Selenocopris Burmeister

Nunes, Rafael V. & Vaz-De-Mello, Fernando Z., 2016, New brachypterous species of Dichotomius (Selenocopris) Burmeister (Coleoptera: Scarabaeidae: Scarabaeinae) with the definition of species groups and taxonomic notes in the subgenus, Zootaxa 4139 (1), pp. -1--1: -1

publication ID

http://doi.org/10.11646/zootaxa.4139.1.4

publication LSID

lsid:zoobank.org:pub:2744EC46-70A1-4267-AAB8-2131F5DC21DC

persistent identifier

http://treatment.plazi.org/id/03C187AE-453A-FF83-0EC8-F8AEFA00F94C

treatment provided by

Plazi

scientific name

Selenocopris Burmeister
status

 

Key for species groups and brachypterous species of Selenocopris Burmeister 

The following key allows the identification of the Selenocopris  species groups and the brachypterous species of the subgenus, now assigned to the quadraticeps  species group. When not specified, the characters fit both for males and females. Females differ from males by always having the 6th abdominal sternite not shortened medially (usually three times wider than the 5th sternite) and/or having modifications on the median portion of it or at the apex of pygidium ( Figs. 1View FIGURE 1 A, 1H–L).

1 Clypeo-genal junction border rounded, lacking angulation ( Fig. 1View FIGURE 1 F) and/or clypeal teeth separated from adjacent clypeal bor- der by external emarginations ( Fig. 1View FIGURE 1 F). Female pygidium swollen, apex excavated in some species ( Figs. 2View FIGURE 2 B–C) and/or female 6th abdominal sternite bearing a rounded central lobe advancing above pygidium apex. ( Fig. 1View FIGURE 1 I). South portion of Atlantic Forest ( Brazil, Paraguay, Argentina) and gallery forests at Central Brazil and South America Chaco ................................................................................................. bicuspis  species group

- Clypeo-genal junction border not rounded, forming at least an obtuse angle, which may be very obtuse on smaller males and females ( Fig. 1View FIGURE 1 G), clypeal teeth never separated from adjacent clypeal border by external emarginations ( Fig. 1View FIGURE 1 G). Female pygidium normally convex, apex never excavated; female 6th abdominal sternite either 3x wider than other segments ( Fig. 1View FIGURE 1 L) or having other modifications at the median portion (emarginations, grooves, projecions or tubercles) ( Figs. 1View FIGURE 1 H, 1K). Forest habitats in South America............................................................................... 2

2 Male 6th abdominal ventrite with strong transverse groove all along its extension ( Fig. 1View FIGURE 1 J). Female 6th abdominal ventrite with a "U" shape emargination at medial portion ( Fig. 1View FIGURE 1 K). Apex of male paramera curved inward (dorsal and ventral view) lacking rounded or circular excavations near basis (dorsal view). Pronotum simply convex, having at most a pair of tubercles at anterior portion......................................................................... ascanius  species group

- Male 6th abdominal segment lacking strong transverse groove all over its extension. Female 6th abdominal sternite with tubercles or other modifications at medial portion ( Figs. 1View FIGURE 1 A,H,L). Male paramera curved inward or outward (dorsal and ventral view) and/or truncated at apex and having rounded or circular excavations near basis (dorsal view) ( Fig. 3View FIGURE 3 C–D). Pronotum simply convex or having excavations, tubercles, lobes or strong declivity at anterior portion......................... 3

3 Male clypeo-genal junction border either in straight sub-straight or even acute angle, in some cases expanded ( Fig. 1View FIGURE 1 G). Genal border usually curved inward and clypeal teeth always curved upward. Female 6th abdominal sternite or 3x larger then 5th segment medially or having a central pair of tubercles ( Figs. 1View FIGURE 1 A, 1L)...... quadraticeps  species group (continues on couplet 5)

