Anthomyza shewelli, Roháćek & Barber, 2016

Roháćek, Jindřich & Barber, Kevin N., 2016, Nearctic Anthomyzidae: a monograph of Anthomyza and allied genera (Diptera), Acta Entomologica Musei Nationalis Pragae (suppl.) 56, pp. 1-412 : 279-296

Anthomyza shewelli sp. nov.

( Figs 2 , 456–458 , 460–481 , 485 , 508, 509 , 516)

Type material. HOLOTYPE: ♂, “CAN: ON: ~74kmNNE The-ssalon, shore of Mississagi R., 17.vii.2010, KNBarber, sweeps, graminoids, herbs, Equisetum spp. 46°53.94’N 83°16.23’W ” and “ Holotypus ♂ Anthomyza shewelli sp. n., J. Roháček & K. N. Barber det. 2014” (red). The specimen is in perfect condition, with exposed genitalia and well-exposed gonostyli (see Fig. 481 ) ( CNCI, intact). PARATYPES: CANADA: MANITOBA: Aweme, Criddle farm, spring nr. Assiniboine River, 49°42'N 99°36'W, sweep vegetation at spring, 13.vi.1999, 1 ♀, T. A. Wheeler leg. ( LEMQ 0039606); ~ 14 km SW Falcon Lake, jct. Hwy#1 & Rd.86E, 49°38.24'N 95°29.89'W, sweeps, graminoids, mostly Carex spp., 29.vii.2011, 1 ♂ 1 ♀, K. N. Barber leg.; 5 mi SW Shilo, 28.vi.1958, 1 ♀, R. Hurley leg., 2. viii.1958, 1 ♂, J. G. Chillcott leg.; ~ 6 km SW Shilo, 49°45.19'N 99°41.28'W, edge of wet area, sweeps, incl. Equi-[setum] ssuviatile , Carex , Calamagrostis , Impatiens , 10.vii.2011, 1 ♀, K. N.Barber leg.(all CNCI). NEW BRUNSWICK: Chamcook, 30.vi.1965, 1 ♂; Chamcook, Glebe Road, 14.vii1965, 2 ♀♀, all G. E. Shewell leg.; Kouchibouguac N. P., Code 5433Y, 27.vi.1977, 1 ♂ 3 ♀♀, J. R. Vockeroth leg. (all CNCI). NEWFOUNDLAND: Aspen Br[oo]k Cp. [P. Pk.], 18.vii.1961, 1 ♂, C. P. Alexander leg. ( USNM); Bay d’Espoir, low veg. on bog, 11.vii.1985, 1 ♀, L. Hollett leg.; Black Head, 21.vii.1967, 1 ♀, J. F. McAlpine leg.; Coal Brook, 24.vi.1983, 3 ♂♂ 1 ♀, A. Borkent leg.; Little H[arbou]r East, moist veg. nr. stream, 10.vii.1984, 1 ♂ 1 ♀, L. Hollett leg. (all CNCI); Junction Pond, Notre Dame Cp. [P. Pk.], 11.vii.1961, 2 ♂♂, C. P.Alexander leg.( USNM);Avalon Pen[insula], Portugal Cove, Indian Meal Line, 15.vii.1982, 1 ♀, 31.vii.1982, 1 ♂, b. l., 15.vii.1982, 2 ♀♀ (1 ♀ genit. prep.), [no collector] ( NFRC); Portugal Cove, Northeast Pond, 11.vii.1987, 1 ♂; Portugal Cove, sedge fen, 27.vi.1987, 1 ♂ (genit. prep.), both T. A. Wheeler leg. (both DEBU); St. John’s, Agric. Exp. Stn., 26.vii.1967, 1 ♀, J. F. McAlpine leg. ( CNCI, genit. prep.). NOVA SCOTIA: CBHNt. [Cape Breton Highlands National] Pk., Cheticamp R., PG563688, forest noodplain, 1.vi.1984, 1 ♀, B. E. Cooper leg.; same locality but Lone Shieling, PG731861, swept along fast rocky stream, 28.vi.1983, 1 ♀, J. R. Vockeroth leg.; same locality but Pleasant Bay, PG684871, wet hardwood forest, 10.vii.1984, 2 ♀♀, H. J. Teskey leg.; Cape Breton, 2 mi W Lone Shieling, Anse R., 29.v.1983, 2 ♀♀, H. Goulet leg.; Kentville, 6.viii.1958, 1 ♂, J. R. Vockeroth leg.; Lawrencetown, Halifax Co., 19–20.vii.1967, 1 ♂, D. M. Wood leg. (all CNCI); Lockeport, 18.vii.1958, 16 ♂♂ 9 ♀♀ (1 ♀ genit. prep.), 25.vii.1958, 3 ♂♂; Cranberry I., Lockeport, 24.vii.1958, 1 ♂, 25. vii.1958, 1 ♀; Mount Uniacke, 5.viii.1958, 1 ♂; Petit Étang, PG565677, marshy river shore, fresh water, 25.vi.1983, 1 ♀; S. Harbour, PG929935, marshy lake shore, brackish water, Carex , Iris , Juncus , 27.vi.1983, 1 ♂ 1 ♀; S. Harbour, beach, PG949958, damp ground with Carex & Iris , 3.vii.1983, 1 ♂, all J. R. Vockeroth leg. (all CNCI). ONTARIO: Algonquin Park, 26–30.vi.1955, 3 ♂♂ 5 ♀♀ (1 ♀ genit. prep.), Sabrosky leg. ( USNM);Algonquin Pk., 23.vi.1962, 1 ♂ 1 ♀, R. J. Pilfrey leg., 20.vi.1978, 2 ♀♀, 4.vii.1978, 2 ♀♀, D. McCorquodale leg.; Algonquin Pk., Swan Lake Stn., Scott Lake Survey, 45°29'15"N 78°43'20"W, forested slope, 17.vi.1993, 1 ♀, S. A. Marshall leg.; Algonquin P. Pk., nr. Crossbar Lake, 45.3271°N 78.2990°W, Malaise trap, hardwood forest, canopy gap ( CR-GAP), 4–17. vii.2008, 1 ♂, E. Proctor leg. (all DEBU); Algonquine [sic], mixed wood, 1.vi.1991, 1 ♀, M. Barták leg. ( MBPC, genit. prep.); 10 km NW Penetanguishene, Awenda P. Pk., Second Lake, sweep sedge at shoreline, 13.vii.1992, 1 ♂ 3 ♀♀, T. A. Wheeler leg. ( LEMQ 0039598–600, 603); 10 km N Bobcaygeon, “Nogie’s Creek”, sweep creekside veg., 1.vii.1995, 1 ♂, J. M. Dow leg. ( DEBU); Brittania, 17.vi.1938, 1 ♀, G. E. Shewell leg. ( CNCI); Bruce Co., Cramer Rd., 29.vii.1997, 1 ♂, S. A. Marshall leg.; Bruce Co., Crane River & Hwy#6, sweep, 23.vi.1995, 1 ♀, both S.A. Marshall leg.; Bruce Co., Hwy#6 @ Crane River, 45°08.91'N 81°28.30'W, sweeps, Carex aquatilis , 21.vi.2008, 1 ♂, K. N. Barber leg. (all DEBU); Bruce Co., Dorcas Bay, 45°11'N 81°35'W, 22.vi.2008, 2 ♂♂ 2 ♀♀, S. A. Marshall leg. ( DEBU 00299106, -12, -15, -16, 1 ♂ 1 ♀ genit. prep.); Bruce Co., Hwy#6 @ Willow River, 45°10.39'N 81°31.23'W, sweeps, mostly Carex , Equisetum , 21.vi.2008, 3 ♂♂ 5 ♀♀, J. Klymko leg. ( DEBU); Bruce Co., Bruce Pen. Natl.Pk., Lower Andrew Lk., portage trail, sweep, 8.vi.2000, 1 ♂, C. S. Onodera leg. ( DEBU 00080050); Bruce Peninsula N. P., Bartley Lake, 45°12.9'N 81°29.3'W, sweeps, shoreline graminoids, 2.vii.1999, 2 ♂♂ 1 ♀, K. N. Barber leg.; Bruce Peninsula N. P., bank of Crane River nr. Hwy 6, 45°08.9'N 81°28.1'W, sweeps/pooter, Calamagrostis & Carex , 31.vii.1997, 3 ♂♂ 3 ♀♀ (1 ♀ genit. prep.), K.N. Barber leg., sweeps, grasses / sedges, 31.vii.1997, 1 ♂, S. A. Marshall leg.; Bruce Peninsula N. P., Crane River below Lake Scugog, 45°07.0'N 81°32.1'W, sweeps, riverside vegetation, 3.vii.1999, 2 ♂♂ 1 ♀ (1 ♂ genit. prep.), K. N. Barber leg. (all DEBU); Bruce Peninsula N. P., Dorcas Bay Rd. at Willow Creek, 45°09.4'N 81°34.4'W, sweeps, mostly creekside graminoids, 20.vi.1999, 1 ♂, S. A. Marshall leg., 3.vii.1999, 5 ♂♂ 6 ♀♀, K. N. Barber leg.; same locality but 45°09.39'N 81°34.43'W, sweeps, Carex aquatilis , 22.vi.1999, 2 ♀♀, K. N. Barber leg. (all DEBU); Bruce Peninsula N. P., Emmett Lake, 45°13.1'N 81°29.0'W, sweeps, graminoids, wet area under Acer , 2.vii.1999, 1 ♀; same locality but 45°13.4'N 81°28.3'W, sweeps, graminoids, edge of marsh, 2.vii.1999, 1 ♂ 1 ♀; Bruce Peninsula N. P., Singing Sands, 45°11.6'N 81°34.4'W, sweeps, grasses along dry ditch, 31.vii.1997, 1 ♂ 4 ♀♀, 4.vii.1998, 1 ♂ 4 ♀♀; same locality but 45°11.50'N 81°34.61'W, sweeps, sedges in fen, 20.vi.2008, 1 ♂, 22.vi.2008, 8 ♂♂ 1 ♀ (1 ♂ genit. prep.), all K. N. Barber leg. (all DEBU); Bruce Co., Singing Sands, 45°11.5'N 81°34.9'W, 27.vii.1997, 1 ♀, S.A.Marshall leg.( DEBU 00075282); ~ 40 km NE Chapleau, 47°59.76'N 82°55.04'W, wet roadside sweeps, mostly Carex utriculata , 23.vi.2013, 1 ♂ (genit. prep.), 13.vii.2013, 4 ♂♂ 2 ♀♀, wet roadside sweeps, mostly Carex utriculata , grasses, 23.vi.2013, 3 ♂♂ 1 ♀; ~ 5 km SE Cochrane, 49°01.16'N 80°57.93'W, sweeps, railside Equisetum spp., graminoids, herbs, 18.vii.2009, 1 ♂; ~13.5 km S Cochrane, 48°56.65'N 81°00.18'W, hydro right-of-way,sweeps, mostly Carex utriculata , 22.vi.2013, 1 ♂, 18.viii.2013, 1 ♂ 1 ♀, all K. N. Barber leg. (all CNCI); Crieff Bog, 3 km W Puslinch, C3bC, pt3b, sedge island, 28.v.–4.vi.1987, 1 ♂, D. Blades leg. ( DEBU); Dubreuilville, along Magpie River, 48°21.12'N 84°34.04'W, sweeps, Equisetum ssuviatile , Carex , 10.vii.2010, 5 ♀♀, K. N. Barber leg. ( CNCI); same locality but sweeping Equisetum ssuviatile , Carex spp. on muddy river bank, 10.vii.2010, 8 ♂♂ 14 ♀♀, J. Roháček leg. ( SMOC, 5 ♂♂ 3 ♀♀ genit. prep., 1 ♂ 1 ♀ photographed); ~ 35 km WSW Dubreuilville, 2 km SE jct. Hwys.#17 & #519, 48°17.16'N 84°53.34'W, sweeps, roadside vegetation incl. wet ditch, 31.vii.2008, 1 ♂ 1 ♀, sweeps, Equisetum sp., 31.vii.2008, 2 ♀♀; ~ 3.8 km ENE Dugwal, 48°35.33'N 80°57.90'W, sweeps, wet ditch, Carex utriculata , Equisetum ssuviatile , Scirpus , 23.vi.2013, 1 ♂, 13.vii.2013, 2 ♀♀, all K. N. Barber leg. (all CNCI); ~ 55 km NNW Elliot Lake, S of Rocky Island Lake, 455 m, 46°49.32'N 82°59.54'W,sweeping, vegetation with predominant Scirpus sp., 3.vii.2010, 1 ♀, J. Roháček leg. ( SMOC); ~ 59 km NNW Elliot Lake, s. of Rocky Is. Lake, 46°50.16'N 83°03.05'W, sweeps, mostly Carex aquatilis in fen, 3.vii.2010, 2 ♂♂ 1 ♀, K. N. Barber leg. ( CNCI); same locality but 455 m, sweeping, mostly Carex rostrata ? [more likely C. utriculata ] in fen, 3.vii.2010, 1 ♂ 1 ♀, J. Roháček leg. ( SMOC); ~ 63 km NNW Elliot Lake, s. of Rocky Is. Lake, 46°49.80'N 83°09.08'W, sweeps, streamside graminoids, 4.vii.2010, 3 ♂♂ 1 ♀, K. N. Barber leg. ( CNCI); same locality but 425 m, sweeping streamside graminoids, 4.vii.2010, 3 ♀♀, J. Roháček leg. ( SMOC); ~ 66 km NNW Elliot Lake, Rocky Island Lake, 46°50.82'N 83°08.76'W, sweeps, Scirpus [ microcarpus ] on dried shoreline, 4.vii.2010, 2 ♂♂ 4 ♀♀, K. N. Barber leg. ( CNCI); same locality but 405 m, sweeping Scirpus sp. [ S. microcarpus ] on dried shoreline, 4.vii.2010, 6 ♂♂ 5 ♀♀, J. Roháček leg. ( SMOC, 1 ♂ genit. prep.); Fathom Five Nat. Pk., N. Cove Island, dry cedar, p.m. sweeps, 25.vi.1995, 1 ♂, fen, Bass Bay shore, p.m. sweeps, 25.vi.1995, 4 ♂♂ 5 ♀♀; Fathom Five Nat. Pk., [N.] Cove Island, shore of Bass Bay, sweep, 26.vii.1995, 1 ♂, all S. A. Marshall leg. (all DEBU); Finland, S of Caliper Lake, sweep sedge at beaver dam, 10.vii.1992, 2 ♀♀, T. A. Wheeler leg. ( LEMQ 0039601, -602, 2 ♀♀ genit. prep.); ~7.0 km E Foleyet, 48°14.34'N 82°20.75'W, hydro right-of-way, sweeps, mostly Carex utriculata , 13.vii.2013, 1 ♀; ~ 2.8 km SE Fraserdale, 49°49.96'N 81°35.03'W, sweeps, mostly Equisetum ssuviatilis in wet ditch, 20.vii.2009, 1 ♂; Goulais River, end of Island Rd., 46°43.57'N 84°24.45'W, sweeps, mud nats, Equisetum ssuviatile , Dulichium arundinaceum , 9.vii.2007, 1 ♂ 1 ♀; Goulais River, Pine Shores Rd., 46°41.67'N 84°25.47'W, sweeps, Thalictrum , Clematis , sedges, edge of Cranberry Ck., 4.vii.2007, 1 ♂ 1 ♀, all K. N. Barber leg.(all CNCI); Goulais River, Sand Bay, 46°44.81'N 84°32.68'W, sweeping Juncus and Carex at margins of fen pools, 10.vii.2010, 9 ♂♂ 3 ♀♀, J. Roháček leg. ( SMOC, 4 ♂♂ 1 ♀ genit. prep.); ~ 7.5 km NE Goulais River, Robertson Rd. below Robertson Cliffs, 46°45.89'N 84°18.40'W, sweeps, Carex gynandra under Acer , 4.vii.2016, 8 ♂♂ 16 ♀♀, K. N. Barber leg. ( CNCI); Greenwater P. Pk., 49°11.05'N 81°16.04'W, sweeps, mostly emergent Equisetum ssuviatile , 18.vii.2009, 2 ♂♂ ( DEBU 01501989, -90); Greenwater P. Pk., 49°10.91'N 81°16.28'W, sweeps, Carex , Calamagrostis , Phalaris , in creek noodplain, 21.vii.2009, 2 ♂♂ 9 ♀♀ ( DEBU 01502129–39); Greenwater P.Pk., 49°10.93'N 81°16.37'W,sweeps, Phalaris arundinacea in creek noodplain, 21.vii.2009, 1 ♀ ( DEBU 01502107); Greenwater P.Pk., Sandbar Lk.Trail, 49°13.10'N 81°17.35'W,sweeps, lakeshore Equisetum spp., graminoids, Caltha , 21.vii.2009, 3 ♂♂ 6 ♀♀ ( DEBU 01502090–98, 1 ♀ genit. prep.), all K. N. Barber leg.; Guelph, 14.vi.1973, 1 ♀, D. H. Pengelly leg., 31.v.1982, 1 ♀, A. W. Schaafsma leg. ( DEBU); ~ 85 km W Hearst, 49°46.14'N 84°49.10'W,gas right-of-way, sweeps, sedges and Equisetum ssuviatile , 21.vi.2013, 1 ♀; ~ 5 km NE Heyden, Hwy 552, 46°41.37'N 84°16.85'W, sweeps, roadside ditch/stream, graminoids, 31.vii.2006, 2 ♀♀; ~ 6 km N Heyden, Hwy #17 base of “Mile Hill”, 46°41.74'N 84°20.60'W, sweeps, graminoids under alder, 7.viii.2005, 1 ♂ 1 ♀; ~ 13 km N Heyden, ~ 3.8 km E Robertson Lake Rd., 46°45.7'N 84°18.4'W, sweeps, Carex , Impatiens , Aster , 1.viii.2005, 1 ♀, all K. N. Barber leg. (all CNCI); Hwy 17N & Trout Lake Road, 46°37.563'N 84°17.019'W, sweep, roadside, 23.vii.2011, 1 ♂ 1 ♀, J. E. Swann & D. R. Edwards leg. ( BDUC); Hurkett, dock area, 48°50.42'N 88°29.38'W, sweeps, emergent Equisetum ssuviatile , 31.vii.2008, 1 ♀; Icewater Creek WS [Watershed], 46°53.72'N 84°03.39'W, sweeps, riparian ferns, graminoids, 5.viii.2006, 1 ♂ 1 ♀; same locality but 12.7 km NNE Searchmont, mi.10.5 Whitman Dam Rd., alder thicket, 21.vi.1986, 6 ♂♂ 10 ♀♀ (1 ♀ genit. prep.), 23.vi.1986, 2 ♂♂ 3 ♀♀, 7.vii.1986, 1 ♀, 21.vii.1986, 1 ♂, all K. N. Barber leg.; same locality but ~ 12.7 km NNE Searchmont, mi.10.5 Whitman Dam Rd., riparian meadow/ alder thicket, Malaise, 24–29.vi.1986, 1 ♂, [K. N. Barber] leg.; same locality but 13.5 km NNE Searchmont, mi.15 Whitman Dam Rd., sandy access road, 20.vi.1986, 2 ♀♀, K. N. Barber leg.; Iroquois Falls, overgrown wet shrubby Sphagnum bog, 18.vi.1987, 2 ♂♂ 13 ♀♀, swept over small sandy stream, 24.vi.1987, 1 ♂, Carex , grass, on unshaded wet soil, 24.vi.1987, 1 ♂, J. R. Vockeroth leg. (all CNCI); ~11.9 km N Kejick, 49°43.89'N 95°04.14'W, sweeps, wet ditch, graminoids/ Equisetum , 30.vii.2008, 2 ♂♂ 2 ♀♀, sweeps, wet ditch, Equisetum ssuviatile , 30.vii.2008, 2 ♂♂ 1 ♀, sweeps, wet ditch, Calamagrostis canadensis , 30.vii.2008, 1 ♀; King Mt., 26 km N S[ault] S[ainte] Marie, riparian sweeps, 16.vi.1987, 12 ♂♂ 18 ♀♀ (1 ♀ genit. prep.), all K. N. Barber leg. (all CNCI); Lake Superior P. Pk., Crescent Lk. campground, 47°16.66'N 84°33.00'W, sweeps, shoreline sedges, 29.v.2010, 1 ♂, K. N. Barber leg. ( DEBU 01502430); same locality but Crescent Lake Trail, 47°16.73'N 84°33.12'W, sweeping Clintonia , ferns, Aralia , Maianthemum under Betula / Acer , 9.vii.2010, 1 ♂, J. Roháček leg. ( SMOC); Lake Superior P. Pk., Hwy 17 near jct. Agawa Rock, 47°22.31'N 84°41.23'W, sweeps, mostly Carex utriculata , 30.vi.2013, 3 ♂♂ 2 ♀♀ ( DEBU 01503939–43, 2 ♂♂ genit. prep.), 12.vii.2014, 1 ♂ 3 ♀♀ ( DEBU 01503950–53), K. N. Barber leg.; Algoma District, Lake Superior Prov. Pk., Rabbit Blanket Lake, 43°41'24"N 80°23'12"W [coordinates do not match locality], 400 m, 20.vi.2001, 1 ♂, S. A. Marshall leg. ( DEBU 00170277); Luther Marsh Bog, sweeps, 27.vi.1985, 1 ♂ 4 ♀♀, K. N. Barber leg. ( DEBU); ~ 6.3 km E Macleod, 49°41.37'N 86°51.41'W, sweeps, graminoids incl. Carex utriculata & C. aquatilis substricta , 6.vii.2012, 1 ♂, K. N. Barber leg. ( CNCI); Manitoulin I., nr. Evansville, Campbell Bay, 45°49'32"N 82°33'14"W, sweeping, lake shore graminoid vegetation, 2.vii.2010, 2 ♀♀, J. Roháček leg. ( SMOC); Manitoulin Is., ~ 2.2 km N Cold Springs, Perch Ck. @ Hwy 540, 45°53.1'N 82°06.2'W, sweeps, various grasses / sedges in noodplain, 4.vii.1999, 2 ♂♂; same locality but 45°53.2'N 82°06.3'W, sweeps/pooter, Calamagrostis canadensis , 1.viii.1997, 1 ♀, K. N. Barber leg. ( CNCI); Manitoulin Island, 10 km W Gore Bay, 45°53'N 82°34'W, sweep in grassland alvar, 20.vi.1996, 5 ♂♂ 4 ♀♀, P. Bouchard leg. ( LEMQ 0039586–94, 1 ♀ genit. prep.); Manitoulin Is., 0.7 km N Michael’s Bay Pk., 45°36.0'N 82°06.3'W, sweeps, lakeside grasses, 4.vii.1999, 1 ♀; same locality but 45°36.5'N 82°06.2'W, sweeps/pooter, graminoids in fen nat, 28.vii.1997, 1 ♂ 4 ♀♀, 30. viii.2004, 1 ♂, sweeps, Carex / Calamagrostis in fen nat, 5.vii.1998, 1 ♂ 4 ♀♀, all K. N. Barber leg. (all CNCI); Manitoulin I., Misery Bay Nature Reserve 45°47.64'N 82°44.93'W,sweeping, mosty Carex from wetlands boardwalk, 2.vii.2010, 11 ♂♂ 12 ♀♀, J. Roháček leg. ( SMOC, 1 ♂ 3 ♀♀ genit. prep.), sweeps, mostly Carex stricta from wetlands boardwalk, 2.vii.2010, 2 ♂♂ 1 ♀, K. N. Barber leg. ( DEBU 01502488–90); Manitoulin I., nr. Misery Bay Nat. Reserve, Little Lake Huron Road, 45°47.93’N 82°45.52’W, sweeping, mostly Carex , Juncus on wet alvar, 2.vii.2010, 1 ♂ 11 ♀♀, J. Roháček leg. ( SMOC, 1 ♀ genit. prep.), sweeps, mostly Carex , Juncus , on wet alvar, 2. vii.2010, 3 ♂♂ 6 ♀♀, K. N. Barber leg. ( CNCI); Manitoulin Island, Poplar, 45°46'N 82°28'W, sweep along grassy trail, 27.vi.1992, 1 ♂ 1 ♀, T. A. Wheeler leg. ( LEMQ 0039604, -05, 1 ♂ genit. prep.); Manitoulin I., Sand Bay, 45°48'06"N 82°47'36"W, dunes, sweep grassy area by stream, 25.vi.2003, 1 ♀, M. Buck leg. ( DEBU 01131800); Marmora, 7.vi.1952, 1 ♀; Mattawa, wet alder thicket, rich undergrowth, 16.vi.1987, 2 ♀♀ (1 ♀ genit. prep.), all J. R. Vockeroth leg.; Hwy#17, ~ 7 km W Mattawa, 46°17.3'N 78°49.0'W, sweeps, Carex gynandra , 30.vi.2005, 5 ♂♂ 4 ♀♀, K. N. Barber leg.; Maynooth, 19.vi.1953, 2 ♀♀, J. F. McAlpine leg.; Mer Bleue, 23.vi.1952, 2 ♂♂, G. E. Shewell leg.; Midland, 30.vii.1950, 1 ♂ 2 ♀♀ (1 ♀ genit. prep., hind legs in microvial), swampy woods, balsam poplar, 2.v.1959, 1 ♀, J. G. Chillcott leg. (all CNCI); Moosonee, 51.27717°N 80.64778°W, Repl. 3 wet, pan traps, 16–19.vi.2010, 1 ♂; Moosonee, 51.24690°N 80.68102°W, Repl.3 mesic, Malaise trap, 23–26.vi.2010, 1 ♂; Moosonee, 51.28034°N 80.64252°W, Repl. 1 wet, Malaise trap, 15–18.vi.2010, 1 ♂; Moosonee, 51.27717°N 80.64778°W, Repl. 3 wet, Malaise trap, 19–22.vi.2010, 1 ♂, all NBP Field Party leg. (all LEMQ); Moosonee, 51°16.07'N 80°38.66'W, sweeps, shoreline graminoids, 9.vii.2014, 1 ♂; Moosonee, 51°16.63'N 80°38.87'W, sweeps, Calamagrostis , Carex , drier edge of sedge meadow, 9.vii.2014, 2 ♂♂ 2 ♀♀; Moosonee, 51°16.69'N 80°38.86'W, general sweeps, sedge meadow, 9.vii.2014, 11 ♂♂ 9 ♀♀ (1 ♂ genit. prep., 2 pairs in copula); Moosonee, 51°16.75'N 80°38.76'W, sweeps, Equisetum [ ssuviatile ] & graminoids, wet edge of sedge meadow, 9.vii.2014, 5 ♂♂ (1 ♂ genit. prep.); Moosonee, 51°16.68'N 80°38.65'W, sweeps, mostly C [arex] utriculata , C. aquatilis , wet sedge meadow, 10.vii.2014, 2 ♂♂ 2 ♀♀ (1 ♂ genit. prep.); Moosonee, 51°16.54'N 80°39.00'W, sweeps, Equisetum , Rubus , Cornus , graminoids, edge of wet forest trail, 9.vii.2014, 2 ♀♀, 10.vii.2014, 3 ♂♂, 11.vii.2014, 2 ♂♂ 2 ♀♀; Moosonee, 51°16.55'N 80°39.01'W, sweeps, mostly Carex spp., wet forest trail, 11.vii.2014, 1 ♂ 1 ♀ (1 ♀ genit.prep.); Moosonee, 51°16.99'N 80°38.37'W, sweeps, mostly Equisetum ssuviatile , Carex spp., wet sedge meadow, 10.vii.2014, 7 ♂♂ 9 ♀♀; Moosonee, 51°16.36'N 80°39.11'W, sweeps, railside ditch, mostly Equisetum ssuviatile , Carex spp., 10.vii.2014, 5 ♂♂ 4 ♀♀, 11.vii.2014, 1 ♂ 1 ♀; Moosonee, 51°16.17'N 80°39.10'W, sweeps, mostly Carex utriculata , Scirpus , in wet hydro cut, 10.vii.2014, 1 ♀, all K. N. Barber leg. (all CNCI); Obatanga P.Pk., 48°20.60'N 85°02.03'W, sweeps, Carex sp., Equisetum ssuviatile , 7.vii.2007, 1 ♂, K. N. Barber leg. ( DEBU 01501890); One-Sided [= Caliper] Lake, 26.vi.1960, 1 ♀, Kelton & Whitney leg.; Ottawa, 19.vii.1954, 1 ♂, W. R. M. Mason leg. (both CNCI); Otter Rapids, 50°10.80'N 81°38.59'W, sweeps, Carex spp., 19.vii.2009, 1 ♀, sweeps, Equisetum ssuviatile , 19.vii.2009, 2 ♂♂ 1 ♀, sweeps, roadside