Stenosfemuraia cuadrata González-Sponga, 2005

Huber, Bernhard A. & Arias, Quintin, 2017, Revision of the Venezuelan spider genus Stenosfemuraia González-Sponga, with new generic and specific synonymies (Araneae, Pholcidae), Zootaxa 4341 (4), pp. 495-508: 500-503

publication ID

publication LSID

persistent identifier

treatment provided by


scientific name

Stenosfemuraia cuadrata González-Sponga, 2005


Stenosfemuraia cuadrata González-Sponga, 2005 

Figs 15–27View FIGURES 15 – 21View FIGURES 22 – 27, 45–48View FIGURES 41 – 52

Stenosfemuraia cuadrata González-Sponga, 2005: 106  , pl. 4, figs 1–9 (♂ ♀). Stenosfemuraia  sp.: Bruvo-Mađarić et al. 2005: 663 ( DNA sequence: 28S); Astrin et al. 2006: 445 ( DNA sequences: 16S, CO1).

Diagnosis. Males are easily distinguished from S. parva  by armature of male chelicerae ( Fig. 24View FIGURES 22 – 27; one pair of simple frontal apophyses; see also fig. 3 in González-Sponga 2005); from S. pilosa  by shape of procursus ( Figs 47–48View FIGURES 41 – 52; slender in dorsal view, without subdistal side branch, with simple tip; possibly indistinguishable from S. parva  ); from both species also by more slender femora (~0.13 versus 0.19–0.21 in S. parva  and 0.29–0.34 in S. pilosa  ). Females differ from congeners by sclerotized lateral elements of epigynal plate ( Figs 19View FIGURES 15 – 21, 26View FIGURES 22 – 27; absent in S. pilosa  ; rather parallel orientation in S. parva  ) and by distinctive shape of anteriorly contiguous pore plates ( Figs 21View FIGURES 15 – 21, 27View FIGURES 22 – 27). Males and females also differ from congeners by longer legs (♂ tibia 1 L/d: ~60 versus <40; ♂ tibia 1:>5.0 versus <4.0; ♀ tibia 1:>3.0 versus <3.0).

Type material. VENEZUELA: Vargas: 22♂ 16♀ types (+8 juvs and 4 prosomata) (see Notes below), MIZA ( GS 1164View Materials part), Hacienda El Limón, Municipio Carayaca, 950 m [~ 10.532°N, 67.119°W; see Notes below], 12.viii.1989GoogleMaps  4.v.1991 (A.R. Delgado de González , M.A. González-Sponga), 3♂ examined by BAH  .

Other material examined. VENEZUELA: Aragua: 4♂ 3♀ 1 juv., ZFMK (Ar 18253), forest above Colonia Tovar (10.417°N, 67.300°W), ~ 2100 m a.s.l., under dead leaves on ground, 26.xi.2002 (B.A. Huber)GoogleMaps  ; 4♂ 6♀ in pure ethanol, ZFMK (Ven 02/100-10), same dataGoogleMaps  . 2♀, MNHN (Ar 10052 part), with Simon’s label “14653 Psil.— Tovar!”, no further data [leg. E. Simon, 1887–88].

Redescription. Male ( ZFMK Ar 18253). MEASUREMENTS. Total body length 2.5, carapace width 1.0. Distance PME-PME 80 µm, diameter PME 100 µm, distance PME-ALE 90 µm, distance AME-AME 20 µm, diameter AME 20 µm. Sternum width/length: 0.75/0.50. Leg 1: 21.1 (4.9 + 0.4 + 5.1 + 8.8 + 1.9), tibia 2: 3.1, tibia 3: 2.4, tibia 4: 2.8; tibia 1 L/d: 60. Femora 1–4 width (at half length): all ~0.13.

COLOR (in ethanol). Carapace ochre-yellow with brown marginal bands and median mark including ocular area and clypeus ( Fig. 15View FIGURES 15 – 21); sternum whitish; legs ochre-yellow, femora and tibiae with subdistal darker rings (and indistinct light tips); abdomen pale gray, dorsally and laterally with dark bluish internal marks, ventrally with indistinct light brown plate in front of gonopore, without mark behind gonopore.

