Eumenes verticalis Say, 1824

Grandinete, Yuri Campanholo, Noll, Fernando Barbosa & Carpenter, James, 2018, Taxonomic Review of Eumenes Latreille, 1802 (Hymenoptera, Vespidae, Eumeninae) from the New World, Zootaxa 4459 (1), pp. 1-52: 30-31

publication ID

https://doi.org/10.11646/zootaxa.4459.1.1

publication LSID

lsid:zoobank.org:pub:633A9498-FCC6-4439-BE33-A45B9E51B550

persistent identifier

http://treatment.plazi.org/id/03C5878F-FFEE-F31E-FF30-9339D7F2F8BA

treatment provided by

Plazi

scientific name

Eumenes verticalis Say, 1824
status

 

Eumenes verticalis Say, 1824 

Eumenes verticalis Say, 1824  , 2: 346.

Type data: Type destroyed.

Type locality: “ Pennsylvania ” 

de Saussure, 1852, 1: 41.

Cresson, 1887: 287 (cat.).

Dalla Torre, 1894, 9: 33 (cat.).

Dalla Torre, 1904, 19: 25 (cat.).

Bequaert, 1928, 101: 1004.

Bequaert, 1938, 33: 61 (key), 64, 70 (type lost).

Bequaert & Taylor, 1942, 42: 68 (list).

Bequaert, 1944, 71: 78 (key), 86.

MacLachlan, 1980, 53: 618, 620 (key).

Buck et al. 2008, 5: 44 (key), 155.

Eumenes coloradensis Cresson, 1875  , 5: 717. REVISED STATUS.

Type data: Lectotype female at ANSP (examined).

Type locality: Colorado ( USA). 

Dalla Torre, 1894, 9: 21 (cat.).

Dalla Torre, 1904, 19: 21 (cat.).

Tucker, 1909, 22: 286.

Cresson, 1916, 1: 104 (designation of Lectotype).

Isely, 1917, 10: 348 (key), 359.

Bequaert, 1938, 33: 65, 69 (note on types).

Bequaert, 1944, 71: 78 (key), 81.

Eumenes (Alpha) cruciferorum Viereck, 1908  , 33: 388. REVISED STATUS.

Type data: Lectotype female (by present designation) and Paralectotype male at KUNHM (examined).

Type locality: “ Thomas’ Ranch, Oak Creek Canon , 6000 ft., 20 miles southwest of Flagstaff , Coconino Co., Arizona ” 

(USA).

Eumenes cruciferorum  ; Bequaert, 1938, 33: 69 (note on types).

Bohart, 1951: 885.

Eumenes (Alpha) enigmatus Viereck, 1908  , 33: 389. NEW STATUS.

Type data: Lectotype male (by present designation) and Paralectotype male at KUNHM (examined).

Type locality: “Thomas’ Ranch, Oak Creek Canon, 6000 ft., 20 miles southwest of Flagstaff, Coconino Co., Arizona”. Eumenes enigmatus  ; Bequaert, 1938, 33: 69 ( enigmaticus  [!]; note on type).

Bohart, 1951: 885.

Eumenes stenogaster Isely, 1917  , 10: 353. NEW STATUS.

Type data: Holotype female at USNM (n° 21379) (examined).

Type locality: “ White Mts , Novo Mexico  ; Rio Ruidoso ; about 6500 ft.” ( USA)  .

Bequaert, 1938, 33: 70 (note on types).

Bohart, 1951: 885.

Eumenes tricinctus Isely, 1917  , 10: 361. REVISED STATUS.

Type data: Holotype female at USNM (n° 21383) (examined).

Type locality: “ Oregon ”. 

Bequaert, 1938, 33: 70 (note on type).

Eumenes verticalis  var. (or subsp.) neoboreus Bequaert, 1944  , 71: 76, 87, NEWAbout NEW STATUS

Type data: Holotype male at MCZAbout MCZ (n° 33185) (examined). Three paratypes male at ROMAbout ROM (examined).

Type locality: “ Canada... Quebec: La Trappe ” (Holotype)  ; “ Canada: Ontario (paratypes)  .

Eumenes verticalis tricinctus  ; (4) 24 (9): 315.

Bohart, 1951: 885 (cat.)

Krombein, 1979: 1508 (cat.).

MacLachlan, 1980, 53: 618.

Ruiz C. et al., 1993, 88: 87 (list).

