Eupines (Eupines) undecim, Shen & Leschen, 2020

Shen, Jia-Wei & Leschen, Richard A. B., 2020, Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae), Zootaxa 4777 (1), pp. 1-84 : 73

Eupines (Eupines) undecim sp. nov.

Fig. 48 ; map Fig. 54

Diagnosis: Head with vertexal foveae punctiform, surrounded by shallow impressions; antennae 11-segmented with 2-segmented antennal clubs; A4, A5, A6, A7, A8 and A9 unmodified, A10 trapezoidal, about twice length of A9, dorsal side truncate, A11 longest, spear-shaped, with basolateral excavation, basal margin truncate at dorsal side. Abdominal ventrite 2 unmodified; ventrite 6 with pair of long and acute median struts, about one-third of ventral length. Protrochanters with short ventral spine, about one-fifth of protrochanteral length; protibiae and mesotibiae unmodified.

Description: Body length 1.25–1.33 mm; body colour reddish brown with lighter coloured tibiae and tarsi. Head wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes moderately developed, prominent in dorsal view, with about 16 facets; vertexal foveae punctiform, surrounded by shallow impressions; antennal tubercles moderately developed; postantennal notches distinctive and broad; postantennal cavities large, slightly shorter than antennal tubercles in dorsal view; antennae 11-segmented with 2-segmented antennal clubs; A4, A5, A6, A7, A8 and A9 unmodified, A10 trapezoidal, about twice length of A9, dorsal side truncate, A11 longest, spear-shaped, with basolateral excavation, basal margin truncate at dorsal side. Pronotum as long as wide, about as wide as head at eye level; lacking lateral antebasal foveae. Elytra slightly wider than long, widest near middle; lacking humeral calli and sutural stria. Abdomen with tergite 1 about 1.2 × longer than tergite 2; ventrite 2 unmodified; ventrite 6 with pair of long and acute median struts, about one-third of ventral length. Legs with protrochanters modified with short ventral spine, about one-fifth of protrochanteral length; protibiae and mesotibiae unmodified. Aedeagus about 0.22 mm long, with stout and symmetrical parameres, apex curved downwards in lateral view, with two pairs of long subapical setae and pair of long apical setae; median lobe with two large and symmetrical sclerites, curved towards each other in dorsal view, widest at base and narrower than parameres.

Comments: This is the only New Zealand species that can be placed into the subgenus Eupines by having an 11-segmented antenna. Ventrite 2 of E. (E.) undecim is unmodified, as it is in E. novum sp. n., while it is modified in all species of the subgenus Byraxis , apart from E. (B.) nemoralis ( Fig. 36F ).

Distribution: North Island. ND.

Etymology: This specific epithet means ‘eleven’ in Latin, referring to the 11-segmented antennae of this species.

Type material ( NZAC): Holotype. ♂, specimen entire, card-mounted: “ NEW ZEALAND, ND, Mangamuka Summit , 400 m, 13 Dec 1976, B.M. May litter 76/108”. Paratypes . 9 ex.: ND: 1, Mangamuka Summit , 400 m, 13/Dec/1976, V.A. May, litter, 76/106; 3, Mangamuka Summit, 400 m, 13/Dec/1976, V.A. May, litter, 76/109; 4, Mangamuka Gorge, 30/Nov/1907 // A. E. Brookes Collection; 1, Ngaiotonga Reserve, 03/Nov/1981, G. Kuschel // litter and decayed wood, 81/119 .

Figures

FIGURE 36. Diagnostic characters of E. (B.) nemoralis (Broun, 1886). A) Habitus. B) Antenna, in dorsal view. C) Same, in lateral view. D) Same, in ventral view. E) Protrochanter. F) Ventrite 2. G) Ventrite 6. H) Aedeagus, in dorsal view. I) Same, in lateral view. J) Same, in ventral view. Scale bars: A = 1 mm, B–F = 0.2 mm, G–J = 0.1 mm.

FIGURE 48. Diagnostic characters of E. (B.) undecim sp. n. A) Habitus. B) Antenna, in dorsal view. C) Same, in lateral view. D) Protrochanter. E) Ventrite 6. F) Aedeagus, in dorsal view. G) Same, in ventral view. H) Same, in lateral view. Scale bars: A = 1 mm, B–D = 0.2 mm, E–H = 0.1 mm.

FIGURE 54. Geographic distributions of E. (B.) pannicula sp. n., E. (B.) petila sp. n., E. (B.) platyarthra (Broun, 1893), E. (B.) protibialis sp. n., E. (B.) setifera (Broun, 1893), E. (B.) waikaremoana sp. n., E. (B.) whirinaki sp. n. and E. (B.) undecim sp. n. in New Zealand.