Amphineurus (Rhamphoneurus) podenasi, Santos & Santos & Ribeiro, 2022

Santos, Daubian, Santos, Rodrigo dos Reis & Ribeiro, Guilherme Cunha, 2022, Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae), Arthropod Systematics & amp; Phylogeny 80, pp. 439-494 : 439

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Amphineurus (Rhamphoneurus) podenasi

sp. nov.

3.3.21. Amphineurus (Rhamphoneurus) podenasi sp. nov.

Material examined.

Holotype. ♂, Chile, Arauco, Nahuelbuta, Pichinahuel [37°48′S 73°02′W], 1400-1600 m, 12/ 20-II-1953, Peña (USNM) GoogleMaps *. Paratypes: 1 ♂, Chile, Chiloé Is., Rio Coluco [42°6′S 73°55′W], 30-I-1952, Peña (USNM) GoogleMaps *; 1 ♂, Chile, Llanquihue, Carelmapu [41°44′S 73°40′W], Coast North of Canal de Chacao , 18-III-1957, Peña (USNM) GoogleMaps *. - Additional material: CHILE. 1 ♀ and 1 [sex unknown], Chiloé Is., Dalcahue [42°22′S 73°39′W], II-1954, Peña (USNM); 1 [sex unknown], Chiloé Is., Aulen [42°2′S 74°1′W], 8-II-1952, L.P. Guzman (USNM); 1 [sex unknown], Chiloé Is., Chaiten [42°58′S 72°32′W], 5/ 8-II-1954, Peña (USNM) GoogleMaps .


The species is named after Sigitas Podenas, a skilled Tipulomorpha expert.


This species is distinguished by only a dark spot on R2 and basal deflection of R5 longer than r-m. Furthermore, this species is characterized by male terminalia with sheath of aedeagus with a spur, tip of right mesal lobe curved downwards and sheath of aedeagus with rounded spur.


Wing length 5.73 mm, width 1.83 mm. - Coloration: General coloration brownish-yellow. Rostrum and palpus brown. First segment of antenna yellow, remainding segmens dark yellow. Head dark brown. Eyes black. Thorax brownish yellow, pleura pale. Contrasting golden setae on prescutum. Scutum without stripes. Halter brownish-yellow. Coxae and legs brownish yellow. Abdomen dark brown. - Head (dorsal view Fig. 26B View Figure 26 , ventral view Fig. 26A View Figure 26 ): Rostrum bifurcated with long appendices; first palpal segment shorter than terminal segment; scape thinner than pedicel; flagellomeres gradually reducing in length, resulting in shorter terminal flagellomeres. - Thorax (lateral view Fig. 26E View Figure 26 , dorsal view Fig. 26F View Figure 26 ): Thorax with several setae, including long setae on prescutum. Anatergite shorter than katatergite. Halter with dilated knob. Wing (Fig. 26D View Figure 26 ) almost clear of markings, except for strong marking covering R2; veins near fork of bM (Fig. 26C View Figure 26 ) faded; M1+2 longer than basal deflection of M1; R2+3+4 shorter than both R2+3 and basal deflection of R5. - Female terminalia (Fig. 26H View Figure 26 ): Female tergite IX longer and wider than tergite X; cercus shorter than hypoginial valves, curved dorsad. Both cercus and hypogynial valve very slender. - Male terminalia (Fig. 26G View Figure 26 ): Male tergite IX slightly darkened along posterior margin, with wide V-shaped median notch. Ventral branch of gonocoxite inclined medially in distal tip. Dorsal branch of gonocoxite nearly half-length of ventral branch. Lobe of gonostylus with rounded lobule; lobule longer than wide, longer than half-length of lobe of gonostylus, narrower than stem. Clasper of gonostylus with differently shaped branches: lateral branch straight; medial branch curved, club shaped. Mesal lobes of gonocoxite asymmetrical; left mesal lobe short with sideways projection; right mesal lobe curved. Sheath of aedeagus straightened, with rounded projection; darkened only on tip.


Specimens of this new species were previously identified by C.P. Alexander as A. (R.) insanus Alexander. This species resembles A. (R.) oosterbroeki sp. nov. but differs mainly in the length of r-m, concentration of setae on the gonocoxite, shape of male tergite IX and right mesal lobe.