Asytesta allisoni Setliff, 2012

Setliff, Gregory P., 2012, 3462, Zootaxa 3462, pp. 1-125 : 77-100

Asytesta humeralis species group

Asytesta allisoni Setliff , new species

( Figures 133–134, 193)

Diagnosis. The anterolateral region of pronotum in front of the procoxae is uniquely clothed in this species with a solid patch of densely distributed white scales that encompasses the entire area from the prosternal canal to the anterior margin of the procoxae. In all other species, these areas either bear a ring of whitish scales with darker background scales in the center or completely lack white or pale scales. Another unique feature of the vestiture is the large black sutural spot near the base of the elytra. These characters will separate A. allisoni from A. lugubris and A. morobeana new species that are otherwise similarly clothed.

Description. Measurements (n = 2): body length 4.7–4.8 mm (mean 4.8 mm), body width 2.2–2.5 mm (mean 2.4 mm), pronotal length 1.8–1.9 mm (mean 1.9 mm), elytral length 2.9 mm, rostral length 1.4–1.5 mm (mean 1.5 mm). Body oval, 1.9–2.2 X longer than broad; integument dark reddish-brown, antennae and tarsi reddish-brown. Densely covered with creamy white (in maculae) to light brown to fuscous scales. Frons, pleura, coxae, and venter more densely clothed in light brown scales; pair of small yellowish-white maculae on temples (as in Figure 104).

Legs with alternating bands of light and dark brown scales, background vestiture interspersed with white, hairlike setae; male with short, suberect, hair-like setae along ventral margin of protibia near apex, setae longer than elsewhere on legs; tarsi clothed with sparse, white, hair-like setae. Dorsum not mottled or dappled with poorly defined, irregular white patches; pronotal disk with pair of large almost black patches at posterior margin on either side of midline, 4–6 irregular dark, almost black maculae in transverse band slightly in front of midline, reaching lateral margins in most specimens, maculae sometimes fused into transverse band (best viewed without magnification), 8–12 small, circular, whitish maculae arranged in three transverse rows of 2–4 maculae each, one row of 2–4 maculae before middle, one pair at middle, one row of 2–4 macular after middle; pair of similarly formed but smaller maculae at apical pronotal margin, sometimes absent; maculae not in sharp contrast to background squamae; anterolateral region of pronotum with very distinct, well defined, solid patch of densely distributed white scales encompassing entire area from lateral margin of prosternal canal to anterior margin of procoxae; dorsal portion of maculae not extending beyond middle of procoxa. Elytra with anterior margin bearing thin band of creamy white to light brown scales; large, irregular, patch of nearly black scales on disk from just posteriad of anterior margin, to slightly before midline, spanning basal area between intervals 3. Scutellum clothed with creamy white scales. Dorsum lacking other vittae or maculae. Mesosternum and metasternum covered with sparse creamy white to tan scales except transverse glabrous line at suture between sclerites, ventrites with sparse, evenly distributed pale scales. Scales course (individually distinct under low magnification), appressed to suberect.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes (as in Figure 104), interrupted at back by glabrous V-shaped area where head articulates on pronotum, thin, medial longitudinal carinae within crown from apex of V-shaped area to center of crown. Eyes weakly convex, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.2–1.4 X broader than long, broadest at base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one tan to dirty white, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle slightly cavernous, lateral margins produced, posterior margin slightly open. Scutellum visible, diameter subequal to punctures at base of elytra. Elytra 1.2–1.3 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri slightly produced laterally, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before midline; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, not elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs moderately long, hind femora exceeding elytral apices by 1/3 their length; mesofemora exceeding elytral apices. Profemora unidentate; profemoral tooth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange distinct, tapering to apex, male flange terminating before tibial apex, female flange tapering to apex (as in Figures 47, 50); dorsal margin weakly convex. Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with distinct supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on significantly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male and female terminalia. As in generic description.

Material examined. Holotype: ♂, " Tree #50, Tray #3, Lithocarpus rufovillosus, A. Allison, Coll., Bishop Museum / Papua New Guinea: Morobe Prov.: Biaru Rd. ; 1970 m, 3.X.1988 / No. 43613, PNG Canopy , Bishop Museum " ( BPBM) . Paratypes: 1 ♀, " Tree #50, Tray #10, Lithocarpus rufovillosus, A. Allison, Coll., Bishop Museum / Papua New Guinea: Morobe Prov.: Biaru Rd. , 1970 m, 3.X.1988 / No. 43841, PNG Canopy, Bishop Museum" ( BPBM) ; 1 ♂, " New Guinea: Wau , Edie Ck., 2000–2150 m. 6.IV.1965 / J. & M. Sedlacek Collectors Bishop Museum " ( BPBM) . 2 ♀♀, " Mt. Kaindi , N. Guinea II.13–III.12.1979, 2300m. J. Sedlacek " ( CWOB) .

