Podaena mariae Delgado and Palma, 2010
publication ID |
https://doi.org/ 10.11646/zootaxa.2678.1.1 |
publication LSID |
lsid:zoobank.org:pub:FB43BE83-EBB8-4D58-85CF-CBE73DD683FE |
DOI |
https://doi.org/10.5281/zenodo.5310472 |
persistent identifier |
https://treatment.plazi.org/id/6451D58F-1CFB-4F94-8109-82343848BE0B |
taxon LSID |
lsid:zoobank.org:act:6451D58F-1CFB-4F94-8109-82343848BE0B |
treatment provided by |
Felipe |
scientific name |
Podaena mariae Delgado and Palma |
status |
sp. nov. |
Podaena mariae Delgado and Palma View in CoL , new species
( Figs 19, 20 View FIGURES 1−24 , 27, 28 View FIGURES 25−30 , 49 View FIGURES 47−51 , 55, 59 View FIGURES 52−60 , 69, 70 View FIGURES 69−74 , 97 View FIGURES 95−98 , 103 View FIGURE 103 )
Description. Male as in Fig. 103 View FIGURE 103 .
Body length (taken from anterior margin of labrum to elytral apex) 2.30mm. Colour: head, pronotum and elytra dark brown to black; elytra uniformly coloured, without lighter marks on the shoulders; legs and palps dark reddish-brown.
Head prognathous, quite elongate, with a short neck region not retracted into the prothorax; evenly convex dorsally with rather fine but distinct punctures, smooth and glabrous; occipital region with the same micro-sculpture as on the dorsal region; postocular ridges present and prominent; clypeus long, 0.7 times as long as frons anterior to compound eyes, glabrous and very sparsely punctuated; frontoclypeal (epistomal) suture straight, not grooved and not deeply impressed; frons with slightly impressed paraocular reticulate areas, continuing towards the genae; compound eyes moderately protruding; labrum wider than long, bilobed, with a deep anteromedian incision; maxillary palps ( Figs 19−20 View FIGURES 1−24 ) with penultimate and distal palpomeres not modified (not grooved).
Thorax: Body elongate with distinct pronoto-elytral angle. Pronotum posteriorly constricted, with distinct pronoto-elytral angle; surface smooth but sparsely punctuated, with shallow anterior and posterior pits; anterior area, next to the neck, markedly convex, forming a gibbosity separated from the remainder of the pronotum by a clear transversal depression. Foretibiae ( Figs 27−28 View FIGURES 25−30 ) not modified, with tibial spines 1 and 2 less developed than in other species (e.g. P.obscura , Fig. 37 View FIGURES 31−38 ). Middle tibiae slightly curved in their distal half. Hind tibiae slightly curved inwards. Middle and hind tarsi rather long. Elytra long, 2.7 times the length of the thorax, with a smooth lateral rim and well defined rows (striae) of serial punctures, fading towards the distal area; a pronounced carina on the 7 th interstria, and a short distal carina on the 6 th interstria. Metaventrite with a subtriangular glabrous area on the disc, including a central, oval and shallow concavity not reaching the apex or the base of the glabrous area ( Fig. 49 View FIGURES 47−51 ).
Abdomen: Last tergite subtrapezoidal, with the distal margin emarginate bearing a tuft of setae of various lengths on each side, including a few strong peg-like setae ( Fig. 55 View FIGURES 52−60 ). Ventrite 7 with a long and firmly attached spiculum ( Fig. 59 View FIGURES 52−60 ). Genitalia ( Figs 69−70 View FIGURES 69−74 ) with a long sclerotized main piece; distal lobe long and wide with a sinuous upper margin on lateral view ( Fig. 69 View FIGURES 69−74 ); transparent ejaculatory pump associated to the everted ejaculatory duct, forming a long, thin, curved flagellum. Parameres long and slender, inserted close to phallobase, with sclerotized and slightly widen apices bearing a tuft of short setae.
Female unknown.
Differential diagnosis. Podaena mariae is the largest and most robust among all species in the genus, and also has the greatest elytron-length / thorax-width ratio of all species (2.7). The other large species, P. trochanteralis , has shorter elytra (elytron-length / thorax-width ratio = 2.5), longer middle and hind legs, especially the tarsi, and the middle and hind tibiae clearly curved inwards. The morphology of the maxillary palps ( Figs 19−20 View FIGURES 1−24 ) allows clear separation of P.mariae from all species with highly modified palps, including P. trochanteralis ( Figs 17−18 View FIGURES 1−24 ). Among the remaining species with slightly or not modified maxillary palps ( P. aotea , P. kuscheli , P. maclellani and P. glabriventris ), P. mariae can be distinguished from P. aotea and P. kuscheli by having slightly modified foretibiae ( Figs 27−28 View FIGURES 25−30 ) and a unique genitalia ( Figs 69−70 View FIGURES 69−74 ); and from P. maclellani and P. glabriventris by tergite X ( Fig. 55 View FIGURES 52−60 ), the glabrous area on the metaventrite ( Fig. 49 View FIGURES 47−51 ), and by the genitalia.
The male genitalia of P. mariae ( Figs 69−70 View FIGURES 69−74 ) are morphologically closest to those of P. maclellani ( Figs 71−72 View FIGURES 69−74 ). Both are well sclerotized in their distal halves; both have long thin parameres, and a permanently everted flagellum. The main difference is the thickness and shape of the flagellum: in P. mariae it is much thinner, curved, and lacks apical microdenticules (pruinosity), which are characteristic in P. maclellani . Furthermore, these two species have a very similar ventrite 7, both with a very long spiculum ( Figs 58−59 View FIGURES 52−60 ).
Etymology. The species epithet is a noun in the genitive case after Mrs María Martínez-Esparza, who assisted in fieldwork, including the collection of the paratype of this species.
Distribution. As in Fig. 97 View FIGURES 95−98 . Known only from two localities in the West Coast of the South Island.
Material examined. Holotype: South Island : Mackenzie ( MK): Male (dry-mounted) ( NZAC): “Mt Cook Nat. Pk. / Black Birch 820 m / 8 tm. 2 Apr. 1977 / J.S. Dugdale ”, “From 2 m / of side strm., funnel”.
Paratype: South Island : Westland ( WD): 1 male (dry-mounted) ( MONZ): “New Zealand, small stream / near Fox Glacier. Westland. / 4−12−1996 / J.A. Delgado & M. Martínez-E. leg” .
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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