Landouria menorehensis, Nurinsiyah & Neiber & Hausdorf, 2019

Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, European Journal of Taxonomy 526, pp. 1-73 : 66-67

publication ID	https://doi.org/ 10.5852/ejt.2019.526
publication LSID	lsid:zoobank.org:pub:3FFC527D-E9CA-4D9F-BF59-076FA60171AC
DOI	https://doi.org/10.5281/zenodo.5587354
persistent identifier	https://treatment.plazi.org/id/E09F6255-8F7A-49E7-A3BC-E5B795EE2015
taxon LSID	lsid:zoobank.org:act:E09F6255-8F7A-49E7-A3BC-E5B795EE2015
treatment provided by	Plazi
scientific name	Landouria menorehensis
status	sp. nov.

Treatment

Landouria menorehensis View in CoL sp. nov.

urn:lsid:zoobank.org:act:E09F6255-8F7A-49E7-A3BC-E5B795EE2015

Figs 47 View Figs 29–47 , 55 View Figs 52–55 , 71 View Figs 63–71 , 88 View Figs 85–88 ; Tables 3–4 View Table 3 View Table 4

Diagnosis

Landouria menorehensis sp. nov. is characterized by a whitish, globose shell with a distinct brown band above the rounded periphery and a moderately narrow umbilicus.

Etymology

Landouria menorehensis sp. nov. is named after its occurrence in the Menoreh karst area.

Material examined

Holotype

INDONESIA • holotype (det. anat.); Yogyakarta, Kulonprogo, Sibolong , agroforestry; 7°44′52″ S, 110°08′12″ E; 687 m a.s.l.; 8 Jan. 2013; A. Nurinsiyah leg.; measurements: D = 13.4 mm, H = 8.6 mm; MZB 19201. GoogleMaps

Paratypes

INDONESIA – Central Java, Yogyakarta • 1 spec.; Kulonprogo, Sibolong, agroforestry along the way to Sibolong Hill ; 7°45′00″ S, 110°07′00″ E; 721 m a.s.l.; ZMH 133441 View Materials GoogleMaps • 2 spec.; same data as for preceding; ZMH 133442 View Materials GoogleMaps • 1 spec.; same data as for preceding; ZMH 133443 View Materials GoogleMaps • 12 spec. (det. anat.); Kulonprogo, Sibolong , agroforestry; 7°44′44″ S, 110°08′02″ E; 716 m a.s.l.; MZB 17686 GoogleMaps • 9 spec.; same data as for preceding; MZB 17702 GoogleMaps • 5 spec.; same data as for preceding; ZMH 133311 View Materials GoogleMaps • 1 spec. (det. anat.); Kulonprogo, Sibolong , snakefruit plantation; 7°44′44″ S, 110°08′02″ E; 716 m a.s.l.; MZB 17724 GoogleMaps • 1 spec.; same data as for preceding; MZB 17725 GoogleMaps • 2 spec. (det. anat.); Kulonprogo, Sibolong, near Kiskendo Cave , shrubs; 7°44′50″ S, 110°08′09″ E; 721 m a.s.l.; MZB 17644 GoogleMaps • 1 spec.; same data as for preceding; ZMH 133444 View Materials GoogleMaps • 6 spec.; same data as for holotype; MZB 19202 GoogleMaps • 3 spec.; same data as for holotype; ZMH 133439 View Materials GoogleMaps • 1 spec.; same data as for holotype; ZMH 133440 View Materials GoogleMaps • 2 spec.; Kulonprogo, Sibolong , agroforestry; 7°44′51″ S, 110°08′12″ E; 673 m a.s.l.; MZB 18039 GoogleMaps • 1 spec.; same data as for preceding; ZMH 133445 View Materials GoogleMaps • 2 spec.; Kulonprogo, Jonggrangan ; 7°44′20″ S, 110°08′32″ E; 730 m a.s.l.; ZMH 140034 View Materials GoogleMaps .

Description

SHELL ( Fig. 88 View Figs 85–88 ; Tables 3–4 View Table 3 View Table 4 ). Slightly depressed conical-globose, with 5.25–5.75 convex whorls; protoconch smooth; teleoconch with fine, irregular wrinkles; apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; without scaly processes; whitish with a distinct brown band above periphery; body whorl rounded; aperture almost circular; upper insertion of peristome distinctly descending; peristome expanded, reflexed and thickened; umbilicus slightly eccentric, moderately narrow, comprising 16–24% of shell diameter, hardly obscured by columellar edge.

GENITALIA ( Figs 47 View Figs 29–47 , 71 View Figs 63–71 ; Table 4 View Table 4 ). Atrium short; penis short, distal part cylindrical, proximal part slightly dilated, without penis coecum; membranaceous sheath around penis connected with proximal part of epiphallus by tissue; penial retractor runs from diaphragm to distal part of short epiphallus; flagellum long, broadest in its middle part, almost smooth, without node, tapering towards its proximal end; vas deferens narrow and long; vagina long; oviduct short; peduncle of bursa copulatrix subdivided into a short, broad distal part and a narrow and long proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.

Remarks

Landouria menorehensis sp. nov. shares with L. pakidulan sp. nov. and L. ciliocincta the whitish shell with a brown band at the periphery. For differences, see remarks under L. pakidulan sp. nov.

Distribution

Landouria menorehensis sp. nov. is endemic to the Menoreh karst area ( Fig. 55) View Figs 52–55 .

Figures

Figs 29–47. Flagella of species of Landouria from Java. 29. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 30. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 31. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 32. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 33. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 34. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 35. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 36. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 37. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 38. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). 39. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 40. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 41. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 42. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 43. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 44. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 45. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 46. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 47. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bar: 1 mm.