Distribution. Papua New Guinea: Morobe Province: Wau area.

Plant associations. Two specimens of this species were taken by kerosene fogging of Lithocarpus rufovillosus (Markgr) Rehder (Fagaceae) .

Etymology. This species is named for Allen Allison who collected the holotype and a paratype of this handsome weevil during canopy fogging studies at Wau Ecology Institute.

Asytesta antica Pascoe

( Figures 135–136, 194)

Asytesta antica Pascoe, 1883: 100 . Holotype in BMNH, labeled “Type. H.T [on disk with red boarder]/ Kaioa [on blue ovalshaped disk]/ A. antica / Pascoe Coll., 93–60.” Type locality: Kaioa Island.

Diagnosis. This species is immediately recognizable by the uniform brown vestiture that lacks maculae or vittae except for the large irregular patch of white scales entirely covering the cervical region of the pronotal disk from the anterior margin to about the midline and terminating on the sides. This species also has the protibial flange very weakly developed and bidentate profemora.

Redescription. Measurements (n = 5): body length 5.0– 5.6 mm (mean 5.2 mm), body width 2.3–3.0 mm (mean 2.6 mm), pronotal length 2.0– 2.2 mm (mean 2.1 mm), elytral length 3.0– 3.5 mm (mean 3.2 mm), rostral length 1.5–1.8 mm (mean 1.7 mm). Body oval, 1.9–2.1 X longer than broad; integument reddish-brown to dark brown, antennae and tarsi reddish-brown. Densely covered with brown to fuscous scales, with more densely distributed dirty white scales in dorsal pronotal maculae; maculae in sharp contrast to background squamae. Frons, pleura, coxae, and venter more densely clothed in tan to brown scales; vestiture on femora and tibia interspersed with white, hair-like setae; male with short, suberect, hair-like setae along ventral margin of protibia near apex, setae longer than elsewhere on legs; tarsi clothed with sparse, white, hair-like setae. Dorsum dark mottled with irregular, maculae of light brown; maculae less densely distributed on sides. Pronotum with large irregular patch of white scales from anterior margin to midline, reaching sides; anterolateral region of pronotum with poorly defined ring of denser brown scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with sparser background scales in center; dorsal portion of ring not extending beyond middle of procoxa. Elytra lacking vittae or maculae. Mesosternum and metasternum covered with sparse brown scales except transverse glabrous line at suture between sclerites, ventrites with sparse, evenly distributed pale scales. Scales course (individually distinct under low magnification), appressed to suberect.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes ( Figure 104), interrupted at back by glabrous V-shaped area where head articulates on pronotum, medial longitudinal carinae absent. Eyes weakly convex, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.2–1.4 X broader than long, broadest at base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one light brown, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle slightly cavernous, lateral margins produced, posterior margin slightly open. Scutellum visible, minute, diameter subequal to punctures at base of elytra. Elytra 1.1–1.3 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before middle of elytra; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, not elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs moderately long, hind femora exceeding elytral apices by 1/3 their length; mesofemora exceeding elytral apices. Profemora bidentate; apical profemoral teeth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange distinct but weakly produced, tapering to apex, male flange terminating before tibial apex, female flange tapering to apex ( Figures 45–46); dorsal margin weakly convex. Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with distinct supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on significantly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia. Tergite VII broader than long, anterior margin convex, posterior margin emarginate at middle, wing binding patch distinct, with two rows of 5–6 minute plectral tubercles reaching posterior margin, tubercles greatly reduced, distinct under high magnification.

Female terminalia. Tergite VII longer than broad, anterior and posterior margins broadly convex, wing binding patch distinct, with two rows of 5–8 minute plectral tubercles; plectral rows not reaching posterior margin; tubercles visible, minute, distinct under high magnification. Bursa more or less indistinct from vagina; vagina with walls moderately sclerotized near junction with oviduct; no distinct internal sclerites visible.

Material examined. Holotype: ♀, "Type. H.T [on disk with red boarder]/ Kaioa [on blue oval-shaped disk]/ A. antica / Pascoe Coll., 93-60" ( BMNH) ; Paratype: 1 ♂, "Kaioa [on blue oval-shaped disk]/ Pascoe Coll. 93-60" ( BMNH) . Non-type material: 1 ♀, "Wallace/ Moluccas / Fry coll., 1905.100/ Asytesta Id. By A.M. Lea , near if not humeralis Pasc. " ( BMNH) ; 1 ♂, "Kaioa" ( BMNH) ; 1 ♂, "Kai Is. [= Kaioa], Coll. Jekel / A. sp./ Museo Genova, coll. Angelo Solari, (acquisto 2000)" ( MSNG) .

