Enghophyllum naxium ( Verhoeff, 1901 ) Lazányi & Vagalinski, 2013

Lazányi, Eszter & Vagalinski, Boyan, 2013, A new genus of the millipede tribe Brachyiulini (Diplopoda: Julida: Julidae) from the Aegean region, European Journal of Taxonomy 70, pp. 1-12 : 6-9

Enghophyllum naxium ( Verhoeff, 1901) comb. nov.

Figs 1E, G , J-K, 3, 4A-D, 5

Brachyiulus (Chromatoiulus) naxius Verhoeff, 1901: 98-99 , fig. 21.

Chromatoiulus naxius – Attems 1927: 222 (in key). — Strasser 1974: 290, 295. — Ceuca 1992: 425. Chromatoiulus (Diaxylus) naxius – Attems 1940: 307, 311-312.

Megaphyllum naxium – Mauriès & Karamaouna 1984: 55-56, figs 7-9. — Enghoff & Kime 2009. — Lazányi et al. 2012: 21 , 41.

Diagnosis

Differs from the only congener E. sifnium gen. et sp. nov. by the apical margin of the promere being concave, the opisthomere not compressed meso-laterad, e.g., the lateral process (lp on Fig. 4 A-D) parallel to the promere, the lateral process (lp) much broader and the apical posterior process (app) broader and blunter.

Material examined

GREECE: 1 ♂, 2 ♀♀, Antiparos Island, litière [litter], 22 Dec. 1982, leg. P. Beron & St. Andreev, det. Mauriès 1983 ( NMNHS); 1 ♂, Sporadhes, Anatoliko Mauri [east Mavri Islet] (Levitha Islands), 18.i.1990 [18 Jan. 1990], S. Sfendourakis leg., ex coll. M. Karamaouna ( ZMUC).

Remarks

Male body length 16.9-17.9 mm; height 1-1.2 mm; body ring number: 39-41 podous rings + 2-3 apodous rings + telson; number of ocelli: right side: 30-38, left side: 32-38; developmental stadium IX-X with

8-9 rows of ocelli. Female body length 13.4-18.3 mm; height 1-1.3 mm; body ring number: 37-41 podous rings + 3-5 apodous rings + telson; developmental stadium VIII-IX with 7-8 rows of ocelli.

This species was described on the basis of two males and four females and has only been found once since its description, viz., by Mauriès & Karamaouna (1984) from Antiparos. The new specimens from east Mavri belong to the collection of Karamaouna, but they are slightly different: the apical posterior process (app) of the solenomere is not bifurcated, but undivided and slightly serrated (serration visible in posterior view) ( Fig. 4A , C-D).

Distribution

Greece: Cyclades: Naxos Island (type locality), Antiparos Island ( Mauriès & Karamaouna 1984); Dodecanese Islands: east Mavri Islet.

Figures

Fig. 1. Non-gonopod characters of Enghopyhllum gen. nov., males.A. Right antenna.B. Gnathochilarium. C. 8th and 9th body rings. D. Telson. E. Subanal scale. F. Mandibular stipital (“cheek”) lobe, lateral view. G. Mandibular stipital (“cheek”) lobe, ventro-lateral view. H. First leg-pair. I. Second leg-pair. J. Penis in the penis sac. K. Pleurotergum of the 7th body ring, ventral view. L. Anterior left leg on the 6th body ring. M. Anterior right leg on the 22nd body ring. – A-D, F, H-I, L-M: E. sifnium gen. et sp. nov., holotype. E: E. naxium, from Antiparos. G, J-K: E. naxium, from Mavri Islet.

Fig. 4. Gonopods of Enghophyllum naxium, male from Mavri Islet. A. Gonopods, posterior view. B. Gonopods, antero-lateral view. C. Right gonopods, mesal view. D. Left opisthomere, lateral view. – app = apical posterior process; bpp = basal posterior process; d = deepening; fc = flagellum channel; lp = lateral process; msp = micro-spinose pillow; P = promere; r = ridge; ri = riffle; s = solenomere; sc = sperm canal; tl = triangular lamella. Scale bar: 0.2 mm.