Equisetum spp., 19.vii.2009, 1 ♂ 1 ♀; Otter Rapids, 50°10.85'N 81°38.65'W, sweeps, grasses in hydro cut, 20.vii.2009, 3 ♂♂ 2 ♀♀ (1 ♀ genit. prep.); Otter Rapids, 50°10.96'N 81°37.88'W, sweeps, mostly Schedonorus arundinaceus , on roadside slope, 20.vii.2009, 1 ♂, all K. N. Barber leg. (all CNCI); 1 km S Ouimet Canyon, sweeping grass/ sedge along river, 1.vii.1992, 1 ♀, T. A. Wheeler leg. ( LEMQ 0039597); Pancake Bay P. Pk., 46°58.00'N 84°42.47'W, Black Creek noodplain, sweeps, Carex / Calamagrostis , 17.vii.2004, 2 ♂♂ 3 ♀♀, K. N. Barber leg. ( DEBU 01500246–50); Pancake Bay P. Pk., 46°58.11'N 84°42.72'W, sweeps from boardwalk, mostly emergent sedges / Equisetum , 17.vii.2004, 2 ♂♂ ( DEBU 01500314, -15), 24.vii.2004, 4 ♂♂ ( DEBU 01500552–55), 2.viii.2004, 5 ♂♂ 2 ♀♀ ( DEBU 01500841–47, 1 ♂ genit. prep.), 7.viii.2004, 1 ♂ 3 ♀♀ ( DEBU 01501070–73, 1 ♀ genit. prep.), 27.vi.2005, 2 ♂♂ 1 ♀ ( DEBU 01501680–82), 26.v.2007, 1 ♂ ( DEBU 01501796), 16.vi.2007, 2 ♂♂ ( DEBU 01501799, -800), 7.vii.2007, 1 ♂ 1 ♀ ( DEBU 01501897, -98), all K. N. Barber leg.; Pancake Bay Prov. Park, 46°58.11'N 84°42.72'W, sweeping from boardwalk, mostly emergent sedges / Equisetum , 9.vii.2010, 7 ♂♂ 8 ♀♀, J. Roháček leg. ( SMOC, 1 ♂ 1 ♀ genit. prep., 1 ♂ 1 ♀ photographed); Pancake Bay P. Pk., 46°58.10'N 84°42.71'W, sweeps, mostly Carex nr. wetland boardwalk, 24.vii.2004, 2 ♂♂ 3 ♀♀ ( DEBU 01500697–701), 2.viii.2004, 1 ♂ 1 ♀ ( DEBU 01500901, -02); Pancake Bay P. Pk., 46°58.12'N 84°42.75'W, sweeps, mostly graminoids/ Typha near wetland boardwalk, 2.viii.2004, 1 ♂ 1 ♀ ( DEBU 01500961, -62), all K. N. Barber leg.; Petawawa, 7.vi.1961, 2 ♂♂ 1 ♀, J. R. Vockeroth leg.; Hwy 101 at Prairie Bee River (west side bridge), 47°51.81'N 83°54.33'W, sweeps, mostly Carex utriculata , 14.vii.2013, 1 ♂ 1 ♀; Hwy #17 at Prairie River mouth, 48°48.32'N 86°46.64'W, sweeps, grasses, composites, Rubus , forest edge, 16.vi.2007, 1 ♀, all K. N. Barber leg.; Puslinch Tsp. [Township], S Lot. 31, Gore Conc[ession], 1 ♂, 26.vi.1981, H. J. Teskey leg. (all CNCI); ~ 10.8 km W Jct Hwys 556 & 129, km 64.3 Ranger Lk. Rd., 46°53.46'N 83°27.01'W, sweeps, Equisetum ssuviatile and Carex sp., 23.vi.2007, 3 ♂♂ 1 ♀ (♀ genit. prep.), sweeps, Equisetum ssuviatile and Dulichium arundinaceum , 23.vi.2007, 1 ♀, K. N. Barber leg.; 10 km S Richmond, ex. fen, 29.v.1987, 1 ♂, J. M. Cumming leg.; ~ 4.5 km E Rosseau, on Aspdin Rd., 45°15.88'N 79°34.88'W, sweeps, mostly Carex in sedge meadow, 7.vii.2005, 1 ♂ 2 ♀♀, K. N. Barber leg.; St. Ignace Il. [ Island], Sturgeon Bay, 2.v.1959, 1 ♀, J. G. Chillcott leg. (all CNCI); S[ault] S[ainte] Marie, S. of Algoma U[niversity] College, 46°29.9'N 84°17.2'W, sweeps, Carex aquatilis , 12.vii.1997, 4 ♀♀ (1 ♀ genit. prep.), 26.vi.1998, 1 ♂ 1 ♀, 9.vi.2001, 2 ♂♂ 3 ♀♀, 10.vi.2001, 5 ♂♂ 4 ♀♀ (1 ♀ genit. prep.), 28.vi.2002, 1 ♂ 2 ♀♀, 29.vi.2002, 3 ♂♂ 5 ♀♀ (1 ♀ genit. prep.), 1.vii.2002, 1 ♂ 1 ♀, sweeps, mostly Carex aquatilis , 5.vi.2001, 1 ♂ 4 ♀♀ (1 ♂ wing illustration), 12.vi.2001, 2 ♂♂ 5 ♀♀ (1 ♀ genit. prep.) ( CNCI), 13.vi.2001, 4 ♂♂ 9 ♀♀ ( CNCI 2 ♂♂ 7 ♀♀, 1 ♀ genit. prep., SMOC 2 ♂♂ 2 ♀♀), 14.vi.2001, 3 ♂♂ 10 ♀♀, 15–16.vi.2001, 5 ♂♂ 6 ♀♀ (incl. 2 pairs in copula), 21.vi.2001, 3 ♂♂ 4 ♀♀, 22.vi.2001, 2 ♂♂, 25.vi.2001, 4 ♂♂ 7 ♀♀, 15.vii.2001, 1 ♂ 3 ♀♀ (1 ♀ genit. prep.), 17.vii.2001, 4 ♂♂ 2 ♀♀, 18.vii.2001, 1 ♂ 4 ♀♀ (1 ♀ genit. prep.), 21.vii.2001, 1 ♂, 29.vii.2001, 1 ♂, sweeps, trailside Carex / Scirpus , 14.vii.1998, 1 ♀, sweeps, Calamagrostis canadensis & Carex aquatilis , 23.vii.1997, 1 ♂ 1 ♀, sweeps, Calamagrostis canadensis , 26.vi.1998, 2 ♂♂ 2 ♀♀, sweeps, mixed graminoids, 18.vi.1998, 1 ♀, sweeps/pooter, Calamagrostis canadensis , 12.vii.1997, 1 ♀, sweeps/pooter, grassy edge Populus / Betula , 11.vii.1997, 1 ♀ ( CNCI), K. N. Barber leg.; same locality but 46°29.88'N 84°17.19'W, sweeps, Carex aquatilis , 1.vii.2003, 1 ♂ 1 ♀, 18.vi.2005, 5 ♂♂ 2 ♀♀, 27.vii.2005, 1 ♀, sweeps, mostly Carex aquatilis , 18.vii.2004, 1 ♀, 2.vii.2007, 1 ♂, sweeps, mostly Calamagrostis canadensis , 12.vi.2008, 1 ♂; same locality but 46°29.82'N 84°17.17'W, sweeps, mostly Carex aquatilis near river, 21.vii.2005, 1 ♂ 3 ♀♀, all K. N. Barber leg. (all CNCI); S[ault] S[ainte] Marie, S. of Algoma University, 46°29.88'N 84°17.19'W, sweeps, Carex aquatilis , 29.vi.2008, 2 ♂♂, sweeps, mostly Carex aquatilis , 18.vi.2016, 1 ♀, sweeps, mostly Carex spp., Scirpus cyperinus , 5.vii.2008, 1 ♀, sweeps, mostly Carex aquatilis , Calamagrostis canadensis , 5.vii.2008, 1 ♀; same locality but 46°29.81'N 84°17.15'W, sweeps, Carex aquatilis , Calamagrostis canadensis by river, 5.vii.2008, 3 ♂♂ 3 ♀♀, all K. N. Barber leg. (all CNCI); S[ault] S[ainte] Marie, Baseline Rd., 46°31.40'N 84°24.40'W, sweeps, Thalictrum , Rubus , Equisetum , Carex , ferns, under aspen, 9.vi.2005, 1 ♂ 2 ♀♀, 10.vi.2005, 1 ♀, sweeps, Rubus , Equisetum , Thalictrum , Impatiens , ferns, under aspen, 17.vi.2005, 1 ♂ 1 ♀, sweeps, Carex gynandra , 8.vii.2006, 1 ♀; same locality but 46°31.40'N 84°24.45'W, sweeps, edge of forest, Solidago , Rubus , Equisetum , grasses, 25.vi.2005, 1 ♀; same locality but 46°31.61'N 84°24.68'W, w. of creek, sweeps, Carex edge of alder thicket, 22.vii.2005, 1 ♂ 1 ♀ (1 ♀ genit. prep.), all K. N. Barber leg. (all CNCI); S[ault] S[ainte] Marie, Bellevue Pk., 46°30.1'N 84°18.1'W,sweeps, mostly Calamagrostis , 7–8.vii.2000, 2 ♂♂; S[ault] S[ainte] Marie, Birchwood Pk., mixed forest, 28.vi.1986, 3 ♂♂ 4 ♀♀, 5.vii.1986, 1 ♀; same locality but 46°30.7'N 84°15.6'W, sweeps, including Impatiens , under Betula / Acer , 19.vi.1998, 1 ♀; S[ault] S[ainte] Marie, Bristol Pl[ace] Pk., 46°30.8'N 84°16.6'W, sweeps, Phalaris arundinacea , 28.v.1999, 1 ♀, all K. N. Barber leg. (all CNCI); S[ault] S[ainte] Marie, Finn Hill, 46°31.6'N 84°17.3'W, sweeps, graminoids/ composites, 19.vii.2005, 1 ♂ 1 ♀; same locality but 46°31.6'N 84°17.4'W, sweeps, mostly graminoids under Populus , 4.vii.2002, 1 ♂ 1 ♀, sweeps, graminoids in wet area under Populus , 1.vii.2002, 1 ♀; same locality but 46°31.7'N 84°17.5'W, sweeps, mostly sedges in trail, 4.vii.2002, 1 ♂ 5 ♀♀, sweeps, mostly sedges in trail, 7.vii.2002, 2 ♀♀; same locality but 46°31.7'N 84°17.3'W, sweeps, meadow graminoids, 1. vii.2002, 1 ♂, sweeps/pooter, Calamagrostis canadensis , 1.vii.2002, 1 ♀; same locality but 46°31.63'N 84°17.33'W, sweeps, Scirpus cyperinus , 19.vii.2004, 6 ♂♂ 4 ♀♀, sweeps, Carex stipata stipata , 19.vii.2004, 5 ♂♂ 3 ♀♀, 20. vii.2004, 2 ♂♂ 5 ♀♀, 6.vii.2008, 1 ♂ 2 ♀♀, 25.vi.2009, 9 ♂♂ 7 ♀♀, 18.vi.2016, 3 ♂♂ 2 ♀♀, 29.vi.2016, 2 ♂♂ 3 ♀♀, 2.vii.2016, 1 ♂ 1 ♀, pooter, Carex stipata stipata , 21.vii.2004, 1 ♂, 22.vii.2004, 1 ♂, 23.vii.2004, 1 ♀, 25. vii.2004, 1 ♂ 1 ♀, sweeps, mostly Carex stipata stipata , 26.vi.2007, 17 ♂♂ 21 ♀♀, 13.vii.2007, 1 ♀, all K. N. Barber leg. (all CNCI), sweeping boggy meadows, mostly Carex stipata stipata , 7.vii.2010, 1 ♂, 12.vii.2010, 2 ♂♂, J. Roháček leg. ( SMOC); sweeps, mostly Carex / Calamagrostis , edge of Populus tremuloides , 25.vi.2009, 17 ♂♂ 12 ♀♀ (incl. 2 pairs in copula, 2 ♀♀ genit. prep.), sweeps, graminoids, herbs, composites, edge of Populus tremuloides , 6.vii.2008, 2 ♀♀, 8.viii.2008, 1 ♀, 25.vi.2009, 1 ♀, 18.vi.2016, 1 ♂ 4 ♀♀, 2.vii.2016, 1 ♂ 2 ♀♀; same locality but 46°31.48'N 84°17.36'W, sweeps, mostly Scirpus microcarpus , 18.vi.2005, 1 ♀, sweeps, mostly Scirpus microcarpus , Impatiens , 27.vi.2007, 1 ♂, all K. N.Barber leg. (all CNCI), sweeping graminoid vegetation, 7.vii.2010, 1 ♀, J. Roháček leg. ( SMOC); same locality but 46°31.63'N 84°17.29'W, sweeps, Scirpus microcarpus , 19.vii.2004, 5 ♀♀, 4.viii.2004, 1 ♀, 29.vi.2016, 2 ♂♂; same locality but 46°31.63'N 84°17.43'W, sweeps, Impatiens , ferns, Carex gynandra , 15.vii.2006, 1 ♂ 1 ♀; same locality but 46°31.67'N 84°17.32'W, sweeps, Calamagrostis canadensis , 30.vii.2004, 1 ♂; same locality but 46°31.66'N 84°17.34'W, sweeps,mostly Calamagrostis canadensis , 12.vi.2008, 1 ♀ (genit. prep.), all K. N. Barber leg. (all CNCI); Sault Ste. Marie, Fish Hatchery Road, near Coldwater Creek, 46°34.29'N 84°17.21'W, sweeping graminoids, Impatiens , 9.vii.2010, 1 ♂ 1 ♀, J. Roháček leg. ( SMOC); S[ault] S[ainte] Marie, Landslide Rd., Coldwater Ck. noodplain, 46°33.8'N 84°16.6'W, sweeps/pooter, Calamagrostis canadensis , 7.viii.1997, 1 ♀, sweeps, mostly Carex sp., 7.viii.1997, 2 ♂♂ ( CNCI), sweeps, mostly Carex aquatilis , 13.vi.2001, 7 ♂♂ 16 ♀♀ ( CNCI 5 ♂♂ 14 ♀♀, 1 ♀ genit. prep., SMOC 2 ♂♂ 2 ♀♀), K. N. Barber leg.; same locality but Coldwater Ck., 46°33.94'N 84°16.66'W, sweeps, graminoids in wet ditch, 26.viii.2008, 2 ♀♀, K. N. Barber leg.( CNCI); S[ault] S[ainte] Marie,River Road, 46°30.4'N 84°15.3'W, sweeps, mostly Carex spp., 11.vi.2001, 3 ♂♂ 5 ♀♀; same locality but 2nd Line E, 46°32.3'N 84°16.6'W, sweeps, graminoids in open meadow, 26.vi.1999, 1 ♀; same locality but 2nd Line Ext., 46°32.5'N 84°17.0'W, sweeps, mostly Carex sp., 13.vi.2001, 1 ♂ 1 ♀, all K. N. Barber leg. (all CNCI); S[ault] S[ainte] Marie, Sault Coll[ege] Outdoor Lab, 46°32.1'N 84°18.2'W, sweeps, short trail grasses, Acer / Betula , 25.vii.1997, 1 ♂ 1 ♀, sweeps, Carex sp., 25.vii.1997, 3 ♀♀ (1 ♀ genit. prep.), sweeps, mixed meadow veg., 18.vi.1998, 4 ♀♀, sweeps, trailside graminoids under Acer / Betula , 19.vi.2002, 1 ♂, sweeps, trailside Carex under Acer / Betula , 30.vi.2002, 1 ♀; same locality but hydro cut nr. Sault Coll[ege] Outdoor Lab, 46°32.1'N 84°18.0'W, sweeps, mostly sedges, 10.vii.2002, 3 ♂♂ 3 ♀♀ (1 ♀ genit. prep.), all K. N. Barber leg. (all CNCI); S[ault] S[ainte] Marie, Hwy #17 city limits, 46°36.58'N 84°17.83'W, sweeps, mostly riparian sedges, 16. viii.2004, 2 ♂♂ 1 ♀, sweeps, Calamagrostis canadensis in wet area, 16.viii.2004, 1 ♂ 3 ♀♀, sweeps, mostly Carex / Calamagrostis in wet area, 16.viii.2004, 1 ♂, 23.viii.2004, 1 ♀, sweeps, mostly Carex / Calamagrostis in wet area, 19.vii.2008, 6 ♂♂ 2 ♀♀, 13.viii.2008, 3 ♂♂ 4 ♀♀, all K. N. Barber leg. (all CNCI); same locality but 46°36.62'N 84°17.85'W, sweeps, mixed graminoids in alder thicket, 4.vii.2016, 2 ♂♂ 5 ♀♀ ( CNCI), sweeps, Carex gynandra in alder thicket, 4.vii.2016, 31 ♂♂ 39 ♀♀ ( AMNH, BDUC, CMNH, CASC, CSCA, INHS, KNWR, LACM, LEMQ 3 ♂♂ 3 ♀♀ each; CNCI 4 ♂♂ 12 ♀♀), 7.vii.2016, 29 ♂♂ 31 ♀♀ ( MCZN, NMPC, OSAC, PMAE, RBCM, SEMC, UBCZ, USNM 3 ♂♂ 3 ♀♀ each; CNCI 5 ♂♂ 7 ♀♀), all K. N. Barber leg.; S[ault] S[ainte] Marie, Voyageur Trail, 46°35.48'N 84°15.23'W, sweeps, Calamagrostis canadensis , 9.vii.2006, 2 ♀♀, [K. N. Barber] leg.; same locality but Wishart Pk., 46°33.86'N 84°17.57'W, sweeps, graminoids/ferns under canopy, 23.vii.2005, 1 ♀; same locality but Whitensh Is./St. Mary’s Is., 46°30.68'N 84°21.20'W, sweeps, riparian graminoids, 8.viii.2004, 4 ♂♂ 4 ♀♀, all K. N. Barber leg. (all CNCI); ~ 10 km W S[ault] S[ainte] Marie, Airport Rd., 46°29.72'N 84°28.96'W, natural gas r[igh]t-of-way, sweeps, mostly ferns, graminoids, 5.viii.2009, 1 ♀, sweeps, graminoids, Equisetum , herbs, 5.viii.2009, 2 ♀♀, K. N. Barber leg. ( CNCI), sweeping graminoids, composites, Equisetum , Rubus , ferns, 12.vii.2010, 2 ♂♂ 1 ♀, J. Roháček leg. ( SMOC, 1 ♂ genit. prep.); ~ 10 km W S[ault] S[ainte] Marie, Sunnyside Beach Rd., 46°29.70'N 84°31.87'W, sweeps, emergent Equisetum ssuviatile in wet ditch, 22.vi.2007, 2 ♂♂ 1 ♀, K. N. Barber leg. ( CNCI); Searchmont, N Hwy #552, 46°50.3'N 84°04.4'W, sweeps, roadside sedges / grasses, 10.vii.1998, 1 ♂ 2 ♀♀; ~ 5 km SE Searchmont, km6.2 Ranger Lk. Rd., 46°45.52'N 83°59.51'W, sweeps, sedges at beaver dam outnow, 23.vi.2007, 1 ♂; 18 km NNE Searchmont, mi.15 Whitman Dam Rd., grassy access road, 19.vi.1986, 3 ♂♂ 7 ♀♀, 24.vi.1986, 1 ♂, all K. N. Barber leg.; Simcoe, 13.vi.1939, 1 ♂, G. E. Shewell leg. (all CNCI); Smooth Rock Falls, 49°16.04'N 81°36.08'W, sweeps, Carex utriculata , 22.vi.2013, 1 ♂; ~ 21 km NNE Smooth Rock Falls, 49°20.91'N 81°32.01'W, sweeps, Equisetum ssuviatile in wet ditch, 19.vii.2009, 2 ♂♂ 2 ♀♀, all K. N. Barber leg. (all CNCI); Thessalon, 16.vi.1963, 1 ♂, R. G. Brumpton leg. ( DEBU); ~61.5 km N Thessalon, Hwy#129, 47°05.00'N 83°09.40'W, sweeps, emergent Equisetum ssuviatile / sedges, 23.vi.2007, 1 ♀; ~ 74 km NNE Thessalon, 46°53.94'N 83°16.23'W, shore of Mississagi R., sweeps, graminoids, herbs, Equisetum spp. [ E. ssuviatile & E. palustre ], 5.vii.2010, 5 ♂♂ 4 ♀♀, 17.vii.2010, 17 ♂♂ 13 ♀♀, all K. N. Barber leg. (all CNCI), sweeping graminoids with Equisetum spp. on muddy shore, 5.vii.2010, 6 ♂♂ 8 ♀♀ (1 ♂ 1 ♀ genit. prep.); ~ 92 km NNE Thessalon, nr. Mountain Ash Lake, 47°02.98'N 83°10.88'W, sweeping Carex aquatilis on edge of wetland, 4.vii.2010, 6 ♂♂ 13 ♀♀ (1 ♂ genit.prep.), all J. Roháček leg. (all SMOC); ~ 34 km N Timmins, 48°45.88'N 81°21.71'W, sweeps, Carex spp., 18.vii.2009, 1 ♂; ~ 29 km SW Timmins, 48°19.12'N 81°44.79'W, sweeps, Carex spp./ Calamagrostis , 18.vii.2009, 1 ♀, both K. N. Barber leg. (both CNCI); Wolf Lake env. nr. Dorset, peat-bog, sweeping over peat-bog meadow, 5.viii.1994, 1 ♂ 1 ♀, J. Roháček leg. ( SMOC); Wylde Lk. Bog, 8 km E Arthur, W2bE, pt2b, sedge meadow hollow, 16–22.vi.1987, 1 ♀, D. Blades leg. ( DEBU). QUEBEC: Beech Grove, 7.vi.1955, 1 ♂ 1 ♀ (1 ♀ genit. prep.), beaver grass, 7.vi.1955, 1 ♀, J. F. McAlpine leg.; Beechgrove, 45°39'N 76°08'W, 29.vii.1962, 1 ♂ 5 ♀♀, 27.vi.1984, 2 ♂♂ 2 ♀♀ (1 ♀ genit. prep.), 24.vi.1988, 2 ♂♂ 2 ♀♀; Farnham, 5.vi.1963, 11 ♂♂ 8 ♀♀ (1 ♂ 1 ♀ genit.prep.), all J. R.Vockeroth leg.; Fairy L.Crk., 30.v.1965, 1 ♀, [no collector] (all CNCI); Gaspesie, near Percé, 48°37'N 64°11'W, sweep, 1.viii.2001, 4 ♂♂, V. Dion & S. Boucher leg. ( LEMQ 0039576–79); Gatineau Co., Masham Township, 27.vii.1995, 2 ♂♂ 1 ♀, E. Ikeda leg. ( LEMQ 0039580–82, 1 ♂ genit. prep.); Gatineau Park, 8.vi.1954, 1 ♂, W. R. M. Mason leg.; Gatineau Pk., Black Lk., 48°29'28"N 75°51'31"W, streams E and below, 24.vi.2009, 2 ♂♂ 1 ♀, B. J. Sinclair leg.; Graceneld, 20.vi.1937, 1 ♂, O. Peck leg.; Gatineau Pk., Harrington L., 8.vi.1954, 1 ♀ (genit. prep.), H. J. Huckel leg., 7.vi.1954, 1 ♀, W. R. Richards leg., 3.vii.1963, 1 ♀, J. R. Vockeroth leg.; Knowlton Ldg., 11.vii.1968, 1 ♀, J. R. Vockeroth leg.; Lac Megantic, 22–24.vi.1982, 1 ♀, H. J. Teskey leg.; Lac Phillipe, 45°37'N 76°W, 7.vii.1968, 1 ♂, J. R. Vockeroth leg. (all CNCI); Lac Roddic, 16 km S Maniwaki, 22.vi.1991, 1 ♂, M. Barták leg. ( MBPC, genit. prep.); Mt. Ste. Marie, Low, 1800', 22.vi.1965, 1 ♀, J. R. Vockeroth leg. ( CNCI, genit. prep.); Meach [sic Meech] Lake, Old Chelsea, 1 ♂ 3 ♀♀, 24.vi.1980, K. N. Barber leg. ( DEBU); Old Chelsea, Summit King Mt., 1150', 14.vi.1963, 1 ♀ (genit. prep.), 18.vi.1963, 1 ♀, 21.vi.1963, 1 ♂, J. R. Vockeroth leg. ( CNCI); St. Charles Bog, 46°45'48.0"N 70°59'34.4"W, sweeping, abandoned site, T2, 22.vi.2006, 1 ♂ 1 ♀, A. G. Taillifer leg. ( LEMQ 0040323, -24, 1 ♂ genit. prep.); near Schefferville, 54°49.507'N 66°51.393'W, sweep, grass/ sedge meadow, 8.vii.2008, 1 ♀, A. Rogic leg. ( LEMQ, genit. prep.). SASKATCHEWAN: Beaver Creek Cons. Area, ~ 13 km S Saskatoon, 51°58.6'N 106°43.0'W, sweeps, mostly grasses under Betula / Populus , 17.vii.1999, 1 ♂, K. N. Barber leg. ( CNCI, genit. prep.). UNITED STATES OF AMERICA: IDAHO: Boundary [Co.], Perkins Lake, 48°45.6'N 116°05.5'W, 760 m, 3.vi.2006, 1 ♂, W. N. Mathis leg. ( USNM, genit. prep., hind leg in microvial). INDIANA: Lafayette, 17.vi.1915, 1 ♀, J. M. Aldrich leg. ( USNM, genit. prep.). MASSACHUSETTS: Concord, marshy pond, 27.vii.1961, 1 ♂ 6 ♀♀ (1 ♀ genit. prep.), W. W. Wirth leg.; Middlesex Co., Belmont, Rock Meadow, 12.vi.1981, 1 ♀, N. E. Woodley leg. (all USNM). MICHIGAN: Cheboygan Co., 17.vii.1942, 1 ♀, C. W. Sabrosky leg. ( USNM); Cheboygan Co., Roberts Road nr. U. Michigan Biol. Stn., 45°32.56'N 84°39.78'W, sweeps, roadside ditch, mostly Equisetum spp., 25.vi.2010, 1 ♀, K. N. Barber leg. ( CNCI); Hunt Ck. Exp. Sta., nr. Lewiston, 20.vii.1942, 3 ♂♂ 2 ♀♀ (1 ♀ genit. prep.), C. W. Sabrosky leg.; Keweenaw Co., 27.vii.1953, 1 ♂ 2 ♀♀ (1 ♀ head glued to point, missing wing), G. Steyskal leg.; Leelanau Co., 21.vi.1937, 1 ♂, C. W. Sabrosky leg.(S. W. Frost det. as Tethina maritima Melander ); Midland Co., 10–11.vi.1951, 1 ♂, R.R. Dreisbach leg. (all USNM). MINNESOTA: Eaglesnest, 25.v.1959, 1 ♀, W. V. Balduf leg. ( INHS 40,085, Sabrosky det. as Anthomyza gracilis Fallén ). MONTANA: 15 mi S Big Fork, 26.vi.1967, 2 ♂♂ 1 ♀ (all genit. prep.), D. Allen leg.; 1.0 mi S Swan Lake, 30.vi.1967, 1 ♂, 2.viii.1966, 1 ♂ (genit. prep.); 20 mi S Swan Lake, 14.vii.1967, 1 ♂ 1 ♀, all B. A. Foote leg. (all USNM). NEW HAMPSHIRE: Pinkham Notch, 9.vii.1931, 2 ♀♀ (1 ♀ genit. prep.), A. L. Melander leg.; White Mtns., Stinson Lake, 23.vii.1961, 1 ♀, W.W.Wirth leg. (all USNM);White Mts.Nat. For., Fabyan, 6.viii.1980, 1 ♂, A. E. Stubbs leg. ( BMNH, BM. 1983-250, genit. prep.). NEW YORK: Bemus Pt., Chautauqua Lake, swampy woods, 31.v.1963, 1 ♀; Lewis Co., Brantingham L., lake margin, 22.vi.1963, 3 ♂♂ 1 ♀, all W. W. Wirth leg. (all USNM); Franklin Co., Paul Smiths, 20.vii.1962, 1 ♂ 1 ♀, J. R. Vockeroth leg. ( CNCI); St. Lawrence Co., Colton, Raquette R., 24.vii.1963, 1 ♂; Chautauqua Co., S. Dayton, marsh area, 1.vi.1963, 1 ♀; Lewis Co., Whetstone Gulf, 20–23.vi.1963, 1 ♂ 1 ♀, all W. W. Wirth leg. (all USNM); Whiteface Mt., 4600–4872', 19.vii.1962, 1 ♀, J. R. Vockeroth leg. ( CNCI); Wilmington Notch, Adirondacks, “ vii-3 ”, 1 ♀, J. M. Aldrich leg. ( USNM). NORTH CAROLINA: Highlands, 31.v.1957, 1 ♂, W. R. M. Mason leg., 3800', 1.vi.1957, 1 ♀, J. R. Vockeroth leg. ( CNCI, both genit. prep.). OHIO: 3.0 mi N. Kent, Herrick Fen, 16.vii.1986, 1 ♂, B. A. Foote leg. ( CNCI). VIRGINIA: Giles Co., M[oun]t[ai]n Lake Biol.Stn., 37°23'33"N 80°33'05"W, cranberry bog, 1469 m, 26.v.1999, 1 ♂ 2 ♀♀, S.A. Marshall leg. ( DEBU). WEST VIRGINIA: Pocahontas Co., Cranberry Glades, 16.vii.1955, 1 ♂, W. W. Wirth leg.; Pocahontas Co., Hills Creek, Rt. 39, 30.vi.1982, 3 ♂♂ 2 ♀♀, O. S. Flint & W. Mathis leg. (all USNM). WISCONSIN: Washburn Co., T39N R12W B32, 13.vi.1953, 1 ♀ (genit. prep.), 4.vii.1953, 2 ♂♂, T39N R12W B33, 29.vi.1953, 1 ♀, 8. vii.1953, 1 ♂, 19.vii.1953, 1 ♂, R. H. Jones leg. ( USNM).