Distribution. Indonesia: Kaioa Island [between Halmahera and Bacan].

Remarks. No other specimen in the BMNH or MSNG collections bear any indication of type status. The male specimen that I treat here as a paratype has identical labels as the holotype and is missing its abdomen ( Figure 135).

Asytesta biakana Setliff , new species

( Figures 141–142, 193)

Diagnosis. This species is most easily confused with A. eudyasmoides , A. humeralis and A. marginalis new species, which also have more or less solid, dark, almost black, background coloration and short, white, rectangular humeral maculae as the only significant dorsal markings. All of three of these species have a visible scutellum, which is lacking in A. biakana . This species has unidentate profemora and small white maculae evenly distributed on the elytra, which further distinguishes it from the latter two species. Several other species of Asytesta also have rectangular humeral maculae (e.g., A. rata , A. signata , and A. thompsoni new species); however, these species all have the dorsal background vestiture strongly mottled with light and dark scales.

Description. Measurements (n = 7): body length 4.8–6.3 mm (mean 5.6 mm), body width 2.8–3.2 mm (mean 3.0 mm), pronotal length 2.0– 2.6 mm (mean 2.3 mm), elytral length 2.8–3.8 mm (mean 3.2 mm), rostral length 1.5–2.0 mm (mean 1.8 mm). Body oval, 1.7–2.0 X longer than broad; integument dark reddish-brown to black, antennae and tarsi dark reddish-brown. Sparsely covered with small, brown to fuscous scales, with more densely distributed dirty white scales in maculae. Frons, pleura, coxae, basal portion of femora, and venter more densely clothed in brown to tan scales; pair of small whitish maculae on temples (as in Figure 104). Vestiture on femora and sometimes tibia interspersed with dirty white, hair-like setae; male with short, suberect, hair-like setae along ventral margin of protibia near apex, setae longer than elsewhere on legs; tarsi clothed with sparse, white, hair-like setae. Dorsum dark; with small, regularly distributed maculae of light brown to dirty white scales; maculae composed of 3–5 scales, less densely distributed on sides, absent on pronotum in some specimens. Anterolateral region of pronotum with faint ring of light brown scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with darker background scales in center (as in Figure 104); dorsal portion of ring not extending beyond middle of procoxa. Elytral humeri with distinct, rectangular, maculae of bright white scales on intervals 5 and 6; macula on intervals 5 longer than macula on intervals 6, maculae often absent; lacking other vittae. Mesosternum and metasternum covered with sparse brown to dirty white scales except transverse glabrous line at suture between sclerites, ventrites with sparse, evenly distributed brown to dirty white scales. Scales fine (indistinct under low magnification), appressed to suberect.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes ( Figure 104), interrupted at back by glabrous V-shaped area where head articulates on pronotum, medial longitudinal carinae absent. Eyes weakly convex, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.2–1.4 X broader than long, broadest at base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one brown, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle slightly cavernous, lateral margins produced, closed to slightly open behind. Scutellum not visible. Elytra 1.0–1.2 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before middle of elytra; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, not elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing vestigial.

Legs moderately long, hind femora exceeding elytral apices by 1/3 their length; mesofemora exceeding elytral apices. Profemora unidentate; profemoral tooth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange distinct, tapering to apex, male flange terminating before middle of tibia, female flange tapering to apex ( Figures 47, 50); dorsal margin dorsal margin strongly convex (as in Figure 48–49). Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with distinct supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on significantly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male and female terminalia. As in generic description.

Material examined. Holotype: ♂, " New Guinea: Neth., Biak I.: Kampong, Landbouw, 50–100m. May 28, 1959 / J.L. Gressit, Collector/ T.C. Maa, Collector, Bishop" ( BPBM) . Paratypes: 1 ♀, " Biak Isl. , Korim, Roidifu, ca. 100m., 2.II.2001 / Irian Jaya, leg. A. Riedel " ( MZB) ; 1 ♂, " Biak Isl. , Korim, Roidifu, ca. 100m., 2.II.2001 / Irian Jaya, leg. A. Riedel " ( SMNK) ; 2 ♂♂, 2 ♀♀, " Irian Jaya , Biak Is., Umg. Sepse, leg. A. Riedel 3.X.1990 " ( ARC) .

Distribution. Biak Island.

Etymology. The specific epithet refers to the species’ type locality.

