Miasa trifoliusa Zheng & Chen

Zheng, Yan-Li, Yang, Lin, Chen, Xiang-Sheng & Luo, Xu-Qiang, 2018, Two new species of the genus Miasa Distant, 1906 from China, with a key to all species (Hemiptera, Fulgoromorpha, Dictyopharidae), ZooKeys 754, pp. 23-32: 25-27

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Miasa trifoliusa Zheng & Chen

sp. n.

Miasa trifoliusa Zheng & Chen  sp. n. Figs 21-32


♂, BL: 14.2 mm; HL: 1.8 mm; HW: 0.4 mm; FWL: 10.7 mm.


General colour in dried specimens ferruginous-brown, marked with faint yellow and reddish brown. Cephalic process of the base brown, terminal black, brown on side. Frons brown with faint yellow marks. Frontoclypeal dark with paired brown blotchy markings. Compound eyes dark brown, ocelli light pink. Antennae brown. Pronotum and mesonotum brown, the middle faint yellow. Forewings with stigmal area and posterior margin broadly dull ochreous, a large oblique triangular apical streak, and a narrow streak along nodal line fuscous; hind wings with an apical fuscous spot. Legs brown with dark spot.

Cephalic (Figs 21-25) process relatively long, distinct upturned, ratio length to length of pronotum and mesonotum combined 0.7. Vertex (Figs 21-25) with lateral margins carinate, sub-parallel at base, sharply sinuate in front of eyes, then narrowing to arrowhead at apex, ratio of length to width between eyes 3.5. Frons (Fig. 26) elongate, median carina complete and elevated, length approx. 4.8 times long than width. Pronotum (Figs 21-24) distinctly shorter than mesonotum medially in the middle line, median carina obscure, lateral carina distinct, ratio length to length approx. 0.3:1. Mesonotum (Figs 21-24) median carina obscure, lateral carina distinct. Forewings (Figs 22, 23) elongate, with ratio of length to width approx. 4.0:1; stigma distinct, with four cells, CuA vein first branched before Sc+R and M veins near middle; crossveins very scarce, forming a nodal line along Sc+R, M and CuA veins at apical 1/3; apical cells approx. 12; Pcu and A1 veins fused into a long Pcu+A1 vein at apical 1/6 in clavus. Legs long and thin, fore femur not flattened and dilated, with one minute, short, blunt spine near apex; hind tibia with five lateral black-tipped spines and six apical black-tipped teeth, hind tibiae I with ten and tarsomeres II with eight black-tipped apical teeth, respectively.

Male genitalia. Pygofer (Figs 27-29) wider ventrally than dorsally (aprpox. 4.5:1), hatchet-shaped in lateral view. Gonostyles (Figs 27, 28) relatively large, broadening towards apex in lateral view (Fig. 27), posterior margin straight, upper margin with dorsally directed, black-tipped process near middle, with ventrally directed, hook-like process near sub-middle on outer upper edge. Anal tube (Figs 27, 29) wide and narrow down in dorsal view, ratio length to width approx. 1.4:1. Aedeagus (Figs 30-32) with one pair of special long endosomal processes, processes with apex acute, sclerotised and pigmented. Phallobase sclerotised and pigmented at base, with three pairs of membranous lobes at apex: the dorsal lobes relatively small and the ventral two pairs of membranous lobes large and connected in ventral view (Fig. 31).

Type material.

Holotype ♂, China, Yunnan, Xishuangbanna Menglun. 18.VIII.2014, Wang Ying-Jian.


China (Yunnan).

Differential diagnosis.

This species is similar to M. wallacei  Muir, but can be distinguished most easily by the phallobase conformation. The former is membranous with a pair of dorsal lobes directed posteriorly, and two pairs of ventral lobes: large, almost equal in length, apically bifurcate, directed dorsally, the latter membranous with a pair of dorsal lobes directed posteriorly, and two pairs of ventral lobes: upper pair large and elongate, directed dorsally; lower pair relatively small and rounded. There have differences in body colour and in the lengths and widths of the forewings, but the differences are not obvious.


This new species is named with the Greek word " trifoliusa  " referring to phallobase with three pairs of membranous lobes at apex.