Other material examined (not included in type series). CANADA: NEWFOUNDLAND: St. Anthony , 8.vii.1951, 1 spec., J. B. Wallis leg. ( CNCI, abdomen missing) . ONTARIO: Algonquin Pk., 4.vii.1978, 1 ♀, D. McCorquodale leg. ( DEBU, missing 3 legs) ; Algonquin Park , 26–30.vi.1955, 1 ♂, C. W. Sabrosky leg. ( USNM, headless) ; Manitoulin I., Misery Bay Nature Reserve 45°47.64'N 82°44.93'W, sweeping, mosty Carex from wetlands boardwalk, 2.vii.2010, 1 ♀, J. Roháček leg. ( SMOC, headless) ; Moosonee , 51.27717°N 80.64778°W, Repl. 3 wet, Malaise trap, 19–22.vi.2010, 1 ♂ (damaged genit. prep.) ; Moosonee , 51.28288°N 80.63926°W, Repl. 2 wet, Malaise trap, 16–19. vi.2010, 1 ♂ (crushed), both NBP Field Party leg. (both LEMQ) ; S[ault] S[ainte] Marie, Finn Hill , 46°31.63'N 84°17.33'W, sweeps, Scirpus cyperinus , 19.vii.2004, 1 ♀, K. N. Barber leg. ( CNCI, headless, genit. prep.) ; ~ 74 km NNE Thessalon, shore of Mississagi River , 46°53.94'N 83°16.23'W, sweeping graminoids with Equisetum spp. on muddy shore, 5.vii.2010, 3 ♂♂, J. Roháček leg.( SMOC, used for molecular work, 1 ♂ headless) . UNITED STATES OF AMERICA: MASSACHUSETTS: Concord , marshy pond, 27.vii.1961, 1 ♀, W. W. Wirth leg. ( USNM, headless) . WISCONSIN: Washburn Co., T39 N R12 W B 32, 13.vii. 1953, 1 spec., R. H. Jones leg. ( USNM, abdomen missing) .