Rodríguez-Palafox, 1996: 479 (list).

Eumenes verticalis verticalis  ; Bohart, 1951: 885 (cat.).

Krombein, 1979: 1508 (cat.).

MacLachlan, 1980, 53: 618.

Ahlstrom, 1995, 1: 102 (list).

Eumenes verticalis neoboreus  ; Bohart, 1951: 885 (cat.).

Krombein, 1979: 1508 (cat.).

MacLachlan, 1980, 53: 618.

Eumenes verticalis coloradensis  ; Bohart, 1951: 885.

Krombein, 1958: 163;

Krombein, 1979: 1508 (cat.).

MacLachlan, 1980, 53: 618.

Ruiz C. et al., 1993, 88: 83, 87 (list).

Rodríguez-Palafox, 1996: 479 (list).

Eumenes cruciferoides Giordani Soika, 1978  , 29: 18 (key), 28. NEWAbout NEW SYNONYMY.

Type data: Holotype female at MSNV (examined).

Type locality: “ Messico: Nogares, dintorni di Orizaba, 1540 m  .” (Mexico).

Rodríguez-Palafox, 1996: 479 (list).

Distribution: Canada (Alta, B.C., Queb, Man*, Ont., Sask*); Mexico (Lower California *, Nuevo León *, Tamaulipas *, Veracruz); U.S.A. (AZ, CA, CT, CO, DC*, DE*, ID, IL, IN*, MA, MD*, MN*, MO, NC, ND, NH, NJ, NM, NV*, NY, OH*, OR, PA*, SD*, UT, VA*, WI*, WV*, WY).

* Records from the literature.

Comments and diagnosis. Eumenes verticalis  was originally described by Say (1824), from Pennsylvania (USA) (the type was destroyed), and currently it has four recognized subspecies: E. verticalis coloradensis Cresson, 1875  ; E. verticalis tricinctus Isely, 1917  , E. verticalis neoboreus Bequaert, 1944  , and E. verticalis verticalis (Say)  , 1824.

We had examined all the type specimens from all the subspecies of E. verticalis  plus 137 specimens (57 females and 80 males, all with male genitalia extracted and examined) and as we stated for E. consobrinus  , E. verticalis  has some intraspecific variation along its distribution, mainly in color and punctation of T2 throughout the east coast of the USA and southeast Canada, which leads us to propose the abandonment of these subspecies.

Because the primary type of the species was destroyed, we compared specimens of the typical E. verticalis  with the original description of Say (1824) and we selected one of these for the redescription of the species. This specimen is established as the lectotype.

Giordani Soika (1978) described E. cruciferoides  ( Fig. 12 View Figure ) and noted that it resembles E. crucifera bolliformis  but also noted that they differ in the “punctation of the clypeus, punctation and color (p. 29)”. We compared the type of E. cruciferoides  and to the type of E. bolliformis  and, although the former is known only from the type, we maintain it as a different species based on the following features: the punctures on clypeus are coarser and extend to the apical region; the punctation on the mesepisternum is denser, with the interspaces forming carinae, particularly in the center-basal region; the pubescence on the head and mesosoma is slightly longer, most evident on the frons and scape.

Although E. cruciferoides  is known only from the type specimen, we compared it with the typical E. verticalis  and no morphological differences were found besides having the ferrugineous marks contiguous with the yellow marks along body. As we observed for E. fraternus  and E. crucifera var. bolliformis  , specimens found the the southern USA and/or Mexico present ferrugineous marks along the body. As with E. verticalis  , E. cruciferoides  also has the clypeus with coarse punctation and long pubescence along the body, including on the scape (longer than the pedicel). For these reasons, we treat E. cruciferoides  as a new synonym of E. verticalis  .

NEW

University of Newcastle

MCZ

Museum of Comparative Zoology

ROM

Royal Ontario Museum

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hymenoptera

Family

Vespidae

Genus

Eumenes

Loc

Eumenes verticalis Say, 1824

Grandinete, Yuri Campanholo, Noll, Fernando Barbosa & Carpenter, James 2018

2018
Loc

Eumenes cruciferoides

Giordani Soika 1978

1978
Loc

neoboreus

Bequaert 1944

1944
Loc

Eumenes stenogaster

Isely 1917

1917
Loc

Eumenes tricinctus

Isely 1917

1917
Loc

Eumenes verticalis

Say 1824

1824