Asytesta bidentata Voss , new status

( Figures 143–144, 193)

Asytesta lugubris ssp. bidentatus Voss, 1958: 219 . Holotype, not examined (depository unknown). Type locality: Deslacs Islands.

Diagnosis. This species was originally described as a subspecies of A. lugubris Heller. The two species are superficially similar; however, A. bidentata differs from A. lugubris by the distinctly bidentate profemora, lack of pale macula on the temples above the eyes, and more distinct humeral macula that stand out from the background vestiture, with part of the macula reaching the sixth elytral interval. The vestiture of A. lugubris is also much darker than in this species.

Redescription. Measurements (n = 1): body length 5.1 mm, body width 2.7 mm), pronotal length 2.1 mm, elytral length 3.0 mm, rostral length 1.8 mm. Body oval, 1.9 X longer than broad; integument dark reddish-brown, antennae and tarsi reddish-brown. Densely covered with tan to dark brown scales. Frons, pleura, coxae, and venter more densely clothed in dirty white to tan scales; lacking maculae on temples.

Legs with alternating bands of tan and dark brown scales, background vestiture interspersed with tan, hair-like setae; male with short, suberect, hair-like setae along ventral margin of protibia near apex, setae longer than elsewhere on legs; tarsi clothed with sparse, tan, hair-like setae. Dorsum mottled with large, poorly defined, irregular patches of brown scales; pair of larger dark patches at posterior margin of pronotal disk on either side of midline; pronotal disk with faint dorsolongitudinal vitta of denser tan scales from apex to base; 6 irregular dark maculae in transverse band slightly in front of midline reaching sides (best viewed without magnification), 6 small, circular, whitish maculae arranged in two transverse rows of 4 maculae each, one row before middle, one row after; lacking pair of maculae at apical pronotal margin in single specimen examined, vitta and maculae not in sharp contrast to background squamae; anterolateral region of pronotum with poorly defined ring of denser dirty white scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with darker background scales in center (as in Figure 104); dorsal portion of ring not extending beyond middle of procoxa. Elytral humeri with indistinct, rectangular, maculae of dense tan white scales on intervals 5 and 6; macula on intervals 5 longer than macula on intervals 6. Dorsum lacking other vittae or maculae. Mesosternum and metasternum covered with sparse tan scales except transverse glabrous line at suture between sclerites, ventrites with sparse, evenly distributed tan scales. Scales course (individually distinct under low magnification), appressed to suberect.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes ( Figure 104), interrupted at back by glabrous V-shaped area where head articulates on pronotum, thin, medial longitudinal carinae within crown from apex of V-shaped area to center of crown. Eyes weakly convex, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.3 X broader than long, broadest at base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one brown, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle slightly cavernous, lateral margins produced, posterior margin slightly open. Scutellum visible, minute, diameter smaller than punctures at base of elytra. Elytra 1.1 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated near basal margin; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, not elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs moderately long, hind femora exceeding elytral apices by 1/3 their length; mesofemora exceeding elytral apices. Profemora bidentate; apical profemoral tooth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia weakly compressed laterally, ventral margin with flange weakly developed, tapering to apex, male flange terminating before middle of tibia (as in Figure 47); dorsal margin weakly convex. Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with distinct supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on significantly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia. Tergite VII broader than long, anterior margin convex, posterior margin emarginate at middle, wing binding patch distinct, with two rows of 5 minute plectral tubercles reaching posterior margin, tubercles greatly reduced, distinct under high magnification.

Female terminalia. Not examined.

Material examined. ♂, " I. Deslacs, Biro 1901/ Paratype, Asytesta lugubris ab. bidentatus, Voss" ( SMTD) .

Distribution. Vitu Islands (= Witu Islands and Deslacs Islands).

Remarks. Voss (1958: 219) mentions 10 specimens of this species in his description. I was only able to examine a single specimen labeled paratype.

Asytesta bivirgata Pascoe

( Figures 145–146, 194)

Asytesta bivirgata Pascoe, 1885: 259 . Lectotype, here designated, in MSNG. Labeled “N. Guinea, Hatam VI, Beccari 1875/ Typus/ Asytesta bivirgata , typus! Pasc./ bivirgata, Pasc. ” Type locality: Hatam.

Diagnosis. Asytesta bivirgata belongs with A. lugubris and its allies. It has broad, divergent, sublateral vittae on the pronotal disk that meet the short humeri maculae on elytral intervals 5 and 6, which are themselves strongly bifurcated and terminate before the middle of the elytra. These characters will distinguish this species from its congeners.

