Philonthus pulcher, Wendeler, 1924

Taphylininae, Iochini, Aorothiini, Thiini, Latyprosopini, Taphylinini, Mblyopinina, Nisolinina, Yptiomina, Hilonthina & Herman, Lee H., 2001, Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina), Bulletin of the American Museum of Natural History 2441 (265), pp. 2435-3021 : 2698-2703

pulcher Wendeler, 1924 , see: quadripunctatus Bernhauer, 1915 .

quadripunctatus Bernhauer, 1915 g: 193 ( Leucitus ; Type locality: Neu-Guinea: Sattelberg).

— Scheerpeltz, 1933: 1380 ( Leucitus ; catalog).

— Cameron, 1937a: 104 ( Leucitus ; New Guinea).

— Last, 1975a: 1 ( Leucitus ).

— Last, 1975a: 3 ( Leucitus ; characters).

— Last, 1987: 55 ( Leucitus ; Papua New Guinea; Irian).

- DISTRIBUTION: New Guinea.

pulcher Wendeler, 1924: 340 ( Leucitus ; Type locality: Neu-Pommern, Süd-See).

— Scheerpeltz, 1933: 1380 ( Leucitus ; catalog).

— Last, 1975a: 1 ( Leucitus ; synonym of quadripunctatus ).

rufipennis Cameron, 1918 , see: Hesperus .

semichalceus Heller, 1910: 9 ( Leucitus ; Type locality: Papua New Guinea, 780–900 m).

— Bernhauer and Schubert, 1914: 373 ( Leucitus ; catalog).

— Last, 1975a: 5 ( Leucitus ; characters).

- DISTRIBUTION: New Guinea.

somoruensis Last, 1968: 360 ( Philonthus ; Type locality: New Guinea: Mobutie Somoru).

— Last, 1975a: 1, 4 ( Leucitus ; characters).

- DISTRIBUTION: New Guinea.

stenoides Gravenhorst, 1806: 90 ( Staphylinus ; Type locality: Iauae).

— Erichson, 1840: 493, 930 ( Philonthus ; Java).

— Fauvel, 1878f: 253 ( Leucitus ; characters; Java).

— Bernhauer and Schubert, 1914: 373 ( Leucitus ; catalog).

— Last, 1975a: 1 ( Leucitus ).

— Schillhammer, 1999b: 95 ( Leucitus ; based on original description, species definitely belongs in Leucitus ; type locality in Java probably erroneous; species probably from New Guinea; type specimen probably lost).

- DISTRIBUTION: Indonesia.

superbus Arrow, 1915: 499 ( Philonthus ; Type locality: Utakwa , River, Dutch New Guinea).

— Scheerpeltz, 1933: 1363 ( Philonthus ; catalog).

— Last, 1975a: 1, 6 ( Leucitus ; characters).

- DISTRIBUTION: New Guinea.

Linoderus

[1 species; Neotropical Region]

Linoderus Sharp, 1885: 452 (species included: gracilipes ). TYPE SPECIES: Linoderus gracilipes Sharp , fixed by monotypy.

— Bernhauer and Schubert, 1914: 367 (world catalog; 1 species).

— Blackwelder, 1944: 136 (checklist of species from Mexico, Central and South America, and

the West Indies).

— Blackwelder, 1952: 221 (type species: gracilipes ).

gracilipes Sharp, 1885: 452 ( Linoderus ; Type locality: Panama, Volcan de Chiriqui, Bugaba). — Bernhauer and Schubert, 1914: 367 ( Linoderus ; catalog).

- DISTRIBUTION: Panama.

Mentophilonthus

[8 species; Ethiopian Region]

Mentophilonthus Levasseur, 1966: 207 (species included: odzalaensis , triseriatus , descarpentriesi , mongendensis , dilutior , lampropterus ). TYPE SPECIES: Philonthus triseriatus Bernhauer , fixed by original designation.

bangoranensis Levasseur, 1980: 354 ( Mentophilonthus ; Type locality: République Centrafricaine: Bangoran , SW de Ndélé).

- DISTRIBUTION: Central African Republic.

centrafricanus Levasseur, 1980: 356 ( Mentophilonthus ; Type locality: République Centrafricaine: Bangoran , SW de Ndélé).

