Scydmaenus (Corbulifer) pseudorobustus Franz

Jałoszyński, Paweł, 2018, ' Monster Scydmaenus' of Australia: revision of subgenus Corbulifer Franz (Coleoptera: Staphylinidae: Scydmaeninae), Zootaxa 4472 (3), pp. 505-531: 516-517

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Scydmaenus (Corbulifer) pseudorobustus Franz


Scydmaenus (Corbulifer) pseudorobustus Franz  

( Figs 30–32 View FIGURES 30–32 , 58 View FIGURES 55–65 , 69 View FIGURES 66–69 , 70 View FIGURE 70 )

Scydmaenus (Corbulifer) pseudorobustus Franz, 1975: 288   , figs 266–267.

Type material. HOLOTYPE: ( AUSTRALIA, QUEENSLAND): ♂, six labels: "Deviding {sic!} Range / Highway to Warwick / Queensld., Austr." [white, printed, with handwritten "Pa82" in blue ink on the reverse side], " Scydmaenus   / ( Corbulifer   ) / pseudorobustus   / det.H.Franz" [white, handwritten and printed], " ♂ " {white, printed], " Typus " [red, handwritten], " SAMA Database / No. 25-037032" [white, printed], " SAMA Digital Image / 21.Aug 2013 " [green, printed and handwritten] ( SAM).

Additional material (2 exx.): QUEENSLAND: 12km SE of Rathdowney Mt Chinighee , 28°19'S, 152°58'E, 720 m, 17.12.1982, Monteith, Yeates & Thompson, QM Berlesate 496, rainforest litter (QM) GoogleMaps   ; NEW SOUTH WALES: 1 ♂, 28.29S 152.23E, Tooloom Scrub Floral Res., rest area, 755m, Beaury SF, 23km W of Urbenville , 16 Jun. 1993, D. S. Chandler, Berlesate ANIC 1648 View Materials , subtropical closed forest Dendrocnide excelsa   (giant stinger) litter ( ANIC) GoogleMaps   .

Revised diagnosis. Male metatibia gradually and moderately broadened, indistinctly broader than metafemur, broadest distinctly distad middle, lateral (external) tibial margin bisinuate with a distinct submedian concavity, mesal tibial margin concave in proximal 1/5 and nearly evenly convex in distal 4/5, dorsolateral glandular 'pocket' present, with mesal margin bearing long setae; aedeagus in dorsal view broadest near base, with apical portion not delimited, apex divided into two very long and pointed lateral projections strongly bent dorsoproximad.

Redescription. Body of male ( Fig. 30 View FIGURES 30–32 ) strongly convex, reddish brown, covered with setae slightly lighter than cuticle; BL 1.83–1.88 mm (mean 1.85 mm).

Head broadest at very small eyes, HL 0.35–0.38 mm (mean 0.37 mm), HW 0.40; tempora in lateral view as long as 5 × length of eye, only slightly convergent posterad and posteriorly bent mesad at a slightly obtuse angle. Punctures on frons and vertex very fine and shallow, inconspicuous; setae short, sparse and suberect. Antennae slender, AnL 0.95–1.03 mm (mean 0.99 mm), antennomeres I –VI each distinctly elongate, VII and VIII each about as long as broad, IX and X much larger than VIII, each slightly transverse, XI indistinctly longer than IX –X combined, nearly twice as long as broad.

Pronotum rounded, broadest slightly behind anterior third; PL 0.50–0.53 mm (mean 0.51 mm), PW 0.45–0.48 mm (mean 0.46 mm). Antebasal pits indistinct. Punctures on pronotal disc very fine and shallow, inconspicuous; setae short, sparse and suberect.

Elytra oval, broadest in front of middle; EL 0.98 mm, EW 0.73 mm, EI 1.34; apices separately rounded. Punctures relatively large and dense but very shallow, inconspicuous; setae longer and denser than those on pronotum, suberect to erect.

Metatibia ( Fig. 58 View FIGURES 55–65 ) in dorsal view gradually and moderately broadened, broadest between middle and distal third, mesal margin concave in proximal 1/5, remaining portion nearly evenly convex, lateral (external) margin bisinuate with distinct submedian concavity; dorsolateral 'pocket' present, with mesal margin bearing long setae.

Aedeagus ( Figs 31–32 View FIGURES 30–32 ) moderately slender; AeL 0.43 mm; in dorsal view median lobe broadest near base, relatively strongly narrowing distad with bisinuate lateral margins and broad, subtrapezoidal apical portion; apical projections conspicuously long, bent dorsoproximad at a sharp angle.

Female. Unknown.

Distribution. Queensland and New South Wales; central-eastern coast of Australia ( Fig. 70 View FIGURE 70 ).

Remarks. Males of several species have the metatibiae modified in a similar way as those of S. pseudorobustus   : S. eungellanus   , S. gibraltar   , S. kiwarrakianus   and S. gushi   (all described as new). All of them have the metatibiae gradually swollen and with a dorsolateral 'pocket' which divides the tibia into proximal and distal portions. In S. pseudorobustus   the lateral (external) metatibial margin is very weekly concave at the 'pocket', in all remaining similar species the concavity is much deeper, so that the tibia appears recurved. Moreover, in S. pseudorobustus   the proximal portion of metatibia is distinctly longer than the distal one, and consequently the tibia is broadest in its distal half. Only in males of S. eungellanus   the proximal portion of metatibia is indistinctly longer than the distal portion, but adults of this species have the metatibiae of a different shape than those in S. pseudorobustus   . Aedeagi of all these species differ markedly in the shape of the median lobe and apical structures.


South Australia Museum


South African Museum


Australian National Insect Collection














Scydmaenus (Corbulifer) pseudorobustus Franz

Jałoszyński, Paweł 2018

Scydmaenus (Corbulifer) pseudorobustus

Franz 1975: 288