Description. Male. Total body length 2.04–2.78 mm; general colour dark brown to brown, distinctly but not densely grey to brownish grey microtomentose (more sparsely on abdomen), subshining to partly dull ( Figs 2 , 456, 457 , 481 ). Head as long as high or slightly higher than long, anteriorly (in pronle) almost rectangular because face very little receding ( Fig. 481 ). Occiput dorsomedially slightly concave, uniformly dark brown and grey microtomentose, at most with small medial area above foramen paler brown. Frons ( Fig. 456 ) yellow only anterolaterally; rest of frons of subtriangular shape (more or less narrowed anteriorly) dark brown to brown, microtomentose and dull, in darkest specimens with this dark area expanded up to anterior frontal margin and covering most of surface of frons. Frontal triangle including ocellar triangle dark brown and both grey microtomentose; orbit and adjacent forefrons pale yellow to brightly yellow and silvery whitish microtomentose up to posterior ors, in darkest specimens only orbit yellow to dirty ochreous yellow slightly behind the anterior long ors (but silvery white microtomentum extended up to posterior ors), dark brown and greyish microtomentose more posteriorly. V-shaped area between frontal triangle and orbits dark brown (posteriorly) to pale brown (anteriorly), with nne darker longitudinal striations particularly in paler anterior area; extent of this dark V-shaped area variable, reaching up to anterior margin of frons in darkest specimens. Frontal triangle relatively small and narrow, with acutely pointed anterior corner, reaching to anterior two-nfths to one-third of frons. Ocellar triangle and/or adjacent parts of frontal triangle often more shining than rest of the latter. Frontal lunule distinct, yellow. Face narrow, pale yellow to deep yellow, often with darker ochreous midline, sparsely whitish microtomentose and dull; parafacialia and gena pale yellow and with distinct and rather broad bordering stripe being pale ochreous and narrower on parafacialia and dark brownish and wider on gena and with golden somewhat glittering microtomentum, rest of gena silvery white microtomentose; postgena dark yellow to ochreous but posteriorly darkened, brownish as adjacent occiput. Mouthparts largely yellow, but palpus paler, clypeus brownish and prementum often partly darkened, ochreous to pale brown. Cephalic chaetotaxy generally as in A. tschirnhausi : pvt moderate to relatively long and slightly (rarely) to strongly crossed; vti and oc longest of cephalic setae; vte and posterior (or both) ors slightly to distinctly shorter than oc; 3 ors but only 2 long, middle subequal or slightly shorter than posterior; 1 shorter anterior ors setula well developed (rarely up to half length of anterior ors) but no microsetula in front of the latter; 1–2 pairs of medial microsetulae in anterior third of frons; 1 weak setula behind vte; postocular setulae (7–8) relatively long, in single row; postgena with several setulae and 2 short (ventral distinctly longer) setae; 1 long vi (about as long as anterior ors); subvibrissa greatly reduced, only as long as anterior peristomal setula; only 3–5 small peristomal setulae. Palpus of moderate width, yellow, with 1 relatively short ventropreapical seta and 6–7 ventral setulae. Eye broadly subovoid, with longest diameter oblique and about 1.3–1.4 times as long as the shortest. Smallest genal height 0.17–0.20 times as long as shortest eye diameter. Antenna distinctly geniculate, normally entirely yellow (in darkest specimens with darker yellow scape and pedicel); 1st nagellomere with moderate (medium-long) white marginal pilosity. Arista 1.8–1.9 times as long as antenna, with both basal segments ochreous yellow and terminal seta brown, both shortly ciliate (cilia shorter than those on 1st nagellomere).