BODY. Habitus as in Figs 15–16View FIGURES 15 – 21; ocular area raised; carapace elevated, with deep median furrow; clypeus and sternum unmodified.

CHELICERAE. With one pair of simple (undivided) frontal apophyses ( Fig. 24View FIGURES 22 – 27).

PALPS. As in Figs 22–23View FIGURES 22 – 27; coxa with retrolatero-ventral apophysis; trochanter barely modified; femur with retrolatero-ventral process proximally and small ventral apophysis distally; procursus very simple, slender in dorsal view, without subdistal side branch ( Figs 47–48View FIGURES 41 – 52); bulb with two dorsal processes, distal apophysis curved, very slender towards tip ( Figs 45–46View FIGURES 41 – 52).

LEGS. Without spines; with curved hairs on metatarsi 1–3, few vertical hairs; retrolateral trichobothrium on tibia 1 at 6%; prolateral trichobothrium present on tibia 1; tarsus 1 with ~25 pseudosegments, distally distinct.

Male (variation). Tibia 1 in 8 other males: 5.0–5.7 (mean 5.3). Internal abdominal marks less distinct in types. One male with few curved hairs also on tibiae 1–3.

Female. In general similar to male ( Figs 17–18View FIGURES 15 – 21), legs without curved hairs. Tibia 1 in 7 females: 3.3–3.7 (mean 3.5). Epigynum as in Figs 19–20View FIGURES 15 – 21, 25–26View FIGURES 22 – 27; with large protruding weakly sclerotized area in front of light brown epigynal plate, with pair of dark lateral sclerites in transversal orientation, posterior margin evenly curved; posterior plate simple. Internal genitalia as in Figs 21View FIGURES 15 – 21 and 27View FIGURES 22 – 27, with pair of anteriorly contiguous pore-plates, membranous median sac, and anteriorly diverging sclerites/‘wings’.

Natural history. All specimens were collected from forest leaf litter. González-Sponga (2005) specifically mentions Cecropia  leaves.

Distribution. Known from medium to high elevation forests (950–2100 m a.s.l.) in Aragua (Tovar area) and Vargas states, Venezuela ( Fig. 1View FIGURE 1).

Notes. González-Sponga (2005) reported 1♂ holotype (“1164a”), 1♀ paratype (“1164b”), and 21♂ 27♀ adult paratypes without specific collection number. Only one vial was found at MIZA, labeled “1164” and containing 22♂ 16♀ 8 juvs and 4 prosomata. Of the 22♂, three belong to S. parva  and were separated (see above) (and clearly labeled as being part of the S. cuadrata  type series). The label says “Hda El Limón, Dpto Vargas, DF” (which differs only with respect to administrative units from the locality as published in the original description: “Hacienda El Limón, Municipio Carayaca, Estado Vargas) and lists four dates: “ 12-8-89, 29-10-89, 10-8-90, 4-5- 91 ” (which agrees with the range in the original description). We suspect that this is the entire type series and that a holotype was never physically separated from this series. Since the identity of the species is beyond doubt, and since the holotype might just be misplaced rather than lost, we prefer not to designate a neotype and to treat the type specimens unspecifically as “ types ”.

González-Sponga (2005) gives the distribution of S. cuadrata  as “Galipán, Parque Nacional El Avila, Estado Vargas ”. This does not agree with the type locality. The exact coordinates of the type locality are not known to us, but Hacienda El Limón is close to the town of Carayaca, while Galipán [San Antonio de Galipán] is a town within the limits of El Ávila National Park and lies about 25 km E of the type locality. We do not have an explanation for this contradiction. In addition, the coordinates in González-Sponga (2005) are clearly wrong (about 32 km SW of Carayaca, in the state of Aragua).

Some measurements in González-Sponga (2005) are clearly wrong, e.g.: female carapace width (0.4; should be ~0.7).


Department of Natural Resources, Environment, The Arts and Sport


Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez


Biologische Anstalt Helgoland Marine Station


Zoologisches Forschungsmuseum Alexander Koenig














Stenosfemuraia cuadrata González-Sponga, 2005

Huber, Bernhard A. & Arias, Quintin 2017

Stenosfemuraia cuadrata González-Sponga, 2005 : 106

Astrin 2006: 445
Gonzalez-Sponga 2005: 106
Bruvo-Madaric 2005: 663