Onthophagus (Parascatonomus) accedens accedens Lansberge, 1883

Krikken, J. & Huijbregts, J., 2008, Two new taxa from the Lesser Sundas in the genus Onthophagus (Coleoptera: Scarabaeidae: Scarabaeinae), Journal of Natural History 42 (43 - 44), pp. 2735-2746 : 2744-2745

Onthophagus (Parascatonomus) accedens accedens Lansberge, 1883

( Figures 15–17 , 23 )

Diagnostic remarks

Length 10.5–16.0 mm. Nominate O. accedens differ from nusatenggaricus in having the anterior clypeal border distinctly emarginate-sinuate on either side of the median tooth. On average, their body size is larger. Median vertexal protrusion very low. Major males with slight transversely arcuate crest on anterior side of pronotum. Black, occasionally brown (the holotype is brown), head and pronotum never cupreous; elytra matt, microreticulate, and finely punctate. Parameres rather similar to nusatenggaricus, parts have different proportions ( Figure 23 ). See also previous subgroup diagnosis.

Measurements in mm of specimen figured (Java). Maximum width of head 4.8. Median length of pronotum 6.1, maximum width 8.4. Sutural length of elytra 6.0, maximum width combined 8.9. Body length (head deflexed) slightly over 13 mm.

Comments

The brown female holotype is from Bandung on Java (the place was not given in the original description, but is clear from the collection event label with the type, in Leiden), and agrees with all the series at hand, from west to east Java and from Bali. Most records with ecological data give upland sites and an association with carrion; no records from Central Java. Boucomont (1914) mentioned Lombok , but his record may pertain to nusatenggaricus as defined above; a sympatric occurrence of both forms may be indicative of full species-level status of O. accedens and nusatenggaricus, respectively .

Material examined

New extralimital data (Java) are given here to settle the limits of the nominate subspecies versus nusatenggaricus.

Bali: Belimbing: Pura Mekori Forest, 27–31 October 1991, Krikken, Huijbregts and De Vries, #lk41a, 630 m, multistratal evergreen forest (remnant), human excrement trap, 1 ex.; Belimbing: Pura Mekori Forest , 27–31 October 1991, Krikken, Huijbregts and De Vries, #lk41b, 630 m, multistratal evergreen forest (remnant), fish trap, 14 exx.; Mt Batukau : Pura Luhur Forest , 29–31 October 1991, Krikken, Huijbregts and De Vries, #lk42b, 800 m, multistratal evergreen forest (nr river), fish trap, 7 exx.; Mt Batukau : Pura Luhur Forest , 29–31 October 1991, Krikken, Huijbregts and De Vries, #lk43a, 800 m, multistratal evergreen forest, human excrement trap, 1 ex.; Mt Batukau : Pura Luhur Forest , 29–31 October 1991, Krikken, Huijbregts and De Vries, #lk43b, 800 m, multistratal evergreen forest, fish trap, 2 exx. Subtotal 25 males and females, from Bali, in RMNH .

Java unspecified, 2 exx. West Java: Holotype female from ‘‘ Sijthoff \ Bandong’ ’ [round label, handwritten]; Garut and envs, February–April,1931, 700– 1000m, Van Heurn, 2 exx., Garut, February 1929, Van Heurn 1 ex., same but March–April, 1929, 1 ex., April 1929, 12 exx.; Cipayung, 10 July 1976, lamp, 1 ex. [there are different places of that name S of Bogor]; Mt Salak, N slope, 900m, 28–31 December 1985, Krikken, #pw72a, multistratal evergreen forest (degraded), human excr trap, 1 ex., same data but #pw72b, fish trap, 22 exx. East Java: Mt Kawi, July1934, Van Doesburg, 5 exx., August 1934, Van Doesburg, 10 exx.; Mt Semeru , 3000ft, A Koller, 1 ex.; Lawang, 1897, ex coll. Frühstorfer; all this Javan material in RMNH. Idjen: Blawan, 5 exx. in ZMA. Subtotal 66 males and females, from Java, in RMNH, ZMAN.

Total 91 males and females, in RMNH, ZMAN .

Figures

Figures 9–20. Onthophagus dux subgroup. (9–14) O. accedens nusatenggaricus, male holotype; (15–17) accedens accedens, male W Java; (18–20) dux, male Sabah. (9) habitus, oblique; (10, 15, 18) head, full-face; (11, 16, 19) pronotum, dorsal; (12, 17, 20) elytron, dorsal. (13) protibia, upperside; (14) metatibia and distal part of femur, underside. Scale with Figure18 51mm.

Figures 21–24. Aedeagus of Onthophagus species. (A) Upper side; (B) lateral; (21) O. montishannoniae, holotype; (22) accedens nusatenggaricus, holotype; (23) accedens accedens, Bali; (24) dux, Sabah.