Redescription. Measurements (n = 15): body length 4.9–6.7 mm (mean 5.4 mm), body width 2.0– 3.2 mm (mean 2.6 mm), pronotal length 1.6–2.9 mm (mean 2.2 mm), elytral length 2.3–3.8 mm (mean 3.1 mm), rostral length 1.3–2.4 mm (mean 1.8 mm). Body oval, 1.2–2.2 X longer than broad; integument dark reddish-brown to black, antennae and tarsi reddish-brown. Sparsely covered with light brown to fuscous scales, with more densely distributed dirty white to yellowish-white scales in pronotal maculae and vittae. Frons, pleura, coxae, and venter more densely clothed in tan to brown scales; pair of small whitish maculae on temples (as in Figure 104).

Legs with alternating bands of tan and dark brown scales, interspersed with white, hair-like setae; male with short, suberect, hair-like setae along ventral margin of protibia near apex, setae longer than elsewhere on legs; tarsi clothed with sparse, white, hair-like setae. Dorsum mottled with irregular patches of light brown scales; patches less densely distributed on sides. Pronotal disk with pair of sublateral longitudinal vittae from anterior margin to base, strongly divergent past cervical constriction; 2–6 evenly distributed, small, circular maculae arranged in transverse rows; maculae in sharp contrast to background squamae; anterolateral region of pronotum with poorly defined ring of denser brown scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with sparser background scales in center (as in Figure 104); dorsal portion of ring not extending beyond middle of procoxa. Elytral humeri with distinct, rectangular, maculae of bright white scales on intervals 5 and 6; macula on intervals 5 much longer than on intervals 6, terminating just before midline, strongly bifurcated at posterior end, with one tail continuing on fourth intervals and other tail continuing on fifth intervals; lacking other vittae or maculae. Mesosternum and metasternum covered with sparse brown scales except transverse glabrous line at suture between sclerites, ventrites with sparse, evenly distributed brown scales. Scales course (individually distinct under low magnification), appressed to suberect.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes ( Figure 104), interrupted at back by glabrous V-shaped area where head articulates on pronotum, thin, medial longitudinal carinae within crown from apex of V-shaped area to center of crown. Eyes weakly convex, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.1–1.3 X broader than long, broadest at base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one light brown to dirty white, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle slightly cavernous, lateral margins produced, posterior margin slightly open. Scutellum visible, minute, diameter subequal to punctures at base of elytra. Elytra 1.0–1.4 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated near basal margin; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, not elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs moderately long, hind femora exceeding elytral apices by 1/3 their length; mesofemora exceeding elytral apices. Profemora unidentate; profemoral tooth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange distinct, tapering to apex, male flange terminating before tibial apex, female flange tapering to apex (as in Figures 47, 50); dorsal margin moderately convex. Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with distinct suprauncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on significantly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia. Tergite VII broader than long, anterior margin convex, posterior margin emarginate at middle, wing binding patch distinct, with two rows of 6–7 minute plectral tubercles reaching posterior margin, tubercles greatly reduced, distinct under high magnification.

Female terminalia. Tergite VII longer than broad, anterior and posterior margins broadly convex, wing binding patch distinct, with two rows of 9–10 minute plectral tubercles; plectral rows not reaching posterior margin; tubercles visible, minute, distinct under high magnification. Bursa more or less indistinct from vagina; vagina with walls weakly sclerotized; no distinct internal sclerites visible.

Material examined. Lectotype (here designated): ♂, "N. Guinea, Hatam VI, Beccari 1875/ Typus/ Asytesta bivirgata , typus! Pasc./ bivirgata, Pasc. " ( MSNG) . Paralectotype (here designated): 1 ♂, " N.G. Hatam / Asytesta bivirgata Pasc. " ( BMNH) . Non-type material: 1 ♂, " Testega , 1100–1200 m, 11.IV/ IRIAN JAYA, Manokwari- Prov., leg. A. Riedel, 1993" ( ARC) ; 1 ♀, " Testega , 1100–1200 m, 11.IV/ IRIAN JAYA, Manokwari-Prov., leg. A. Riedel, 1993" ( GPSC) ; 1 ♂, " Testega , 1100–1300m, 30.III–2.IV/ IRIAN JAYA, Manokwari-Prov., leg. A Riedel, 1993" ( ARC) ; 1 ♂, " Tetaho-area , Kosmena, 1400–1750m, 26-28.III./ IRIAN JAYA, Manokwari-Prov., leg. A Riedel, 1993" ( ARC) ; 1 ♂, " Epomani , Ugida, Km. 179, 1350- 1400m, 19–20.I.1996 / IRIAN JAYA, Paniai-Prov., leg. A Riedel, " ( GPSC) ; 3 ♂♂, 3 ♀♀, " Epomani , Ugida, Km. 179, 1350- 1400m, 19–20.I.1996 / IRIAN JAYA, Paniai-Prov., leg. A Riedel, " ( ARC) ; 1 ♂, "IRIAN JAYA, Manokwari-Prov. Ransiki , Membey 800–1200 m, leg. A. Riedel, 31.VIII.1991 " ( ARC) ; 1 ♀, " West New Guinea, Vogelkor; Kebar Val., W of Manokwari, 550m, 4–31.I.1962 / S. Quate, Collector" ( BPBM) .

