Calapnita dayak , Bernhard A. Huber, 2017

Bernhard A. Huber, 2017, Revision and cladistic analysis of the Southeast Asian leaf-dwelling spider genus Calapnita Simon (Araneae, Pholcidae), Zootaxa 4219 (1), pp. 1-63: 39

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Calapnita dayak

sp. nov.

Calapnita dayak  sp. nov. Figs 164–171View FIGURES 156 – 171

Calapnita vermiformis  ” (misidentification; only specimens listed below): Deeleman-Reinhold 1986b: 212. Huber 2011: 48.

Diagnosis. Distinguished from most species of vermiformis  group (except C. bugis  ) by wide obtuse tip of ventral flap of procursus ( Fig. 164View FIGURES 156 – 171); from most (except C. bario  ) also by long and slender palpal tibia (tibia length/width>2.5; Fig. 286View FIGURES 285 – 288); from several species also by very small proximal part of bipartite cheliceral apophyses ( Fig. 168View FIGURES 156 – 171), by narrow and distinct epigynal ‘knob’ ( Fig. 170View FIGURES 156 – 171), and by oval pore plates converging anteriorly and very close together ( Fig. 171View FIGURES 156 – 171).

Etymology. Dayak  is a loose term for over 200 riverine and hill-dwelling ethnic subgroups located principally in the interior of Borneo; noun in apposition.

Material examined. Holotype. INDONESIA-BORNEO: ♂, RMNHAbout RMNH ( ARA 17814View Materials), Central Kalimantan, Tumbang Tahai (2.033°S, 113.583°E), primary marshy forest, from leaves, 3/ 13.ix.1985 (S. Djojosudharmo).GoogleMaps 

Other material. INDONESIA-BORNEO: 1♂ 1♀, RMNHAbout RMNH ( ARA 17424View Materials), same data as holotypeGoogleMaps  . 1♂ 1♀, RMNHAbout RMNH ( ARA 17421View Materials), Central Kalimantan, Kaharian (2.033°S, 113.667°E), swampy primary forest, in foliage, 2– 16.ix.1985 (S. Djojosudharmo)GoogleMaps  . 1♂ 1♀, RMNHAbout RMNH ( ARA 17427View Materials), Central Kalimantan, 40 km NW Palangkaraya [~ 1.96°S, 113.75°E?], secondary forest, from leaves, ix.1985 (S. Djojosudharmo).GoogleMaps 

Assigned tentatively. INDONESIA-BORNEO: 1♂ 1♀, ZFMKAbout ZFMK (Ar 5330), East Kalimantan, Balikpapan [~ 1.25°S, 116.833°E], primary forest, 20.vii.1982 (Murphy, 11872 part).GoogleMaps 

Description. Male (holotype)

MEASUREMENTS. Total body length 3.5, carapace width 0.5. Leg 1: 25.4 (6.5 + 0.3 + 5.5 + 10.9 + 2.2), tibia 2: 4.2, tibia 3: 2.6, tibia 4: 4.7; tibia 1 L/d: 110. Distance PME-PME 160 µm, diameter PME 70 µm, distance PME- ALE ~30 µm; no trace of AME.

COLOR. Entire animal pale gray to whitish, legs slightly yellowish with brown patellae and tibia-metatarsus joints.

BODY. Habitus as in close congeners (cf. Figs 122, 124View FIGURES 120 – 127); ocular area barely elevated, each triad on very low hump; carapace without median furrow; clypeus unmodified; sternum as wide as long (0.40), unmodified.

CHELICERAE. As in Figs 168–169View FIGURES 156 – 171, apophyses near lamellae bipartite with small proximal part, with pair of barely visible lateral processes; without modified hairs; without stridulatory ridges.

PALPS. In general similar to C. vermiformis  (cf. figs 139 and 140 in Huber 2011); trochanter apophysis as in Fig. 167View FIGURES 156 – 171; femur as in Fig. 166View FIGURES 156 – 171, distal process at 50% of femur length; tibia very slender ( Fig. 286View FIGURES 285 – 288), length/diameter 0.44/0.15; procursus as in Figs 164–165View FIGURES 156 – 171, with simple ventro-distal flat sclerite; bulb length 0.30; embolus length 0.58; embolus tip with distinct fringes.

LEGS. Without spines and curved hairs; few vertical hairs; trichobothria on tibia 1 and tarsus 1 pseudosegments not seen.

Male (variation). Tibia 1 in 2 other males: 5.5, 6.1. The male from East Kalimantan shares the distinctively slender palpal tibia but is significantly larger and therefore assigned tentatively (tibia 1 missing; palpal tibia length/ diameter 0.59/0.18; bulb length 0.38; embolus length 0.80).

Female. In general similar to male, also eye triads at same distance. Tibia 1 in 3 females: 5.5, 5.6, 5.8. Epigynum as in Fig. 170View FIGURES 156 – 171, weakly sclerotized triangular plate with small and slender anterior ‘knob’; internal genitalia as in Fig. 171View FIGURES 156 – 171, longer than wide, with oval pore plates close together.

Distribution. Known from Central Kalimantan and (specimens assigned tentatively) East Kalimantan ( Fig. 283View FIGURE 283).


National Museum of Natural History, Naturalis


Zoologisches Forschungsmuseum Alexander Koenig














Calapnita dayak

Bernhard A. Huber 2017


Calapnita vermiformis

Huber 2011: 48
Deeleman-Reinhold 1986: 212