Carabus (Lipaster) stjernvalli paravanicus, Fominykh & Zamotajlov & Khomitskiy & Titarenko, 2020

Fominykh, Dmitriy D., Zamotajlov, Alexandr S., Khomitskiy, Eugeniy E. & Titarenko, Andrey Yu., 2020, Contribution to the knowledge of Carabus (Lipaster) stjernvalli Mannerheim, 1830 in the Caucasus, with description of two new subspecies (Coleoptera: Carabidae: Carabini), Zootaxa 4729 (3), pp. 301-328 : 305-306

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Carabus (Lipaster) stjernvalli paravanicus

subsp. nov.

2. Carabus (Lipaster) stjernvalli paravanicus   subsp. nov.

( Figs. 1 View FIGURE 1 , 3–6 View FIGURES 3–6 , 92 View FIGURES 92–101 , 115–118 View FIGURE 115 View FIGURE 116 View FIGURE 117 View FIGURE 118 )

Type material. Holotype: ♂, Georgia, Samtskhe-Dzhavakheti, W bank of Paravani Lake near Aspara vill., E slopes of Kyerogly Mt., 2100-2200 m, N 41°28’24.99” / E 43°46’19.65”, 04.- 05.06.2015 (D. Fominykh, A. Zubov & A. Safronov) ( ZISP) GoogleMaps   . Paratypes: 444♂♂, 490♀♀, same data as holotype, pitfall traps ( ZISP, MPSU, CFDD, CTAY, CZAM, CKEE, CSAA) GoogleMaps   .

Description. Medium sized subspecies ( Figs. 3–6 View FIGURES 3–6 ). GBL = 26.8–34.8 mm, males 26.8–30.0 mm, females 28.6– 34.8 mm, SBL = 23.6–31.4 mm in both sexes. Body rather robust, ovate. Underside black, dorsum generally with bright metallic luster, green, bronze, more seldom violet or blue, in females less bright, somewhat mat; pronotum basally and elytra with metallic luster. Head broad, a little narrower than pronotum, HL = 4.0– 4.8 mm in males, 4.2–5.6 mm in females, HW = 4.8–6.0 mm in males, 5.4–6.6 mm in females; antennae short. Pronotum transverse, PW = 6.6–8.6 mm in males, 7.8–9.8 mm in females, PB = 5.4–6.8 mm in males, 6.0– 7.8 mm in females, PL = 4.0–5.0 mm in males, 4.4–5.6 mm in females, PW/HW = 1.2–1.6 in both sexes, PW/PL = 1.2–1.6 in both sexes, PW/PB = 1.1–1.4 in both sexes; hind angles faintly protruding posteriorly; disk rugose-punctate. Legs short. Elytra convex, EW/PW = 1.2–1.5 in both sexes; EL = 15.6–17.8 mm in males, 17.2–20.8 mm in females, EW = 9.8–11.4 mm in males, 10.4–12.6 mm in females, EL/EW = 1.5–1.8 in both sexes; elytral punctation faint and sparse, usually with only several rarefied impressions behind.

Male genitalia ( Fig. 92 View FIGURES 92–101 ) in general shape of the projections and lobes of endophallus resemble other forms of C. stjernvalli   .

Differential diagnosis and remarks. This subspecies differs from the other C. stjernvalli   taxa, first of all, in its body proportions ( Table 1 View TABLE 1 , Fig. 116 View FIGURE 116 ). In particular, it is distinguishable from C. stjernvalli stjernvalli   and C. stjernvalli gvalijai   by broader elytra. Among all known forms it is the closest morphologically to C. stjernvalli tabackuriensis   , but is distinguishable by body size and some proportions. The new subspecies also differs from all other taxa of C. stjernvalli   in the structure of endophallus ( Fig. 92 View FIGURES 92–101 ), particularly ventroapical and ventrobasal lobes are more pronounced, also a small projection, missing in the other populations from Trialeti Mt. Range and Dzhavakheti Upland, is present beneath ventrobasal lobe.

Distribution and habitat. The new subspecies populates west spurs of the Samsar Mt. Range at western bank of Paravani Lake. It inhabits alpine meadows at 2100–2200 m. The imago are active from the beginning of June. The following Carabus   species occur together with this subspecies: C. armeniacus Mannerheim, 1830   , C. calleyi Fischer von Waldheim, 1823   , C. cribratus Quensel, 1806   , and C. maurus Adams, 1817   .

Etymology. The subspecific epithet refers to the type locality, vicinity of Paravani Lake in Georgia.


Zoological Institute, Russian Academy of Sciences


Department of Microbiology, Songkla University