. 74 – 76 View FIGURES 83 – 91. 83 – 85 View FIGURES 92 – 100. 92 – 94 )........................... 20
19. Pronotum yellowish brown ( Fig. 50 View FIGURES 47 – 55. 47 – 49 ). Parameres distinctly dilated in apical third ( Figs 185–186 View FIGURES 178 – 186. 178 – 182 ). TL: 2.34 mm, TW: 2.14 mm, TH: 1.35 mm ................................................................ S. dilatata Wang & Ren sp. nov.
- Pronotum dark brown to black ( Figs 53–55 View FIGURES 47 – 55. 47 – 49 ). Parameres not dilated ( Figs 189–190 View FIGURES 187 – 196. 187 – 191 ). TL: 2.01–2.37 mm, TW: 1.85–2.11 mm, TH: 1.19–1.35 mm .................................................................... S. cinctipennis Weise
20. Pronotum black or yellow with black spot ( Figs 56–67 View FIGURES 56 – 64. 56 – 58 View FIGURES 65 – 73. 65 – 67 )...................................................... 21
- Pronotum uniformly yellow without spots ( Figs 68–94 View FIGURES 65 – 73. 65 – 67 View FIGURES 74 – 82. 74 – 76 View FIGURES 83 – 91. 83 – 85 View FIGURES 92 – 100. 92 – 94 )...................................................... 25
21. Pronotum black ( Figs 59–67 View FIGURES 56 – 64. 56 – 58 View FIGURES 65 – 73. 65 – 67 )........................................................................... 22
- Pronotum yellow with a black spot ( Figs 56, 58 View FIGURES 56 – 64. 56 – 58 ). TL: 2.24 mm, TW: 2.08 mm, TH: 1.19 mm .................................................................................................. S. illecebrosa Wang & Ren sp. nov.
22. Pronotum and elytra with very narrow yellow lateral margins. ( Figs 59–61 View FIGURES 56 – 64. 56 – 58 )...................................... 23
-. Pronotal and elytral lateral margins black ( Figs 62–67 View FIGURES 56 – 64. 56 – 58 View FIGURES 65 – 73. 65 – 67 )....................................................... 24
23. Apex of elytra with large quadrate black spot ( Fig. 59 View FIGURES 56 – 64. 56 – 58 ) TL: 2.10–2.49 mm, TW: 1.70–2.11 mm, TH: 1.15–1.25 mm ............................................................................................... S. punctata Crotch
- Apex of elytra without spots. TL: 2.0 mm.................................................. S. klapperichi Mader
24. Elytral disc with spots very large and elongate oval ( Fig. 62 View FIGURES 56 – 64. 56 – 58 ). Elytra covered with uniformly fine punctures ( Fig. 62 View FIGURES 56 – 64. 56 – 58 ). TL: 2.14–2.21 mm, TW: 1.81–1.91 mm, TH: 1.03–1.05 mm ........................ S. magnopunctata Wang & Ren sp. nov.
- Elytral disc with spots relatively small and rounded ( Fig. 65 View FIGURES 65 – 73. 65 – 67 ). Elytra covered with uniformly fine punctures and two rows of large punctures along margin of sutural spot ( Fig. 65 View FIGURES 65 – 73. 65 – 67 ). TL: 2.14 mm, TW: 1.88 mm, TH: 1.15 mm ............................................................................................... S. acutifolia Wang & Ren sp. nov.
25. Anterior elytral spot meeting the margin ( Figs 71–94 View FIGURES 65 – 73. 65 – 67 View FIGURES 74 – 82. 74 – 76 View FIGURES 83 – 91. 83 – 85 View FIGURES 92 – 100. 92 – 94 )........................................................ 26
- Anterior elytral spot not touching the basal elytral margin ( Fig. 68 View FIGURES 65 – 73. 65 – 67 ). TL: 2.11–2.47 mm, TW: 1.88–2.21 mm, TH: 1.15–1.32 mm ......................................................................... S. tortus Wang & Ren sp. nov.