- Male clypeo-genal junction border always obtuse and never expanded. Genal border straight and clypeal teeth curved upward or not. Female 6th abdominal sternite with a pair of triangular projections advancing beneath pygidial apex ( Fig. 1View FIGURE 1 H)..... 4

4 Body almost always over 15 mm in length. Elytral striae and its punctures shallowly and weakly impressed ( Fig. 1View FIGURE 1 D). Elytral interstriae flat, with either silky aspect given by its microsculpture (chagrination) and/or bearing brown or blue reflections. Male paramera bearing longitudinal excavations (dorsal and ventral view) (as those on Fig. 3View FIGURE 3 C–D)...... fissus  species group

- Body almost always under 15 mm in length. Elytral striae and its punctures deeply impressed ( Fig. 1View FIGURE 1 E). Elytral interstriae shinny, not bearing microsculpture (chagrination), sometimes with color reflections. Male paramera lacking longitudinal excavations.......................................................................... spadiceus  species group

5 Brachypterous species (atrophied hind wings) ( Fig. 2View FIGURE 2 E): lacking humeral callus; pronotum wider than elytra (dorsal view); elytra strongly convex (lateral view) ( Fig. 1View FIGURE 1 B).............................................................. 6

- Macropterous species (fully developed hind wings) ( Fig. 2View FIGURE 2 E): having humeral callus; pronotum with the same width as elytra (dorsal view); elytra normally convex (lateral view).............................. macropterous species not dealt here

6 Pygidium bearing ocellate punctures near anterior margin (basis) ( Fig. 2View FIGURE 2 D). Chapada dos Guimarães, Mato Grosso state, Brazil........................................................... Dichotomius (S.) ingens (Luederwaldt 1935)  ( Fig. 3View FIGURE 3)

- Pygidium lacking ocellate punctures ( Fig. 2View FIGURE 2 C), at most having very fine punctures (20x)............................. 7

7 Posterior portion of pronotal disc bearing ocellate punctures ( Figs. 6View FIGURE 6 A–B). Elytral striae widened, lacking distinct punctures ( Figs. 6View FIGURE 6 A–B, E). Chapada dos Parecis at Mato Grosso state, Brazil................ Dichotomius (S.) paresi  n.sp. ( Fig. 6View FIGURE 6)

- Pronotal disc lacking ocellate punctures, those being sometimes restricted to posterior margin, anterior angles and/or to the anterior excavation. Elytral striae narrow and always bearing distinct punctures all over its extension ( Fig. 1View FIGURE 1 E).......... 8

8 Cephalic carina surface wrinkled (striated) ( Figs. 4View FIGURE 4 A–B, E). Anterior excavation of pronotum bearing coarse punctures ( Fig. 4View FIGURE 4 F). Chapadão do Céu, Goiás state, Brazil.............................................. Dichotomius (S.) darwini 

- Cephalic carina lacking striation. Anterior excavation of pronotum bearing at most fine punctures (20 x) (as on Fig. 7View FIGURE 7 E)... 9

9 Clypeo-frontal horn carinate, strongly transverse, widely emarginated at the apex and producing two tubercles at each apex ( Fig. 7View FIGURE 7 A). On smaller individuals, cephalic horn not producing lateral tubercles but still with a conspicuous emargination at apex. Smaller species (maximum body length: 19.5 mm; maximum pronotum width: 13 mm). Descalvado Municipality, São Paulo state, Brazil..................................................... Dichotomius (S.) periotoi  n. sp. ( Fig. 7View FIGURE 7)

- Clypeo-frontal horn carinate but not strongly transverse, lacking emargination and tubercles at the apex ( Fig. 5View FIGURE 5 A). Females and smaller individuals unknown. Larger species (maximum body length: 25 mm; maximum pronotum width: 14 mm). Serra da Canastra, Southwest of Minas Gerais state, Brazil.............................. Dichotomius (S.) manni  n. sp. ( Fig. 5View FIGURE 5)