Horaeomorphus wailimae (Lhoste) Jałoszyński, 2014

Jałoszyński, Paweł, 2014, Oriental Horaeomorphus Schaufuss: a new species, synonymic notes and distributional synopsis (Coleoptera: Staphylinidae: Scydmaeninae), Raffles Bulletin of Zoology 62, pp. 83-88 : 85

Horaeomorphus wailimae (Lhoste) , new combination

( Fig. 1 D–G)

Stenichnus (Cyrtoscydmus) wailimae Lhoste, 1939: 1 , figs. 1–2; Franz, 1970: 546, fig. 11.

Stenichnus waihinae Lhoste (misspelling in Franz, 1970, legend to fig. 11).

Horaeomorphus (Pseudosyndicus) heissi Franz, 1985: 124 , fig. 36, new synonym.

Horaeomorphus heissi Franz ; Jałoszyński, 2006: 39, figs. 1, 8, 22, 50, 83, 86–89, 124.

Material examined. Lectotype (here designated). Male, drymounted beetle ( Fig. 1D ) and genital preparation on microscope slide ( Fig. 1F ), card-mounted specimen with five labels ( Fig. 1G ): “Wai Lima Z. Sum. / Lampongs [i.e., Lampung] No 156 / Karny. XI–XII.1921” [white, printed except for handwritten number], “ Stenichnus / wailimae / J. LHOSTE det. n. sp.” [white, handwritten and printed], “ Holotypus ” [red, printed], “Lhoste det.” [white, printed], “ HORAEOMORPHUS / wailimae ( Lhoste, 1939) / LECTOTYPUS / det. P. JAŁOSZYŃSKI, 2013 [white, printed] ( SDEI).

Paralectotypes. 1 male and 1 female ( Fig. 1E ), same data except for “ Paratypus ” and “ Paralectotypus ” labels ( SDEI) .

Remarks. Lhoste (1939) did not specify the number of individuals included in the type series, therefore the three specimens preserved at SDEI have the status of syntypes. In order to ensure the stability of nomenclature and provide a unique name-bearing type for Stenichnus wailimae a lectotype is here designated.

Franz (1970) redescribed and illustrated the aedeagus of Stenichnus wailimae and did not express any doubts concerning the generic placement of this species. Later Franz (1985) described Horaeomorphus heissi from Sumatra, which was redescribed by Jałoszyński (2006a). In the latter study, which is a comprehensive revision of Horaeomorphus from Malaysia, Singapore, Indonesia, and Brunei, the syntypes of Stenichnus wailimae were not studied, as it seemed unlikely that Franz, who himself published a number of papers on Stenichnus and Horaeomorphus , might have misidentified these genera. However, examination of the type series of Stenichnus wailimae carried out during the present study revealed that this species not only belongs to Horaeomorphus , but is also conspecific with H. heissi .

Horaeomorphus wailimae (= H. heissi ) can be easily identified on the basis of a hook-like projection on the ventral surface of male metafemora ( Fig. 1D , marked with an arrow), missing in females ( Fig. 1E ), non-modified metatrochanters in both sexes, three ante-basal pits on the pronotum, of which the middle one is anteriorly connected to a short longitudinal groove, and a remarkable and unique aedeagus (illustrated in Jałoszyński, 2006a, Figs. 86–89). The only other known species of Horaeomorphus with the hook-like metafemoral projection is H. samosirensis Jałoszyński, 2009 from the Samosir Island, North Sumatra, but it has the pronotum nearly circular and nearly as broad as elytra, and not strongly elongate and much narrower than elytra, as in H. wailimae .

Figures

Fig. 1. Horaeomorphus solodovnikovi, new species, (A–C); and Horaeomorphus wailimae (Lhoste), new combination, (D–G). Dorsal habitus of holotype male (A), lectotype male (D; arrow indicates femoral projection) and paralectotype female (E); right metatrochanter and femur in ventral view (B; arrow indicates the apex of trochanteral projection); internal armature of aedeagus in ventral view (C; arrow indicates median tubular structure); genital preparation of lectotype male (F); and original set of labels of lectotype male (G). Scale bars = 0.5 mm (A, D, E), 0.1 mm (B), 0.2 mm (C).