Thorax very slightly narrower than head, unicolourous brown to dark brown (always darker than abdomen), greyish (but not heavily) microtomentose, subshining (particularly on anterior part of mesonotum) to almost dull (on pleura). Propleuron sometimes with small paler (ochreous) spot above fore coxa. Thoracic chaetotaxy closely resembling that of A. tschirnhausi : 1 relatively long hu (almost as long as anterior npl) and 1 microseta on humeral callus; 1 weak prs (only as long as or shorter than sa); 2 npl, anterior distinctly longer than posterior; 1 sa (shorter than pa) and 1 pa; 2 postsutural dc, anterior dc usually longer and thicker than anterior npl, posterior dc very long, longest of thoracic setae; 5–6 dc microsetae in front of anterior dc but the hindmost (situated in front of anterior dc) variably enlarged, sometimes even resembling a 3rd (but thinner) dc macroseta (as in A. tschirnhausi ); ac microsetae relatively sparse, in 2–4 rather irregular rows on suture, in 2 rows more posteriorly but never reaching beyond level of posterior dc; ppl microseta relatively long though pale and very nne; 2 sc, laterobasal weak (about as long as prs), apical almost as long as posterior dc; 2 relatively strong stpl, anterior very slightly to distinctly shorter than posterior, plus a few (1–3) upcurved pale setulae below stpl and 3–5 longer setae on ventral corner of sternopleuron. Scutellum rounded triangular, distinctly convex dorsally. Legs bright to pale yellow, usually with only last tarsal segment of all tarsi (variably) darkened (often only distally), ranging from dark yellow to pale brown but in darkest specimens with tarsi gradually brownish-darkened towards apex (2 or 3 distal segments darkest). f 1 with ctenidial spine variable, ranging from as long as to distinctly longer than maximum width of t 1; t 2 with very short but distinct posterior ventroapical seta but the other of the pair (anterior) reduced to small setula (in contrast to that of A. tschirnhausi ); f 3 posteroventrally with a row of nne, unmodined setae; fore and hind basitarsus with slightly longer proximoventral setulae. Claws relatively long (as in A. tschirnhausi ). Wing ( Fig. 485 ) elongate and relatively narrow; both veins and membrane pale brown to ochreous. C with distinct sparse spinulae between apices of R 1 and R 2+3. R 2+3 very long, bent parallel to C and apically slightly to distinctly upcurved to it; R 4+5 very slightly sinuous, bent in the middle and apically somewhat upcurved to almost straight, distally slightly divergent from M; the latter almost straight or very slightly bent posteriorly; dm cell medium long, with r-m situated distinctly in front of its middle; apical portion of CuA 1 slightly to distinctly longer than dmcu, almost reaching wing margin and A 1 ending far from it. Alula small but not very narrow. Wing measurements: length 2.18–2.80 mm, width 0.68–0.91 mm, Cs 3: Cs 4 = 0.98–1.31, rm\ dm-cu: dm-cu = 2.58–3.47. Haltere whitish yellow to dirty yellow, in darkest specimens with bases of knob and stem more or less ochreous grey darkened.