Distribution. West Papua: Manokwari Province: Mokwam [type locality], Hatam, Kebar Valley, Kosmena, Membey, Ransiki, Testega; Paniai Province: Epomani.

Remarks. Pascoe (1885: 259) did not indicate how many specimens were in the type series. Thus a lectotype is selected for the purpose of fixing the name to a single specimen.

Asytesta brevipennis Faust

( Figures 147–148, 194)

Asytesta brevipennis Faust, 1898: 166 [key], 173 [description]. Lectotype, here designated, in SMTD. Labeled “Mailu, Brit. N. G., July 95., (Anthony)/ brevipennis, Faust / Coll. J. Faust, Ankauf 1900.” Type locality: Mailu Island.

Diagnosis. This small species is more or less clothed uniformly in light brown to fuscous scales mottled with poorly defined, irregular patches of brown scales and lacks major maculae or vittae on the dorsum. There is a pair of larger dark patches at posterior margin of pronotal disk on either side of midline, 4–8 irregular dark maculae in transverse band slightly in front of midline and 6–8 small, circular, whitish maculae arranged in two transverse rows of 2–4 maculae each. The scutellum is absent in all but one specimen examined. It is most similar to A. signata , which also appears to lack a scutellum, however, A. signata is smaller, has 10–14 small, white maculae on the pronotal disk that are more distinct than in A. brevipennis , and has distinct, rectangular, maculae of dense white scales on intervals 5 and 6 at the elytral humeri.

Redescription. Measurements (n = 5): body length 4.8–5.2 mm (mean 5.0 mm), body width 2.5–3.0 mm (mean 2.6 mm), pronotal length 2.1–2.2 mm (mean 2.1 mm), elytral length 2.7–3.0 mm (mean 2.8 mm), rostral length 1.5–1.7 mm (mean 1.6 mm).Body oval, 1.7–1.9 X longer than broad; integument dark reddish-brown, antennae and tarsi reddish-brown. Densely covered with light brown to fuscous scales. Frons, pleura, coxae, and venter more densely clothed in tan to brown scales; pair of small yellowish-white maculae on temples (as in Figure 104).

Legs with alternating bands of tan and dark brown scales, background vestiture interspersed with white, hairlike setae; male with short, suberect, hair-like setae along ventral margin of protibia near apex, setae longer than elsewhere on legs; tarsi clothed with sparse, white, hair-like setae. Dorsum mottled with poorly defined, irregular patches of brown scales; pair of larger dark patches at posterior margin of pronotal disk on either side of midline, 4–8 irregular dark maculae in transverse band slightly in front of midline reaching sides in most specimens (best viewed without magnification), 6–8 small, circular, whitish maculae arranged in two transverse rows of 2–4 maculae each, one row before middle, one row after; pair of similarly formed but smaller maculae at apical pronotal margin, sometimes absent, maculae in sharp contrast to background squamae; anterolateral region of pronotum with poorly defined ring of denser dirty white to tan scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with darker background scales in center (as in Figure 104); dorsal portion of ring not extending beyond middle of procoxa, absent in some specimens. Dorsum lacking other vittae or maculae. Mesosternum and metasternum covered with sparse tan scales except transverse glabrous line at suture between sclerites, ventrites with sparse, evenly distributed tan scales. Scales course (individually distinct under low magnification), appressed to suberect.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes ( Figure 104), interrupted at back by glabrous V-shaped area where head articulates on pronotum, thin, medial longitudinal carinae within crown from apex of V-shaped area to center of crown. Eyes weakly convex, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.2–1.4 X broader than long, broadest at base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one tan to dirty white, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle slightly cavernous, lateral margins produced, posterior margin slightly open. Scutellum not visible except very minute scutellum visible in single specimen, diameter smaller than punctures at base of elytra. Elytra 1.0–1.1 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated near basal margin; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, not elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs moderately long, hind femora exceeding elytral apices by 1/3 their length; mesofemora exceeding elytral apices. Profemora unidentate; profemoral tooth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange distinct, tapering to apex, male flange terminating before tibial apex, female flange tapering to apex (as in Figures 47, 50); dorsal margin weakly convex. Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with distinct supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on significantly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia. Tergite VII broader than long, anterior margin convex, posterior margin emarginate at middle, wing binding patch distinct, with two rows of 7–8 very minute plectral tubercles reaching posterior margin, tubercles greatly reduced, distinct only under high magnification.

