Manota montivaga, Jaschhof & Hippa, 2005

Jaschhof, Mathias & Hippa, Heikki, 2005, The genus Manota in Costa Rica (Diptera: Mycetophilidae), Zootaxa 1011 (1), pp. 1-54 : 29

Manota montivaga sp. n. ( Fig. 14 )

Description. Large species.

Male. Head. Antenna with fourth flagellomere 1.9 times as long as wide. Maxillary palpus with third palpomere bearing 3 curved sensilla; fourth palpomere with a parasegment apically.

Thorax. Anterior basalare and laterotergite each bare. Episternum 3 with ventral portion setose. Wing. Length 2.2–2.6 mm. Membrane with a few setae along posterior margin. Sc usually bare, occasionally with 1 seta ventrally. CuA­fork complete. Presence of A1 occasionally indicated by 2 or so setae in line between CuA and A2.

Terminalia ( Fig. 14 ). Sternite 9 with only posterior margin distinct; margin emarginate medially, situated below mid length of gonocoxites. Ventro­lateral lobes of gonocoxites subrectangular; interiorly with 1 strong, curved seta on a separate rim. Disto­lateral lobes of gonocoxites large, bilobed; on dorsal lobe with large, straight setae pointing posteriorly; on ventral lobe with smaller, curly setae pointing inwards. Gonocoxites in position I with 1 large megaseta hook­like apically; in position II with 4 blunt megasetae on lobe; in position III with 2 flat megasetae with one megaseta pointed and the other blunt; in position IV with 1 flat, spoon­shaped megaseta. Gonostyli small, flat; ventro­apically with 1 large and 1 smaller setae, followed dorsally by 3 curved and 1 straight setae, most dorsally with 1 seta strongly re­curved. Parastylar lobes hidden behind gonostyli, large, very weak; each with a marginal comb of some 15 trichia (not illustrated). Tegmen long and narrow, almost evenly tapering to tip. Hypoproct rounded apically; with long setae laterally.

Diagnostic characters. Among the species with both anterior basalare and laterotergites bare, it has the terminalia with gonocoxites bearing a quite distinct set of megasetae in positions I to IV, including 1 peculiar hooked megaseta in position I.

Etymology. From Latin, montivagus, mountain­roaming, referring to the occurrence of this species in high­altitude mountains.

Types. Holotype. Male, Costa Rica, Cartago province, El Guarco, Rio Macho Forest Reserve , Estación Ojo de Agua , 2950–3000 m, 22–28 July 1999, by Malaise trap, A. Picado ( INBio sample #52878) . Paratypes. 2 males, Cartago province, Madreselva, Finca los Lagos , 2000–2600 m, April–May 1994, by Malaise trap, M.M. Chavarria & A. Solano ( INBio sample #5494) ; 1 male, San José province, Pérez Zelodón, Chirripó National Park, Llano Bonito , 2400 m, 14 Sep.–14 Oct. 2001, by Malaise trap, M. Alfaro .

Figures

FIGURE 14. Manota montivaga sp. n. A: Male terminalia, ventral view (holotype). B: Male terminalia, central portion, ventral view (holotype). C: Male terminalia, dorsal view (paratype from Madreselva). Scale = 0.1 mm. 1 = posterior margin of sternite 9, 2 = tergite 9, 3 = disto­lateral lobe of gonocoxites, 4 = ventro­lateral lobe of gonocoxites, 5 = dorsal margin of gonocoxites, 6 = gonocoxal apodeme, 7 = gonostylus, 8 = ejaculatory apodeme, 9 = tegmen, 10 = parameral apodeme, 11 = apical process of tegmen, 12 = ventro­lateral lobe of tegmen, 13 = hypoproct (sternite 10), 14 = cercus.