Abdomen with preabdominal terga brown but usually paler, less microtomentose and more shining than thorax ( Fig. 481 ). T1 short (less than two-thirds of T2), narrower and distinctly separate from T2 (only laterally partly fused), the latter almost as wide as T3; T3 and T4 subequal in size; T5 somewhat longer than T4 and posteriorly slightly tapered; T3–T5 often darker than T1 and T2. All preabdominal terga relatively shortly and sparsely setose. Preabdominal sterna light ochreous to pale brown. S1 pale ochreous with narrowly darkened posterior margin, transversely trapezoidal (wider posteriorly) to suboblong, distinctly shorter and slightly wider than S2. S2–S5 relatively large, slightly trapezoidal with anterior corners rounded, about as long as wide or slightly wider than long and becoming larger posteriorly, all shortly and more densely setose than associated terga. S2 usually darker (only in darkest specimens almost concolourous with) than S3–S5, with pale brown to brown pigmentation being darkest in anterior third and the widest and most transverse S5 sometimes (in dark specimens) laterally more or less brown-darkened. T6 relatively large (as characteristic for the A. tschirnhausi group; thus distinctly larger and less transverse than in congeners of other groups), about half length of T5, bare, usually paler brown than T5 and dorsomedially partly or completely narrowly desclerotized and unpigmented. Pregenital sterna S6–S8 relatively long, pale brown (S6) to brown (S7, S8 darkest); S6 and S7 with narrow dark brown anterior marginal ledge; S6 with 1 (rarely)–5, S7 with 1 (rarely)–4 short setae. S8 longer than S7 and about as long as epandrium, tapered posteriorly and setose in posterior two-thirds.