- DISTRIBUTION: Central African Republic.

descarpentriesi Levasseur, 1966: 208 , 209 ( Mentophilonthus ; Type locality: Congo ex français, Odzala. Also cited from Belgian Congo).

— Cameron, 1929: 61 ( Philonthus ; [Note: Misidentified as triseriatus according to Levasseur, 1966: 209]; characters; Belgian Congo).

— Tottenham, 1949b: 300 ( Philonthus ; [Note: Misidentified as triseriatus according to Levasseur, 1966: 209]; characters; Belgian Congo).

— Levasseur, 1966a: 1211 ( Mentophilonthus ; Belgian Congo; Congo).

- DISTRIBUTION: Congo Republic, Congo.

dilutior Bernhauer, 1928 g: 113 ( Philonthus ; Type locality: Belgisch-Kongo: Mongende).

— Scheerpeltz, 1933: 1340 ( Philonthus ; catalog).

— Levasseur, 1966: 208, 211 ( Mentophilonthus ; characters; Belgian Congo).

- DISTRIBUTION: Congo.

seriatipennis Bernhauer, 1928 g: 114 ( Philonthus ; Type locality: Belgisch-Kongo: Mongende).

— Cameron, 1929: 61 ( Philonthus ; characters; Belgian Congo).

— Scheerpeltz, 1933: 1360 ( Philonthus ; catalog).

— Tottenham, 1949b: 300 ( Philonthus ; characters).

— Levasseur, 1966: 211 ( Mentophilonthus ; synonym of dilutior ).

lampropterus Bernhauer, 1934 g: 239 ( Philonthus ; Type locality: Kivu: Kibumba ).

— Cameron, 1950g: 35 ( Philonthus ; Belgian Congo).

— Levasseur, 1966: 213 ( Mentophilonthus ; characters; Belgian Congo).

- DISTRIBUTION: Congo.

minor Bernhauer, 1928 , see: triseriatus Bernhauer, 1928 .

mongendensis Bernhauer, 1928 g: 114 ( Philonthus ; Type locality: Belgisch-Kongo: Mongende). — Scheerpeltz, 1933: 1351 ( Philonthus ; catalog).

— Levasseur, 1966: 208, 211 ( Mentophilonthus ; characters; Belgian Congo).

- DISTRIBUTION: Congo.

odzalaensis Levasseur, 1966: 208 , 211 ( Mentophilonthus ; Type locality: Odzala , Congo ex française. Also cited from Gabon).

— Levasseur, 1966a: 1211 ( Mentophilonthus ; Congo; Zaire; Gabon).

— Levasseur, 1968a: 53 ( Mentophilonthus ; Central African Republic).

- DISTRIBUTION: Gabon, Congo Republic, Congo, Central African Republic.

parcior Bernhauer, 1928 , see: triseriatus Bernhauer, 1928 .

seriatipennis Bernhauer, 1928 , see: dilutior Bernhauer, 1928 .

triseriatus Bernhauer, 1928 g: 112 ( Philonthus ; Type locality: Belgisch-Kongo: Mongende).

— Scheerpeltz, 1933: 1365 ( Philonthus ; catalog).

— Levasseur, 1966: 208 ( Mentophilonthus ; characters; Congo; Ruanda).

— Levasseur, 1966a: 1210 ( Mentophilonthus ; Congo; Zaire).

- DISTRIBUTION: Congo Republic, Congo, Rwanda.

minor Bernhauer, 1928 g: 113 ( Philonthus ; [preoccupied]; Type locality: Belgisch-Kongo: Mongende).

— Scheerpeltz, 1933: 1365 ( Philonthus ; variety of triseriatus ).

parcior Bernhauer, 1928 g: 113 ( Philonthus ; Type locality: Belgisch-Kongo: Mongende).

— Scheerpeltz, 1933: 1365 ( Philonthus ; variety of triseriatus ).

Moeocerus

[13 species; Ethiopian Region]

Moeocerus Fauvel, 1899: 100 (replacement name for Homoeocerus Fauvel ). TYPE SPECIES: Homoeocerus mimus Fauvel , fixed by subsequent designation Blackwelder, 1952: 250.

