Dolicholatiridae Kantor, Fedosov, Kosyan, Puillandre, Sorokin, Kano, R. Clark & Bouchet,

Harzhauser, Mathias, Landau, Bernard M. & Vermeij, Geerat J., 2024, The Dolicholatiridae and Fasciolariidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea, Zootaxa 5470 (1), pp. 1-92 : 7

Family Dolicholatiridae Kantor, Fedosov, Kosyan, Puillandre, Sorokin, Kano, R. Clark & Bouchet,

2021

Original diagnosis. “ Shell small to medium-large (60 mm), from broad to narrow fusiform or turriform. Spire tall, siphonal canal from short to long, constituting nearly half of SL. Protoconch paucispiral, with smooth initial part and later whorls sometimes bearing axial ribs. Spiral sculpture of fine to strong cords, sometimes pronounced only in interspaces between axial ribs. Axial sculpture of strong, broad axial ribs present in most species. Columella sometimes with two weak columellar plaits ” ( Kantor et al. 2021: 803).

Genus Dulaiania nov. gen.

Type species. Fasciolaria pleurotomoides Hoernes & Auinger, 1890 ; Middle Miocene , Romania .

Diagnosis. Small fusiform shell with conical spire, protoconch low, dome-shaped, sculpture of prominent primary and weaker secondary spiral cords, no axial ribs, whorl profile changing from nearly flat-sided to weakly shouldered with steep subsutural ramp; strong growth lines causing scabrous sculpture on subsutural ramp on last whorls, columella with three columellar folds, very weak parietal swelling, outer lip lyrate within.

Description. As for type species.

Etymology. In honor of Alfréd Dulai (Hungarian Natural History Museum, Budapest), who kindly helped us many times with type material for our revisions of Paratethyan gastropods.

Included species. Only the type species is known.

Stratigraphic and geographic range. Badenian/Langhian (Middle Miocene), Central Paratethys Sea.

Paleoenvironment. Unknown.

Discussion. Dulaiania nov. gen. is characterized by its lack of axial sculpture and presence of prominent spiral sculpture on early teleoconch whorls, which is an unusual feature in Fasciolariidae but frequent in Dolicholatiridae . In addition, the dome-shaped protoconch is characteristic for this family. Teralatirus Coomans, 1965 [type species Latirus ernesti Melvill, 1910 (= Teralatirus roboreus (Reeve, 1845) ; present-day, western Atlantic] differs from Dulaiania in its broader apex, which has a large and blunt protoconch, and in its less incised suture, the lower last whorl and the shorter siphonal canal (see Simone et al. 2013). Similarly, Dolicholatirus Bellardi, 1884 [type species Turbinella bronni Michelotti, 1847 , Miocene, Italy] has predominant spiral sculpture on early teleoconch whorls and a broad, dome-shaped protoconch but is usually more fusiform. Strongly raised growth lines along the subsutural collar comparable to those of Dulaiania also occur in the extant Dolicholatirus celinamarumai Kosuge, 1981 from the Western Pacific. Crassicantharus Ponder, 1972 [type species Crassicantharus norfolkensis Ponder, 1972 ; present-day, Australia], as reviewed by Fraussen & Stahlschmidt (2015) differs from Dulaiania in its squatter shape and prominent, axially arranged nodes. Dulaiania bears three weak columellar folds, whereas most extant Dolicholatiridae typically have two folds.

Some extant species of the fasciolariid Turrilatirus Vermeij & Snyder, 2006 develop similar early teleoconch whorls with dominant spiral sculpture and subobsolete axial ribs [e.g., Turrilatirus turritus ( Gmelin, 1791) ]. However, Turrilatirus differs from Dulaiania in its adpressed suture, and it lacks prominent growth lines.