Sanguinobolus Wesener, 2009

Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra, 2009, Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda), ZooKeys 19 (19), pp. 1-128 : 12-14

publication ID 10.3897/zookeys.19.221

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Sanguinobolus Wesener

gen. nov.

Sanguinobolus Wesener View in CoL , gen. n.

Type species: Sanguinobolus maculosus View in CoL sp. n., monotypic

Diagnosis: distinguished by male tarsal pads less prominently developed than in other large-bodied Malagasy genera of Spirobolida ( Figs 7C, D View Figure 7 ), and by anterior telopodite process with a wide and apically swollen tip, unlike in any other Malagasy spirobolid genus ( Figs 6A, B View Figure 6 ). Sperm canal on posterior gonopod discharging into membranous area of telopodite main branch ( Fig. 6D View Figure 6 ). Shares a large body size (> 100 mm) and an aposematic red-black colour pattern with Aphistogoniulus Silvestri, 1897 , Madabolus Wesener & Enghoff, 2008 , Corallobolus and Colossobolus gen. n.. Differs from Aphistogoniulus Silvestri, 1897 and Colossobolus gen. n. by the absence of a mesal branch on the posterior gonopod telopodite ( Fig. 6C View Figure 6 ). The sperm canal in Sanguinobolus runs through the lateral branch, in Aphistogoniulus and Colossobolus gen. n. through the mesal branch. Differs from Colossobolus gen. n. in the absence of a central membranous area basally to the main branch of the posterior gonopod telopodite. Sanguinobolus resembles Colossobolus gen. n. in the presence of a short and wide mesal process on

the coxite of the anterior gonopod which is shorter or only slightly longer than the telopodite and sternite.

Distribution and ecology: only known from the isolated rainforest in Montagne d’Ambre, north Madagascar ( Fig. 5 View Figure 5 ). Species of the genus are probably active on the soil surface because of their relatively long legs and aposematic colour pattern.

Etymology: Sanguinobolus , masculine, is composed of sanguinus, blood, which refers to the blood-red colour pattern of the single constituent species, and - bolus a common suffix for genera of the order Spirobolida .

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