— Bernhauer and Schubert, 1916: 410 (world catalog; 7 species).

— Scheerpeltz, 1933: 1420 (world catalog supplement).

— Gridelli, 1939: 322 (characters; key to species).

Homoeocerus Fauvel, 1899a: 27 (species included: mimus , simplex , ciliatus , robustus , spinulosus , puncticollis ; preoccupied by Burmeister, 1835, and Kolenati, 1859). TYPE SPECIES: Homoeocerus mimus Fauvel by objective synonym with Moeocerus for which Blackwelder fixed the type species. [Note: Blackwelder stated that R. Lucas, 1920: 421, designated the type species but the designation was ambiguous].

— Fauvel, 1907: 48 (synonym of Moeocerus ).

— Bernhauer and Schubert, 1916: 410 (synonym of Moeocerus ).

— Blackwelder, 1952: 190 (type species; synonym of Moeocerus ).

Polystictomoeocerus Scheerpeltz, 1977: 145 (subgenus of Moeocerus ; species included: puncticollis , griseosericans ). TYPE SPECIES: Homoeocerus puncticollis Kraatz , fixed by original designation.

abessinus Bernhauer, 1917 g: 46 ( Moeocerus ; Type locality: Abessinien: Harrar ).

— Scheerpeltz, 1933: 1420 ( Moeocerus ; catalog).

— Gridelli, 1939: 326, 329 ( Moeocerus ; characters; Abyssinia).

— Scheerpeltz, 1977: 146 ( Moeocerus ; subgenus Moeocerus ).

- DISTRIBUTION: Ethiopia.

ciliatus Roth, 1851: 118 ( Acylophorus ; Type locality: Abyssinien).

— Fauvel, 1899a: 30 ( Homoeocerus ; characters; Tigre; Abyssinie).

— Bernhauer and Schubert, 1916: 411 ( Moeocerus ; catalog).

— Gridelli, 1939: 326 ( Moeocerus ; characters; Abyssinia).

— Scheerpeltz, 1977: 146 ( Moeocerus ; subgenus Moeocerus ).

- DISTRIBUTION: Ethiopia.

doriae Gridelli, 1939: 335 ( Moeocerus ; Type locality: Cheren , Eritrea; regione dell’Eritrea, Paese dei Bogos).

— Scheerpeltz, 1977: 146 ( Moeocerus ; subgenus Moeocerus ).

- DISTRIBUTION: Eritrea.

fauveli Gridelli, 1939: 337 ( Moeocerus ; Type locality: Maritzburg, Natal).

— Scheerpeltz, 1977: 146 ( Moeocerus ; subgenus Moeocerus ).

- DISTRIBUTION: South Africa.

griseosericans Scheerpeltz, 1977: 143 ( Moeocerus ; subgenus Polystictomoeocerus ; Type locality: Ed. Luja in der Umgebung von Kondué im Kongogebiet).

- DISTRIBUTION: Congo.

lineiventris Fauvel, 1907: 48 ( Moeocerus ; Type locality: Kilimandjaro: Kibosho , 1700 m).

— Bernhauer and Schubert, 1916: 411 ( Moeocerus ; catalog).

— Scheerpeltz, 1977: 146 ( Moeocerus ; subgenus Moeocerus ).

- DISTRIBUTION: Tanzania.

mimus Fauvel, 1899a: 29 ( Homoeocerus ; Type locality: Natal).

— Bernhauer and Schubert, 1916: 411 ( Moeocerus ; catalog).

— Gridelli, 1939: 326, 334 ( Moeocerus ; characters).

— Scheerpeltz, 1977: 146 ( Moeocerus ; subgenus Moeocerus ).

- DISTRIBUTION: South Africa.

overlaeti Bernhauer, 1935d: 104 ( Moeocerus ; Type locality: Lulua: Kapanga).

— Gridelli, 1939: 326 ( Moeocerus ; Abyssinia).

— Scheerpeltz, 1977: 146 ( Moeocerus ; subgenus Moeocerus ).

