Chimeromyina concilia Arillo & Grimaldi

Grimaldi, David A., Cumming, Jeffrey M. & Arillo, Antonio, 2009, Chimeromyiidae, a new family of Eremoneuran Diptera from the Cretaceous, Zootaxa 2078, pp. 34-54 : 50-52

Chimeromyina concilia Arillo & Grimaldi , sp. n.

( Fig. 13 )

Diagnosis. As for genus.

Description. HEAD: Fairly broad, width more than 1.5x width of notum; eyes large, bare, facets not differentiated. Ocelli on small tubercle, tubercle with 2 pairs long setae. Antenna with apical arista; 3 aristomeres present. Vibrissae not present. THORAX: 0.44 mm long, bristly, each side of notum with 2 long notopleural setae and supra-alar seta; 2 pairs of dorsocentrals, row of long setulae anterior to dorsocentrals. Acrostichal setulae in one, extensive row, extended to posterior margin of notum. Scutellum with 2 pairs setae, apical pair long (ca. 2x length of other pair), cruciate. Wing 0.93 mm long; fairly broad, width less than half the length. Vein C extended to apex of M1, though obscured because tip of wing is folded; Sc present but very incomplete and faint; R1 virtually straight; R2+3 parallels R1, curved costad only at very apex; fork of R4 and R5 widely divergent, nearly at right angle; crossveins bm-cu slightly oblique to r-m (not parallel) and proximal to r-m by distance equal to length of r-m; cell dm and crossvein dm-cu present, cell dm large; vein M2 complete, closing cell dm with crossvein dm-cu; CuA1 complete; CuA2 present, complete, forming cell cu p, with incomplete “spur” of veins A1[+CuA2?] originating from cell cu p. Anal lobe of wing well developed. Legs long, bristly, with row of stiff setae on ventral margin of hind tibia. ABDOMEN: Very folded and not visible in dorsal view, unclear in lateral view.

Type. Sex? (tip of abdomen not preserved), Holotype: SPAIN: ÁLAVA , near Peñacerrada, Escucha Formation, Lower Cretaceous; MCNA 8882.

Etymology. From the Latin, concilio, unite separate parts into a whole, connect; in reference to the plesiomorphic features of this genus that link Chimeromyia with other basal Eremoneura.

Comments. See discussion of phylogenetic relationships, below.

Figures

FIGURE 13. Chimeromyina concilia, n. gen., n. sp., in Spanish amber. Holotype, MCNA 8882. A. Dorsal view of anterior half of body. B. Detail of antennae and vertex of head. C. Left habitus (terminalia lost at surface of amber).