Genitalia. Epandrium ( Figs 460, 462 , 467, 469 ) blackish brown, relatively large, high and distinctly tapered dorsally but usually shorter than in both closest relatives, densely but shortly setose and with 2 or 3 pairs of longer and thicker dorsolateral setae; anal nssure small, narrow, rounded triangular (narrower than that of A. tschirnhausi ). Cercus small, with nne setae, apical and subapical longest. Medandrium very high, dorsally tapered and with narrow end dorsally emarginate as in relatives, with darkened medial ledge of variable length ( Figs 460 , 467 ). Gonostylus ( Figs 460, 466 , 467, 473 , 508, 509 ) largely yellow to distinctly darkened apically, somewhat shorter than epandrial height, slender, elongate, gradually tapered but apically variable, ranging from elongately pointed ( Fig. 466 ), shortly pointed ( Fig. 508 ) to distinctly blunt (cut) with small anterior tooth ( Figs 473 , 509 ) in darkest specimens; always with anterior side convex, in contrast to both A. tschirnhausi and A. gilviventris ; micropubescence restricted to posterior half of outer side of gonostylus and longer setae situated on its inner side. Hypandrium ( Figs 463 , 470 ) of moderate size, with anterior internal lobes distinct, dark and sclerotized, larger than in A. tschirnhausi and sometimes projecting above dorsal hypandrial margin ( Fig. 470 ). Transandrium narrow ( Figs 464 , 471 ), medially with a pair of small, low, dark dorsal lobes (but not tubercle-like projecting as those of A. tschirnhausi ); caudal process distinctly developed but rather variable ( Figs 464 , 471 ), narrow, nat, ventrally slightly narrowed and cut or rounded on tip and medially somewhat less sclerotized, in lateral view more ( Fig. 470 ) or less ( Fig. 463 ) bulging. Pregonite ( Figs 463 , 470 ) fused with hypandrium, low, posteroventrally with a small, inconspicuous lobe carrying 2 (1 longer) setae; anterior nat part of pregonite with 3 setae (all on inner side, posterior seta longest). Postgonite ( Fig. 463 ) very slender and elongate, dark proximally and pale distally, slightly bent but apically not pointed (sometimes almost lanceolate – Fig. 470 ), with 1 longer seta at middle of anterior margin. Basal membrane ventrally slightly bilobed, covered with nat spines being pigmented laterally and hyaline medially ( Figs 464 , 471 ), sometimes also with a group of setiform spines in lateral membrane ( Fig. 470 ); dorsolateral parts of basal membrane adjacent to caudal process of transandrium more or less distinctly secondarily sclerotized, pigmented and dorsally connected with transandrium ( Figs 463, 464 , 470, 471 ) as characteristic for the A. tschirnhausi group. Aedeagal part of folding apparatus with numerous, dense but nat spine-like excrescences ( Figs 465 , 472 ); connecting sclerite slender and rather weakly sclerotized, bordering the former. Phallapodeme slender with robust fulcrum, ventrolaterally projecting apex and shortly forked base. Aedeagus slender ( Figs 465 , 472 ), with small (but rather elongate) phallophore indistinctly separate from large distiphallus. Saccus relatively slender, shorter than nlum, sclerotized in proximal half where it is terminated by a lateral rounded sclerite, distally membranous and armed with 5 thick spines on apex; an additional spine is hidden also behind the rounded basal sclerite (i.e. most similar to that of A. gilviventris ). Filum formed by single, long, twisted and largely dark sclerite terminating in curved, acutely pointed ( Fig. 461 ) to nnely bicuspid ( Fig. 468 ) apex provided with 3–6 small subterminal spines; most of nlum (including its membranous proximal part) with a double to triple row of spinulae (less numerous than in A. gilviventris ). Ejacapodeme very small, slender, with a small, relatively pointed projection ( Figs 465 , 472 ).

Female ( Figs 2 , 456, 458 ). Similar to male unless mentioned otherwise. Total body length 2.38–3.41 mm. Face often with stripe bordering parafacialia darker.Antenna with 1st nagellomere more or less orange-ochreous to orange-brown darkened (at least anterodorsally), usually more extensively on inner side with only base remaining narrowly yellow. Ctenidial spine on f 1 variable in length, as long as to markedly longer than maximum width of t 1, but usually more robust than in male. Wing measurements: length 2.62–3.39 mm, width 0.85–1.05 mm, Cs 3: Cs 4 = 1.03–1.31, rm\dm-cu: dm-cu = 2.48–3.09. Abdomen with preabdominal terga (T1–T5) slightly to distinctly paler than in male, with variable pattern. Typical specimens have terga uniformly brown to partly or largely pale ochreous, each brownish posteriorly and laterally (though sometimes only narrowly) and usually also with a darker medial spot (cf. Figs 2 , 458 ). In very rare extremely pale specimens T1–T5 can be almost completely orange ochreous with these brownish darkenings reduced. All preabdominal terga more transverse than in male; T1 distinctly separate and narrower than T2; T2–T5 subequal in length but T2 and T5 usually slightly narrower than T3–T4; all rather shortly and sparsely setose, with longest setae on posterior margin of T5 (and T6). Preabdominal sterna pale ochreous to yellow, only S2 brownish-darkened in anterior half or third and S1 (entirely bare) with very narrowly darkened posterior margin; S1 relatively long but distinctly shorter than wide, trapezoidal (wider posteriorly); S2–S5 more transverse than in male, becoming wider posteriorly, S2–S4 about as long as wide or slightly wider than long, S5 largest and distinctly transverse (up to more than 1.5 times wider than long).

Postabdomen ( Figs 474, 475 ) very long, slender and elongate as characteristic for the A. tschirnhausi group, tapered posteriorly, telescopic, resembling in form that of A. gilviventris thus generally more slender than that of A. tschirnhausi . T6 broad, tapered posteriorly, transversely trapezoidal, completely brownish or brown spotted ( Fig. 474 ) to uniformly ochreous in lightest specimens, setose in posterior two-thirds (longest setae posteriorly). S6 narrower and less transverse than in A. tschirnhausi , pale ochreous yellow ( Fig. 475 ) to brownish in darkest specimens, sometimes with posterolateral margins darkened, with setae usually sparser than in both relatives. Tergosternum T7+S7 yet longer and narrower than in A. tschirnhausi , conical, emarginate anterodorsally, typically with posterior third to three-fourths dark brown ( Fig. 474 ), very rarely almost entirely yellow with posterior dark ring reduced to about apical one fourth, usually with pale brown to ochreous orange (broader) anterior part and posteriorly with variable unpigmented medial narrow area reaching up to half or more of original T7; ventral part of T7+S7 distinctly prolonged to form anterolateral lobes ( Figs 475 , 516) and with variable (usually large – Fig. 475 , reduced in dark specimens) paler area in anterior half being covered by distinctive whitish micropubescence – this area is unpigmented and depressed below 7th spiracle and somewhat projecting anteriorly, where it is usually separated from darker dorsal part by a more or less elevated ridge (Fig. 516). T7+S7 otherwise without micropubescence, shining, and with setae largely in posterior two-thirds, longest at posterior margin. T8 ( Figs 474, 477 ) yet more elongate than that of A. tschirnhausi , very narrow and usually slightly widened posteriorly and dark brown; without micropubescence and with most of setae reduced to microsetulae or setiform sensillae, thus with only a single pair of exclinate setae. S8 ( Figs 477, 478 ) also longer and narrower than in A. tschirnhausi , almost half length of T8, convex, longitudinally divided into two dark brown and largely bare (lacking micropubescence) sclerites except for sparse microsetulae or setiform sensillae and only 1–2 short and weak setae, all in posterior half; their posterior ends somewhat bent inside the 8th segment. Genital chamber ( Figs 476–478 ) with one pair of posterior elongate nattened sclerites (also longer than those of A. tschirnhausi ); annular sclerite somewhat bent and asymmetrical, situated ventrally, below anterior part of nat sclerites. Ventral receptacle ( Fig. 476 ) membranous, tubular, with longer and broader proximal sinuous part being distally attenuated and ending in slender twisted vermicular distal part. Accessory gland vesicular, subglobular, nnely granulose, on a subterminally dilated and partly ringed duct. Spermathecae (1+1) elongately subovoid ( Figs 479–480 ), one usually somewhat larger than other, the larger as long as or longer (0.9–1.2 times) than S10 medially, on very long ducts, well sclerotized and blackish brown, with sparsely transversely striated surface except basal part bearing a number of short spinulae (usually shorter and thicker than in relatives). T10 ( Figs 474, 477 ) small, elongately suboblong to pentagonal (with poorly denned posterior margin), brown, with a pair of long medial setae (rarely with 3 setae, Fig. 474 ) and 1–2 pairs of short setulae, otherwise without micropubescence. S10 ( Figs 475, 477 ) larger than T10, also brown, distinctly longer than wide, posteriorly rounded, bare except for a pair of moderate to short (sometimes very short, Fig. 477 ) setae and a few microsetulae and some micropubescence in posterior third. Cercus ( Figs 474, 477 ) small, moderate, slender, somewhat laterally nattened, with several short setae (including apical, distinctly exclinate setae) and with very reduced micropubescence restricted to ventral side ( Fig. 477 ).

Variability. This widespread species displays considerable variability in the brown colouration of the frons, terminal tarsal segments, preabdominal sclerites and other structures (cf. the description above) but also in the colouration and sclerotization of some male and female genitalic structures. Interestingly, the darkest male specimens from populations at the southwestern limit of the known distribution of the species ( United States of America: Idaho, Montana) have the apex of the gonostylus roundly to natly blunt. This variant ( Figs 473 , 509 ) is considered to represent the extreme limit of the variability of this structure because intermediate forms (with shortly pointed apex of gonostylus, Fig. 508 ) were found in the United States of America (Colorado) and Canada (Ontario). Typical specimens that have the gonostylus elongately acute ( Fig. 466 ) occur across the entire species range. The darkest males with the blunt gonostylus also have a more sclerotized basal membrane with spines and darker armature of the aedeagal part of the folding apparatus. The females with a darker frons and terminal tarsal segments (syntopic with males discussed above) do not seem to have particularly darkened postabdominal sclerites (including T7+S7 having only the posterior dark brown ring longer). As given in the description above, female T1–T6 and T7+S7 rarely have the brown colour reduced to become largely to almost entirely orange ochreous, thereby resembling in colour typical females of A. gilviventris , although they lack the anteroventrolateral pigmented spots as do the palest A. gilviventris females from populations in Canada: Alberta (see below). A singularly dark male (Nova Scotia: Cranberry I., Lockeport) has the scape and pedicel brown, the 1st nagellomere is anterodorsally brown on the medial surface, the tarsi are gradually brownish-darkened apically, the distal third of the tibiae is also somewhat darkened, the haltere is almost entirely brownish, and the darkened gonostylus is natly blunt apically (as in Fig. 473 ).