- DISTRIBUTION: Congo, Ethiopia.

puncticollis Kraatz, 1858: 363 ( Homorocerus ; Type locality: Senegambia).

— Fauvel, 1899a: 32 ( Homoeocerus ; Senegambie portugaise).

— Bernhauer and Schubert, 1916: 411 ( Moeocerus ; catalog).

— Gridelli, 1939: 325 ( Moeocerus ; Abyssinia).

— Tottenham, 1961: 205 ( Moeocerus ; Senegal).

— Scheerpeltz, 1977: 145 ( Moeocerus ; subgenus Polystictomoeocerus ).

- DISTRIBUTION: “Senegambia”, Senegal, Ethiopia.

robustus Gestro, 1881: 658 ( Homorocerus ; Type locality: Let Marefia).

— Fauvel, 1899a: 30 ( Homoeocerus ; characters; Abyssinie; Choa: Let Marefia; Giuba).

— Bernhauer and Schubert, 1916: 411 ( Moeocerus ; catalog).

— Gridelli, 1939: 330 ( Moeocerus ; characters; Abyssinia).

— Scheerpeltz, 1977: 146 ( Moeocerus ; subgenus Moeocerus ).

- DISTRIBUTION: Ethiopia.

simplex Fauvel, 1899a: 30 ( Homoeocerus ; Type locality: Angola).

— Bernhauer, 1915o: 363 ( Moeocerus ; Belgian Congo).

— Bernhauer and Schubert, 1916: 411 ( Moeocerus ; catalog).

— Scheerpeltz, 1977: 147 ( Moeocerus ; subgenus Moeocerus ).

- DISTRIBUTION: Angola, Congo.

spinulosus Solsky, 1864a: 375 ( Homorocerus ; Type locality: Insel Gorea an der Westküste

von Afrika).

— Fauvel, 1899a: 31 ( Homoeocerus ; characters; Natal; Abyssinia).

— Bernhauer and Schubert, 1916: 411 ( Moeocerus ; catalog).

— Bernhauer and Chapman, 1939: 73 ( Moeocerus ; Kenya).

— Scheerpeltz, 1977: 147 ( Moeocerus ; subgenus Moeocerus ).

- DISTRIBUTION: Senegal, Ethiopia, Kenya, South Africa.

zavattarii Gridelli, 1939: 332 ( Moeocerus ; Type locality: Moyale; Arero).

— Scheerpeltz, 1977: 147 ( Moeocerus ; subgenus Moeocerus ).

- DISTRIBUTION: Ethiopia.

Mysolius

[3 species; Oriental and Australian Regions]

Mysolius Fauvel, 1878f: 255 (species included: aurichalceus ). TYPE SPECIES: Mysolius aurichlaceus Fauvel , fixed by monotypy.

— Duvivier, 1883: 152 (catalog).

— Olliff, 1887: 497 (characters; Australian species).

— Bernhauer and Schubert, 1914: 373 (world catalog; 2 species).

— Blackwelder, 1952: 256 (type species: aurichalceus ).

aurichalceus Fauvel, 1878f: 256 ( Mysolius ; Type locality: Nouvelle-Guinée, Romoi;

Iles Arou, Wokan; Mysol; Salvatti).

— Fauvel, 1879: 97 ( Mysolius ; characters; Fly River).

— Bernhauer and Schubert, 1914: 373 ( Mysolius ; catalog).

— Cameron, 1924: 87 ( Philonthus ; New Guinea).

— Cameron, 1931: 363 ( Mysolius ; New Guinea).

— Last, 1987: 55 ( Mysolius ; Irian).

- DISTRIBUTION: New Guinea, Indonesia.

diversipes Fauvel, 1879: 98 ( Mysolius ; race of aurichalceus ; Type locality: Iles Arou; Nouvelle Guinée; Fly River).

— Bernhauer and Schubert, 1914: 373 ( Mysolius ; variety of aurichalceus ).

chalcopterus Olliff, 1887: 497 ( Mysolius ; Type locality: Mulgrave River, North Queensland). — Bernhauer and Schubert, 1914: 373 ( Mysolius ; catalog).

— Bernhauer, 1920: 16 ( Mysolius ; Australia).