26. Mid area of elytral suture darker but without spots ( Figs 71–76 View FIGURES 65 – 73. 65 – 67 View FIGURES 74 – 82. 74 – 76 )................................................ 27
- Mid area of elytral suture with a black spot ( Figs 77–94 View FIGURES 74 – 82. 74 – 76 View FIGURES 83 – 91. 83 – 85 View FIGURES 92 – 100. 92 – 94 )..................................................... 28
27. Pronotum yellow ( Fig 71, 73 View FIGURES 65 – 73. 65 – 67 ). Penis thick and penis guide extremely broad ( Figs 219–220 View FIGURES 217 – 226. 217 – 221 ). TL: 2.67–2.90 mm, TW: 2.37– 2.54 mm, TH: 1.19–1.38 mm .................................................. S. latissima Wang & Ren sp. nov.
- Pronotum dark red ( Fig. 74 View FIGURES 74 – 82. 74 – 76 ). Penis short and stout, penis guide thin ( Figs 224–225 View FIGURES 217 – 226. 217 – 221 ). TL: 2.24–2.97 mm, TW: 2.01–2.57 mm, TH: 1.19–1.62 mm ....................................................... S. maolanensis Wang & Ren sp. nov.
28. Parameres very short, less than 1/2 of penis guide length ( Figs 229, 234 View FIGURES 227 – 236. 227 – 231 )......................................... 29
- Parameres long, almost equal to penis guide ( Figs 239, 244 View FIGURES 237 – 246. 237 – 241 , 249, 253 View FIGURES 247 – 254. 247 – 250 )........................................... 30
29. Elytron disc and base with large and rounded spots ( Fig. 77 View FIGURES 74 – 82. 74 – 76 ). Penis very long and slender ( Fig. 228 View FIGURES 227 – 236. 227 – 231 ). Parameres slender ( Figs 229–230 View FIGURES 227 – 236. 227 – 231 ). TL: 2.08 mm, TW: 1.81 mm, TH: 1.05 mm ................................ S. tenuis Wang & Ren sp. nov.
- Elytron disc and base with small spots ( Fig. 80 View FIGURES 74 – 82. 74 – 76 ). Penis short ( Fig. 233 View FIGURES 227 – 236. 227 – 231 ). Parameres broad ( Fig. 234–235 View FIGURES 227 – 236. 227 – 231 ). TL: 2.27–2.54 mm, TW: 2.11–2.41 mm, TH: 1.22–1.38 mm ................................................... S. brachyloba sp. nov.
30. Pronotum dark red. Parameres slender ( Figs 86–94 View FIGURES 83 – 91. 83 – 85 View FIGURES 92 – 100. 92 – 94 )......................................................... 31
- Pronotum yellow ( Fig. 83 View FIGURES 83 – 91. 83 – 85 ). Parameres very broad ( Fig. 239 View FIGURES 237 – 246. 237 – 241 ). TL: 2.87 mm, TW: 2.60 mm, TH: 1.42 mm ..................................................................................................... S. denticulata Hoàng
31. Elytra with spots on humeral callus sligthly separated from basal margins ( Fig. 86 View FIGURES 83 – 91. 83 – 85 ). TL: 2.67–2.97 mm, TW: 2.54–2.84 mm, TH: 1.22–1.48 mm ............................................................. S. pengi Wang & Ren sp. nov.
- Elytra with spots on humeral callus that distinctly touch basal margins of elytra ( Figs 248, 252 View FIGURES 247 – 254. 247 – 250 )....................... 32
32. Elytron with spots on base and disc larger than sutural spots ( Fig. 89 View FIGURES 83 – 91. 83 – 85 ). TL: 2.80–2.90 mm, TW: 2.54–2.67 mm, TH: 1.46–1.48 mm ..................................................................... S. wuzhishana Wang & Ren sp. nov.
- Elytron with spots on base and disc distinctly smaller than sutural spots ( Fig. 92 View FIGURES 92 – 100. 92 – 94 ). TL: 2.44 mm, TW: 2.18 mm, TH: 1.29 mm ....................................................................... S. nanlingensis Wang & Ren sp. nov.
