Dryptus pardalis (Férussac, 1821)

Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, Zootaxa 3054, pp. 1-59: 10-11

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Dryptus pardalis (Férussac, 1821)


Dryptus pardalis (Férussac, 1821) 

( Figs 2View FIGURE 2 A–B, 4 A)

Helix pardalis Férussac 1821: 48  . Type locality: not given. Plekocheilus pardalis (Férussac)  ; Tello 1975: 201. Dryptus pardalis (Férussac)  ; Breure 1984: 490, 495; Richardson 1995: 201 (references).

Bulimus leptochilus Pfeiffer 1848 b: 48  . Type locality: La Baja, Prov. Pamplona.

Bulimus lindeni Reeve 1848  : pl. 31 fig. 189. Type locality: La Baja, Prov. Pamplona.

Bulimus plumbeus Pfeiffer 1855 b: 114  . Type locality: Venezuela. Lectotype BMNH 1975130 ( Breure 1978: 31, pl. 11 fig. 1).

Material. Cesar, foothills of Sierra de Perijá, 10–30 km E of Becerril, El Roncón, B. Malkin leg./ 1968–1969: FMNHAbout FMNH 163692–163694; Socorpa Mission trail [approx. 09° 47 ’ N 73 ° 02’ W, 1350 m; see Harvey et al. 2008], Becerril, Sierra de Perijá, Malkin leg./ 1968: FMNHAbout FMNH 159207; Guajira, [Arroyo] Potrero de Venancio, 1100 m [not seen] ( Breure 1984); hills S of Carraipía, O.L. Haught leg./ 1943: USNMAbout USNM 599500; near Carraipía, Haught leg.: USNMAbout USNM 543494; vicinity of Carraipía, Haught leg./ 1947: MCZAbout MCZ 183880; Norte de Santander, Cúcuta, Hno. Nicéforo María leg./ 1936: ANSPAbout ANSP 168419; Hacienda Villa Felisa, 18 km S of Cúcuta, 1100 ft [~ 333 m, erroneous; actual elevation approx. 560 m], M.A. Carriker leg./ 1947: USNMAbout USNM 590659; Palogordo, Táchira Valley, S of Cúcuta, 4500– 5500 ft [~ 1360–1660 m], Carriker leg./ 1947: USNMAbout USNM 590635; Petrolia [sic, = Petrólea], Olsson leg./ 1935: ANSPAbout ANSP 166132; Petrólea, near sea level, Carriker leg.: USNMAbout USNM 517813; Ocaña, G. Wallis leg./ 1875: ZMUZAbout ZMUZ 511536; Buenos Aires, Highway Abrego–Sardinata, 6000–7000 ft [~ 1800–2120 m], Carriker leg./ 1947: USNMAbout USNM 486845; “Sta Librada” [= Santa Librada] (Sarare), Rochereau leg./ 1937: DMNHAbout DMNH 141725; Santander, La Baja ( Baker 1926).?, New Granada: CMCAbout CMC C 11024View Materials. Venezuela, Táchira, La Fría, H.B. Baker leg.: UMMZAbout UMMZ 244280;?, 120 km E of Pamplona, Hno. Nicéforo María leg.: ANSPAbout ANSP 170736.

Altitudinal range. The localities reported here range between “near sea level” and 2435 m.

Remarks. Bulimus leptochilus Pfeiffer, 1848  and Bulimus lindeni Reeve, 1848  were placed in the synonymy of this species by Pilsbry (1895: 40). The type locality for both is “La Baja, Province of Pamplona.” Richardson (1995: 202) was the first to group Bulimus plumbeus Pfeiffer, 1855  , hitherto considered a species of Thaumastus Albers, 1860  , together with Dryptus pardalis  . See also the remarks under D. moritzianus  , some records of which might be misidentified given that these species are closely related. There seem to be two distinct shapes of what is referred to as Dryptus pardalis  : (1) some are cigar-shaped, tapering about evenly at either side, with the central part having near-parallel outlines; and (2) some with the spire substantially less wide, and the body whorl wider, so the whole shell is more “bulimuliform” in appearance. In addition, there is substantial variation in sculpture, with some specimens being nearly smooth (even fresh ones), and others are very sculptured.

Two samples are tentatively referred to this species. One is from near Cúcuta ( UVZ 8101), corresponding to the second shape variation above. The other is a lot from Guajira, near Carraipía, ( USNMAbout USNM 543492). While these specimens look very different from other Dryptus  , they might be misshapen D. pardalis  . Specifically, they are too short, and have a very globose last whorl, which includes a much shorter aperture. One of the specimens has a strong parietal callus. Two reasons make it probable that they are D. pardalis  : (1) this species is known from the same locality (it is the only Dryptus  known from Guajira); (2) two other lots of D. pardalis  are known from Carraipía: USNMAbout USNM 543494 (2 specimens) and USNMAbout USNM 599500 (1 specimen). Furthermore, one could argue that the deformed specimens are shorter and globose due to climatic differences. Interestingly, the spire is not globose, it seems to be just the body whorl having this characteristic. However, the following arguments are against this classification: (1) two specimens are available and both are somewhat similarly misshapen; (2) these individuals are umbilicate (more so than any specimen of D. pardalis  ); (3) neither one has much of a columellar fold. As is the case for several other species of Dryptus  , D. pardalis  may be widely distributed in Venezuela; Tello (1975) listed four Venezuelan Estados (states) in the distribution of this species, including Táchira and Zulia (bordering with Colombia), with Cumaná (Sucre) being the easternmost record. However, these records are in need of critical review.


Field Museum of Natural History


Smithsonian Institution, National Museum of Natural History


Museum of Comparative Zoology


Academy of Natural Sciences of Philadelphia


Zoologisches Museum der Universitaet Zuerich


Delaware Museum of Natural History


Coleccion Mastozoologica del Centro de Investigacion en Biodiversidad y Conservacion,Universidad Autonoma del Estado de Morelos


University of Michigan, Museum of Zoology














Dryptus pardalis (Férussac, 1821)

Borrero, Francisco J. & Breure, Abraham S. H. 2011


Helix pardalis Férussac 1821: 48

Richardson 1995: 201
Breure 1984: 490
Tello 1975: 201


Bulimus leptochilus

Pfeiffer 1848: 48


Bulimus plumbeus

Pfeiffer 1855: 114