Female terminalia. Tergite VII longer than broad, anterior and posterior margins broadly convex, wing binding patch distinct, with two rows of 1–3 very minute plectral tubercles; plectral rows not reaching posterior margin; tubercles greatly reduced, distinct only under high magnification. Bursa more or less indistinct from vagina; vagina with walls weakly sclerotized; no distinct internal sclerites visible.

Material examined. Lectotype (here designated): ♂ "Mailu, Brit. N. G., July 95., (Anthony)/ brevipennis, Faust / Coll. J. Faust, Ankauf 1900" ( SMTD) . Paralectotypes (here designated): 1 ♂, 1 ♀, "Mailu, Brit. N. G., July 95., (Anthony)/ brevipennis, Faust / Coll. J. Faust, Ankauf 1900" ( SMTD) ; 1 ♂, "Mailu, Brit. N. G., July 95., (Anthony)/ Museum Paris, ex. Coll., R. Oberthur " ( MNHN) .

Distribution. Papua New Guinea: Mailu Island.

Remarks. Three additional specimens with identical label data as the lectotype were also examined by me in the MNHN collection but were not borrowed for further study. These specimens are also paralectotypes. Since the number of specimens in the type series is unknown, I select one of the three specimens in Faust’s collection bearing a small square of gold foil as the lectotype in order to fix the name to that specimen.

Asytesta humeralis Pascoe

( Figures 3–8, 23, 30, 37, 45–46, 64, 81–92, 163–164, 193)

Asytesta humeralis Pascoe, 1865: 426 , pl. xvii: Figure 13. Holotype in BMNH, labeled “Type, H.T/ Batchian/ Asytesta humeralis, Pasc. ” Type locality: Bacan Island.

Asytesta definita Faust, 1898: 166 [key], 172 [description], new synonymy. Lectotype, here designated, “Maliase, Boucard/ definita, Faust / Coll. J. Faust, Ankauf 1900.” Type locality: Andai, Halmahera, and Moluccas [see discussion below].

Diagnosis. Asytesta humeralis may be confused with a few other similar species (viz., A. eudyasmoides , A. biakana new species, and A. marginalis new species). All of which have more or less solid, dark (nearly black) background coloration and short, white, rectangular humeral maculae. It differs from A. eudyasmoides by its bidentate profemora and shorter body form. Asytesta biakana and A. marginalis are more similar in shape to A. humeralis than the previous species; however, A. biakana lacks a visible scutellum, has unidentate profemora, and very small white maculae that are evenly distributed on the elytra. Asytesta marginalis is the most closely related species to A. humeralis . In addition to its similar shape and vestiture, already discussed, both species have a visible scutellum and bidentate profemora and have similar protibial shape as well. Asytesta humeralis is distinguished from A. marginalis by the more sparsely distributed, darker brown (fuscous) scales on the dorsum and the distinct and strongly contrasting humeral maculae. Asytesta marginalis has more densely and uniformly distributed light brown to tan scales on the dorsum and greatly reduced or absent humeral maculae. Several other species of Asytesta also have rectangular humeral maculae (e.g., A. rata , A. signata , and A. thompsoni new species); however, these species all have the dorsal background vestiture strongly mottled with light and dark brown scales.