— Last, 1987: 55 ( Mysolius ; Papua New Guinea).

- DISTRIBUTION: Australia, New Guinea.

cupreosplendens Cameron, 1943a: 344 ( Actinus ; Type locality: New Guinea: Milne Bay).

— Last, 1987: 55 ( Mysolius ; Papua New Guinea).

- DISTRIBUTION: New Guinea.

diversipes Fauvel, 1879 , see: aurichalceus Fauvel, 1878 .

Neobisnius

[72 species; Neotropical, Nearctic, Palaearctic, Ethiopian,

Madagascan, Oriental, and Australian Regions]

Neobisnius Ganglbauer, 1895: 464 (species included: villosulus ; procerulus ; prolixus ). TYPE SPECIES: Gabrius villosulus Stephens , fixed by subsequent designation by R. Lucas, 1920: 434.

— Thomson, 1859: 25 (cited as Bisnius ; [Note: Thomson’s use of Bisnius was a misidentification of Neobisnius ; Tottenham, 1949 cited it with a type species and as a synonym of Neobisnius as though it were an available name]; characters; type species: cephalotes ).

— Thomson, 1860: 168 (cited as Bisnius ; [Note: Thomson’s use of Bisnius was a misidentification of Neobisnius ; Tottenham, 1949 cited it with a type species and as a synonym of Neobisnius as though it were an available name]; characters).

— Heyden, 1880: 73 (cited as Actobius ; list of species of Siberian region).

— Heyden, Reitter, and Weise, 1906: 163 (list of species of Europe and Caucasus).

— Reitter, 1909: 116, 123 (characters; key to species).

— Petri, 1912: 71 (list of species of Siebenbürgen).

— Bernhauer and Schubert, 1914: 322 (world catalog; 39 species).

— Leng, 1920: 106 (catalog of North American species).

— Winkler, 1925: 374 (catalog for Palaearctic region).

— Cameron, 1925: 54 (catalog of Indian species).

— Porta, 1926: 99 (characters; key to species of Italy).

— Bruch, 1928a: 427 (catalog; Argentina).

— Portevin, 1929: 356 (key to French species).

— Roubal, 1930: 366 (catalog; Slovakia).

— Cameron, 1932: 56 (characters; key to species of British India).

— Joy, 1932: 126 (key to British species).

— Scheerpeltz, 1933: 1326 (world catalog supplement).

— Normand, 1935: 370 (list of species of Tunisia).

— Blackwelder, 1943: 440 (synonym of Erichsonius ; type species: cinerascens ).

— Gridelli, 1943: 111 (characters; notes; key to a few species).

— Kloet and Hincks, 1945: 165 (list of British species).

— Last, 1948: 148 (key to European species).

— Tottenham, 1949: 371, 417 (type species: villosulus Stephens = elongatulus Erichson ; checklist of British species).

— V. Hansen, 1952: 29 (characters; key to Danish species).

— Blackwelder, 1952: 258 (type species).

— Hatch, 1957: 173, 174 (characters; key to species of Pacific Northwest).

— Kocher, 1958: 142 (checklist of species; Morocco).

— Smetana, 1958: 134 (characters; habitus of one species; type species: villosulus ; key to Czechoslovakian species).

— Smetana, 1959a: 209 (list of species collected in Albania).

— Ferreira, 1962a: 17 (list of Portuguese species).

— Coiffait, 1963: 8, 18 (characters; key to French species groups and included French species).

— Székessy, 1963: 53 (characters).

— Palm, 1963b: 58 (characters; key to species of Sweden).

— Lohse, 1964: 171 (characters; key to central European species).

— Moore, 1965: 98, 100 (characters; notes).

— Coiffait and Saiz, 1968: 348, 360 (characters; notes).

— Scheerpeltz, 1968a: 64 (catalog of Austrian species).

— Shibata, 1973c: 56 (checklist of species of Taiwan).

— Tikhomirova, 1973a: 182 (checklist of species of USSR).

— Coiffait, 1974: 322 (characters; key to species of western Palaearctic region).

— Moore and Legner, 1975: 18 (catalog; North America north of Mexico).