33. Elytron with 4 or less rounded or irregular compact black spots, although additional black streaks may be present on lateral or sutural margins ( Figs 95–106 View FIGURES 92 – 100. 92 – 94 View FIGURES 101 – 109. 101 – 103 )........................................................................... 34
- Elytron with more than 4 rounded or irregular compact black spots ( Figs 107–118 View FIGURES 101 – 109. 101 – 103 View FIGURES 110 – 118. 110 – 112 )................................. 40
34. Elytron with less than 4 black spots...................................................................... 35
- Elytron with 4 black spots ( Figs 95–106 View FIGURES 92 – 100. 92 – 94 View FIGURES 101 – 109. 101 – 103 ).................................................................. 36
35. Elytron with 2 large black spots. TL: 1.75–1.9 mm, TW: 1.55–1.6 mm ............................ S. pictipennis Lewis
- Elytron with 3 relatively small black spots, although additional black streaks are present on lateral or sutural margins. TL: 2.4 mm .............................................................................. S. maculata Korschefsky
36. Elytron with spots touching basal margin or suture ( Figs 98–106 View FIGURES 92 – 100. 92 – 94 View FIGURES 101 – 109. 101 – 103 ).............................................. 37
- Elytron without spots touching basal margin or suture ( Figs 95–96 View FIGURES 92 – 100. 92 – 94 ). TL: 2.51–2.93 mm, TW: 2.24–2.60 mm, TH: 1.48–1.71 mm .................................................................... S. octopunctata Wang & Ren sp. nov.
37. Elytral spots not touching suture at basal 2/5. TL: 2.0– 2.26 mm, TW: 1.76–1.82 mm ( Yu 1995).......... S. linguiformis Yu
- Elytral spots touching suture at basal 2/5, forming a common sutural spot ( Figs 98–106 View FIGURES 92 – 100. 92 – 94 View FIGURES 101 – 109. 101 – 103 )............................ 38
38. Parameres slightly shorter than or equal to penis guide ( Figs 267, 272 View FIGURES 265 – 274. 265 – 269 ).......................................... 39
- Parameres very short, about 1/2 of penis guide length ( Fig. 262 View FIGURES 255 – 264. 255 – 259 ). TL: 2.41–2.80 mm, TW: 2.18–2.47 mm, TH: 1.32–1.48 mm ....................................................................... S. jinpingensis Wang & Ren sp. nov.
39. Penis guide thin and slender, apices of parameres moderately broadening, clavate ( Fig. 267 View FIGURES 265 – 274. 265 – 269 ). TL: 2.26–2.70 mm, TW: 2.08– 2.44 mm, TH: 1.22–1.52 mm ................................................... S. clavata Wang & Ren sp. nov.
- Penis guide wide and short, apices of parameres not broadening ( Fig. 272 View FIGURES 265 – 274. 265 – 269 ). TL: 2.31–2.54 mm, TW: 2.08–2.34 mm, TH: 1.35– 1.42 mm ..................................................................... S. crassa Wang & Ren sp. nov.
40. Elytra yellow to yellowish brown, with eleven or twelve black spots ( Figs 110–118 View FIGURES 110 – 118. 110 – 112 )................................ 41
- Elytra reddish brown, with two large yellow irregular spots, and sixteen small black spots around their margins ( Figs 107– 109 View FIGURES 101 – 109. 101 – 103 ). TL: 2.70–2.90 mm, TW: 2.31–2.67 mm, TH: 1.42–1.45 mm ....................... S. taenia Wang & Ren sp. nov.
41. Pronotum with two black spots before base. TL: 3.2 mm, TW: 2.65 mm ···················································· S. undeci- mpunctata Miyatake
- Pronotum without spots ( Figs 110–118 View FIGURES 110 – 118. 110 – 112 )................................................................... 42
42. Spots on elytron touching the basal margin and suture ( Figs 113–118 View FIGURES 110 – 118. 110 – 112 )........................................... 43
- Spots on elytron not touching the basal margin and suture ( Figs 110–112 View FIGURES 110 – 118. 110 – 112 ). TL: 2.93–3.13 mm, TW: 2.67–2.80 mm, TH: 1.55– 1.71 mm ................................................................. S. neckiformis Wang & Ren sp. nov.
43. Penis guide very thick at base, apex strongly curved in lateral view, and very wide in ventral view ( Figs 287–288 View FIGURES 285 – 294. 285 – 289 ). TL: 2.3 4– 2.70 mm, TW: 2.11–2.41 mm, TH: 1.12–1.25 mm .................................... S. obesa Wang & Ren sp. nov.
- Penis guide distinctly thiner than S. obesa , apex not curved, and narrow in ventral view ( Figs 292–293 View FIGURES 285 – 294. 285 – 289 ). TL: 2.47–3.07 mm, TW: 2.21–2.84 mm, TH: 1.25–1.68 mm ........................................... S. robusta Wang & Ren sp. nov.
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