Discussion. Anthomyza shewelli sp. nov. is one of three species (see above) forming the A. tschirnhausi group. Because all these species are very similar in male and female postabdominal structures, and because of a similar variability in colour and other external features, it is sometimes difncult to differentiate them. In the male sex, A. shewelli can be recognized safely by the shape of the gonostylus, more precisely by its convex anterior margin (usually visible also in air-dried specimens), despite its variably ending apex (see also the key). The identincation of females is complicated by the high variability of pigmentation of the abdominal sclerites, including T7+S7. Females of A. shewelli and A. gilviventris differ from the Palaearctic A. tschirnhausi structurally in the narrower (and more tapered) and less setose postabdominal 7th and 8th segments. Furthermore, dark females of A. shewelli and the darkest females of A. gilviventris have postabdominal sclerites very similarly formed and pigmented (see Figs 502, 503 , 515 ), and the palest females of A. shewelli (particularly those dried from specimens preserved in alcohol) with largely ochreous preabdominal terga very closely resemble the palest females of A. gilviventris . In these cases, A. shewelli females can be differentiated from those of A. gilviventris by their spermathecae, which are as long as or longer than S10, relatively narrower and more densely striated ( Figs 479, 480 vs. Figs 494, 498 , 501, 505 , 514 ); see also below. Because the identincation of A. shewelli based on characters of the spermathecae is not practical (requiring study under high magnincation, although relative size can sometimes be seen through the pale tergites of intact specimens), external characters are often useful and can be examined nrst, if exposed and visible. Anthomyza shewelli females are typically smaller, less densely microtomentose and hence more shining (particularly anteriorly on mesonotum), with relatively dark or dark-spotted preabdominal terga, T7+S7 blackish brown posteriorly with the lateroventral lobes not darkened, and a lateral ridge present on T7+S7. If the lateral ridge on T7+S7 is not evident, only the remaining specimens that are duller, larger and with either a pale abdomen or with a completely dark T7+S7, need to be dissected.

As given above under the diagnosis of the A. tschirnhausi group, A. shewelli is most closely related to A. gilviventris based on the shared strongly narrowed terminal segments of the female postabdomen, a relatively longer (compared with T8) female S8, more reduced setae on the female T8 and S8, and 5–8 spines on the saccus of the distiphallus.

Etymology. This common species is named in honour of G. E. Shewell, whose preliminary work on Nearctic Anthomyzidae has innuenced species concepts in the present monograph and previous published revisions by the authors.

Biology. Those collection records with habitat details are mostly from the east (see Distribution) and frequently mention damp to wet sites supporting a variety of plants. Open wet areas are represented by “cranberry bog”, “overgrown wet shrubby Sphagnum bog”, “peat bog meadow”, “fen”, “ sedge fen”, “fen nat”, “margins of fen pools”, and “riparian”, areas. Specinc mention is made of sedges ( Carex , Scirpus ) and grasses ( Calamagrostis , Phalaris ), horsetails ( Equisetum ) or other plants ( Juncus , Iris ) of open wet sites. The type locality (Ontario: ~ 74 km NNE Thessalon, Fig. 459 ) for A. shewelli (and A. furvifrons ) is probably representative of these sites where various Carex spp. predominate the shoreline plant community, with C. aquatilis being most abundant. But there are also several references to wooded areas such as “forest noodplain”, “wet hardwood forest”, “forested slope”, “wet forest trail”, “forest edge”, “mixed wood”, and “swampy woods”. These latter collections include a wider variety of plants of which the graminoids are probably the most likely host plants.

A particularly large pair of series was taken from Carex gynandra within an alder ( Alnus ) thicket (Ontario: Sault Ste. Marie, city limits), and is notable for the absence of A. gilviventris since these two species are often represented in mixed series. There are three additional independent collections of A. shewelli from C. gynandra while there are no records for A. gilviventris on this plant. There is a suggestion of spatial separation of these two species at one of these three sites (Ontario: Sault Ste. Marie – Finn Hill, Fig. 421 ). Here, both species occurred together in an open meadow on Carex stipata var. stipata but A. gilviventris heavily predominated in a neighbouring open area with a thick growth of Scirpus microcarpus and soil that was usually at or near saturation. On the opposite side of the meadow, bordering the aspen ( Populus tremuloides ) woodlot, A. shewelli predominated where C. gynandra was found in a graminoid and herb mix. Altogether this might suggest that, in the east, A. shewelli is more likely to be found in more sheltered or shaded areas on sedges such as C. gynandra while A. gilviventris is more likely to be found in the open on coarser species of sedge. However, exceptions include collections of A. shewelli from the open type locality where only a single female A. gilviventris was found, and those specimens swept from rather isolated clumps of Scirpus cyperinus (a slightly nner species of Scirpus ) in open meadow (Sault Ste. Marie – Finn Hill, Fig. 421 ) where A. gilviventris was not found. The dried shoreline of a drained reservoir (Ontario: ~ 66 km NNW Elliot Lake, Fig. 548 ) supported almost exclusively S. microcarpus but A. shewelli was the only species of the A. tschirnhausi group collected here. Factors such as light intensity, soil saturation, and available plant species may contribute to spatial separation of these two species but their relative importance may only become apparent where strong populations of both nies occur in close proximity.

Of the collections from western regions where A. shewelli is either absent or poorly sampled, only those from Manitoba have provided suggestions of habitat and these similarly involve graminoids alone (Falcon Lake) or mixed with Equisetum and Impatiens (Shilo) , as well as “vegetation at spring” (Aweme). A single male was collected in Saskatchewan (Beaver Creek Cons. Area) from “mostly grasses under Betula / Populus ”. The night period for A. shewelli runs from 2 May (Ontario: Midland; St. Ignace Island) to 30 August (Ontario: Manitoulin Is., 0.7 km N Michael’s Bay Pk.).

Distribution. Anthomyza shewelli has a fairly broad distribution in eastern North America from Newfoundland to North Carolina, but only as far west as Saskatchewan, Montana and Idaho. Records are available for Canada: Manitoba, New Brunswick, Newfoundland, Nova Scotia, Ontario, Quebec, Saskatchewan; United States of America: Idaho, Indiana, Massachusetts, Michigan, Minnesota, Montana, New Hampshire, New York, North Carolina, Ohio, Virginia, West Virginia, Wisconsin (see Table 2).

Figures

Figs 2–3. Nearctic species of the genus Anthomyza. 2 – A. shewelli sp. nov., a pair (male on left) on leaf of Carex sp. (Canada:Ontario:Dubreuilville, Magpie River), body length ca.2.4 mm (male) and 3.1 mm (female); 3 –A.mcalpinei sp. nov., living male, laterally (Canada:Ontario:Sault Ste. Marie, Finn Hill), body length ca.2.8 mm. Photo by J. Roháček.

Figs 456–459. Anthomyza shewelli sp. nov. and its habitat. 456 – A. shewelli sp. nov. in copula on Equisetum stem, body length ca. 2.4 mm and 3.1 mm; 457 – same, male laterally, body length ca. 2.5 mm (Canada: Ontario: Pancake Bay Prov. Park); 458 – female on leaf of Carex, body length ca. 3.2 mm (Ontario: Dubreuilville env.); 459 – marshy riverbank of Mississagi River (Canada: Ontario) with predominant Carex and Equisetum, habitat of A. shewelli in the type locality. Photo by K. N. Barber (Fig. 459) and J. Roháček (others).

Figs 460–466. Anthomyza shewelli sp. nov., paratype male, typical form (Canada: Ontario). 460 – external genitalia, caudally; 461 – apex of nlum, subventrally; 462 – external genitalia, laterally; 463 – hypandrial complex, laterally; 464 – transandrium, caudally; 465 – aedeagal complex, laterally; 466 – gonostylus, ventrolaterocaudally (widest extension). Scales = 0.05 mm (Figs 461, 466) and 0.1 mm (others).

Figs 467–473. Anthomyza shewelli sp. nov., paratype male, darkest form (USA: Idaho). 467 – external genitalia, caudally; 468 – apex of nlum, subventrally; 469 – external genitalia, laterally; 470 – hypandrial complex, laterally; 471 – transandrium, caudally; 472 – aedeagal complex, laterally; 473 – gonostylus, ventrolaterocaudally (widest extension). Scales = 0.05 mm (Figs 468, 473) and 0.1 mm (others).

Figs 474–480. Anthomyza shewelli sp. nov., paratype female, typical form (Canada: Ontario). 474 – postabdomen, dorsally; 475 – the same, ventrally; 476 – distal part of female genital chamber, laterally; 477 – end of postabdomen with internal sclerites, laterally; 478 – internal sclerites and S8, ventrally (micropubescence omitted); 479 – spermathecae; 480 – spermatheca of another specimen. Scales = 0.2 mm (Figs 474, 475), 0.1 mm (Figs 476–478) and 0.05 mm (Figs 479, 480).

Figs 481–484. Primary types and females of the Nearctic species of the Anthomyza tschirnhausi group. 481 – A. shewelli sp. nov., holotype male, laterally, body length 2.30 mm; 482 – A. gilviventris sp. nov., dark form, paratype female, laterally, body length 3.42 mm (USA: Colorado); 483 – A.gilviventris sp. nov., holotype male, laterally, body length 2.89 mm; 484 – same species, topotypic paratype female, laterally, body length 3.57 mm. Photo by J. Roháček.

Figs 485–486. Wings of the Nearctic species of the Anthomyza tschirnhausi group. 485 – A. shewelli sp. nov., paratype male, wing length 2.7 mm (Canada: Ontario); 486 – A. gilviventris sp. nov., paratype male, wing length 3.0 mm (Canada: Ontario). Photo by K. N. Barber.

Figs 508–515. Gonostyli (Figs 508–512) of paratypes ventrolaterocaudally (in widest extension) and female postabdominal structures (Figs 513–515) of the Nearctic species of the Anthomyza tschirnhausi group. 508 – A. shewelli sp. nov.(Canada:Ontario); 509 – same species (USA:Montana); 510 – A.gilviventris sp.nov.(Canada:British Columbia); 511, 512 – same species (USA: Colorado); 513 – A. gilviventris sp.nov., paratype female,dark form (USA:Colorado), 8th abdominal segment with internal sclerites, laterally; 514 – same specimen, spermatheca; 515 – same specimen, 7th and 8th abdominal segments, dorsally. Scales = 0.1 mm (Fig. 513), 0.2 mm (Fig. 515) and 0.05 mm (others).

Figs 501–507. Anthomyza gilviventris sp. nov., dark form, paratype female (USA: Montana). 501 – spermatheca; 502 – postabdomen, dorsally; 503 – the same, ventrally; 504 – distal part of female genital chamber, laterally; 505 – spermatheca; 506 – end of postabdomen with internal sclerites, laterally; 507 – internal sclerites and S8, ventrally (micropubescence omitted). Scales = 0.05 mm (Figs 501, 505), 0.2 mm (Figs 502, 503) and 0.1 mm (others).

Figs 494–500.Anthomyza gilviventris sp. nov., paratype female (Canada:Ontario). 494 – spermatheca; 495 – postabdomen, dorsally; 496 – the same, ventrally; 497 – distal part of female genital chamber, laterally; 498 – spermatheca; 499 – end of postabdomen with internal sclerites, laterally; 500 – internal sclerites and S8, ventrally. Scales = 0.05 mm (Figs 494, 498), 0.2 mm (Figs 495, 496) and 0.1 mm (others).

Figs 419–421. Living Anthomyza gibbiger sp. nov. and its habitat. 419 – A. gibbiger sp. nov., male, pale form, dorsally, body length ca. 2.6 mm; 420 – same specimen, laterally (Canada: Ontario: Sault Ste. Marie, Finn Hill); 421 – wet meadow at Sault Ste. Marie (Finn Hill), habitat of three species of the A. neglecta group, viz. A. gibbiger, A. orthogibbus and A. dichroa. Photo by J. Roháček.

Figs 547–550. Anthomyza vulgaris sp. nov. and some of its habitats. 547 – bubbling female on Carex leaf laterally, body length ca. 2.8 mm (Canada: Ontario: Sault Ste Marie); 548 – pure growth of Scirpus microcarpus on exposed lake bottom (Canada: Ontario: Rocky Island Lake); 549 – graminoid vegetation on alvar near Misery Bay (Canada: Ontario: Manitoulin I.); 550 – anterior half of the same female as in Fig. 547, dorsally. Photo by J. Roháček.