— Pope, 1977: 29 (list of British species).

— Topp, 1978: 312 (larval characters).

— Moore and Legner, 1979: 24 (characters; notes).

— Muona, 1979: 14 (list of Scandinavian species).

— Uhlig, Vogel, and Sieber, 1980: 246 (some species collected in Germany).

— Burakowski, Mroczkowski, and Stefańska, 1980: 29 (catalog; Poland).

— Frank, 1981: 2 (revision of and key to New World species; characters; checklist of New World species; species group classification).

— Uhlig and Vogler, 1981: 110 (list of some species collected in Germany; habitat and collecting notes).

— Shibata, 1983: 88 (checklist of species of Japan).

— Hammond, 1984: 206 (checklist of species of Borneo).

— Tóth, 1984: 34 (characters; habitus; key to species of Hungary).

— Frank, 1986a: 370 (checklist of species of Florida).

— Segers, 1986: 46 (checklist of species; Belgium).

— Drugmand 1987b (comments on distribution of Belgian species).

— Campbell and Davies, 1991: 117 (checklist of species; Canada).

— Silfverberg, 1992: 17 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Smetana, 1993b: 47 (list of species of Czech Republic and Slovakia).

— Hernández, Outerelo, and Gamarra, 1994: 188 (checklist of species; Canary Islands).

— Smetana, 1995e: 50 (characters).

— Ciceroni and Zanetti, 1995: 27 (list of species of Italy).

— M. Hansen, 1996: 104 (list of Danish species).

— M. Hansen, Mahler, Palm, and Pedersen, 1996: 243 (additions to list of Danish species).

— Cho, 1996a: 159 (characters; key to species of Korea).

— Downie and Arnett, 1996: 374 (characters; key to species of Northeastern North America).

— Lundgren, 1998: 51 (list of species from Florida; records for other States listed for each species).

— Bain, 1998: 39 (17th century remains; Massachusetts).

— M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (list of species collected in 1997; Denmark).

— Newton, Thayer, Ashe, and Chandler, 2000: 396 (14 Nearctic species; notes; characters in key).

— Hayashi, 2000: 23 (notes on empodial setae).

adustus Sharp, 1887 , see: terminalis LeConte, 1863 .

afghanicus Coiffait, 1979d: 559 ( Neobisnius ; Type locality: entre Camdech et Kouchtous, 1400 à 2000 m).

— Tronquet, 1981: 71 ( Neobisnius ; Afghanistan).

— Coiffait, 1982b: 76 ( Neobisnius ; note).

- DISTRIBUTION: Afghanistan.

agnatus Erichson, 1840 , see: sobrinus Erichson, 1840 .

alaiensis Bernhauer, 1914b: 66 ( Neobisnius ; Type locality: Nord-Alai: Fergana; [Note: See lectotype designation by Gusarov, 1995a]).

— Scheerpeltz, 1933: 1326 ( Neobisnius ; catalog).

— Coiffait, 1974: 329 ( Neobisnius ; characters; Transcaspian region; Afghanistan).

— Gusarov, 1995a: 92 [= 1996: 55] ( Neobisnius ; lectotype designation).

- DISTRIBUTION: Uzbekistan, Afghanistan.

alternans Sharp, 1885 , see: jocosus Horn, 1884 .

angusticollis Kolenati, 1846 , see: prolixus Erichson, 1840 .

angustus Cho, 1996a: 162 ( Neobisnius ; Type locality: Korea: Chejudo , Kyoraeri).

- DISTRIBUTION: Korea.

arduus Sharp, 1885 , see: sobrinus Erichson, 1840 .

armuellensis Bierig, 1933: 56 ( Neobisnius ; Type locality: Panama: Puerti Armuelles ).

— Frank, 1981: 28 ( Neobisnius ; characters; Costa Rica; Panama; Colombia).

- DISTRIBUTION: Colombia, Panama, Costa Rica.

bakeri Bernhauer, 1916 h: 424 ( Neobisnius ; Type locality: Luzon : Mt. Makiling).

— Scheerpeltz, 1933: 1326 ( Neobisnius ; catalog).

- DISTRIBUTION: Philippines.