Redescription. Measurements (n = 15): body length 5.0– 7.1 mm (mean 5.9 mm), body width 2.2–3.1 mm (mean 2.8 mm), pronotal length 2.1–3.5 mm (mean 2.4 mm), elytral length 2.9–4.5 mm (mean 3.5 mm), rostral length 1.6–2.0 mm (mean 1.9 mm). Body oval, 1.9–2.3 X longer than broad; integument dark reddish-brown to black, antennae and tarsi reddish-brown. Sparsely covered with small, brown to fuscous scales, with more densely distributed dirty white scales in vittae and maculae, in sharp contrast to background squamae. Frons, pleura, coxae, basal portion of femora, and venter more densely clothed in white to tan scales; vestiture on femora and tibia interspersed with white, hair-like setae; male with short, suberect, hair-like setae along ventral margin of protibia near apex, setae longer than elsewhere on legs; tarsi clothed with sparse, white, hair-like setae. Dorsum dark; occasionally with small, irregular, maculae of light brown to dirty white scales; maculae less densely distributed on sides. Pronotal disk usually with pair of small dirty white maculae on cervical constriction at either side of midline. Anterolateral region of pronotum with well defined ring of dirty white scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with darker background scales in center (as in Figure 104); dorsal portion of ring not extending beyond middle of procoxa. Elytra with thin fascia spanning entire anterior margin in some specimens, margin between intervals 5 bare in most specimens, fasciae reaching epipleural margin when present; humeri with distinct, rectangular, maculae of bright white scales on intervals 5 and 6; macula on intervals 5 longer than macula on intervals 6; lacking other vittae. Mesosternum and metasternum covered with dense brown to dirty white scales except transverse glabrous line at suture between sclerites, ventrites with sparse, evenly distributed pale scales. Scales fine (indistinct under low magnification), appressed to suberect.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes (as in Figure 104), interrupted at back by glabrous V-shaped area where head articulates on pronotum, medial longitudinal carinae absent. Eyes flat, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.1–1.2 X broader than long, broadest subbasally; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one white, hair-like seta ( Figure 3). Prosternal canal deep, lateral walls vertical; mesosternal receptacle slightly cavernous, lateral margins produced, posterior margin slightly open. Scutellum visible, minute, diameter subequal to punctures at base of elytra. Elytra 1.1–1.5 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before middle of elytra; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, not elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs moderately long, hind femora exceeding elytral apices by 1/3 their length; mesofemora exceeding elytral apices. Profemora bidentate; apical profemoral teeth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange distinct, tapering to apex, male flange terminating before middle of tibia, female flange tapering to apex ( Figures 45–46); dorsal margin weakly convex. Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with distinct suprauncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened ( Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on significantly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia ( Figures 81–87). Tergite VII broader than long, anterior margin convex, posterior margin emarginate at middle, wing binding patch distinct, with two rows of 5–8 minute plectral tubercles reaching posterior margin, tubercles greatly reduced, distinct under high magnification.

Female terminalia ( Figures 88–92). Tergite VII longer than broad, anterior and posterior margins broadly convex, wing binding patch distinct, with two rows of 5–8 minute plectral tubercles; plectral rows not reaching posterior margin; tubercles visible, minute, distinct under high magnification. Bursa more or less indistinct from vagina; vagina with walls weakly to moderately sclerotized; no distinct internal sclerites visible.

Material examined. Holotype: ♂, "Type, H.T/ Batchian/ Asytesta humeralis, Pasc " ( BMNH) . Paratypes: 1 ♂, "Batchian/ Asytesta humeralis, Batchian " ( UMO) ; 1 ♀, "Batch., Wall./ Coll., Roelofs/ A. humeralis, Pascoe, D. Mollucas / Type" ( ISNB) . Non-type material: 2 ♂♂, "Wallace/ Moluccas, Gilolo" ( BMNH) ; 1 ♀, " Asytesta humeralis " (ZMUC) ; 1 ♀, " Moluccas / Asytesta I.D. by A.M.Lea" ( BMNH) ; 1 ♂, "Batchian/ Bowring 6347" ( BMNH) ; 1 ♂, "Batchian, Coll. Jekel / Asytesta humeralis , [illegible]" ( MSNG) ; 1 ♂, "19991/ Wallace/ Moluccas, Batchian/ Asytesta humeralis , Moluccas " ( MSNG) ; 1 ♂, "Molueche, Coll. Jekel / Moluccas, Wallace / Asytesta humeralis Pascoe, Ent.II , 426" ( MSNG) ; 1 ♂, "Moluccan, 5801" ( SMTD) ; 1 ♂, "Gorham, Collection, acc. 68498/ Asytesta humeralis Pasc. , [illegible] Archipel. " ( USNM) ; 1 ♂, "Bacan, 6 km N. Labuha / Maluku, leg. R. Gerstmeier, 4.IV.1998 " ( ARC) ; 1 ♀, "Batchian/ Coll. Castelnau, Coll. Roelofs / G: Asytesta humeralis " (ISNB) ; 1 ♀, "44464/ Wallace/ Moluccas, Bachian" ( BMNH) . Material examined of A. definita syn. nov.: Lectotype for A. definita Faust (here designated): 1 ♀, "Maliase, Boucard/ definita, Faust / Coll. J. Faust, Ankauf 1900" ( SMTD) . Paralectotypes: 2 ♂♂, " Asytesta definita, Faust / Coll. J. Faust, Ankauf 1900" ( SMTD) . Non-type material: 1 ♂, " Batch. / Asytesta definita Pasc., Det. K. M. Heller 07." ( BMNH) ; 2 ♀♀, " Museum Paris , Moloque, Gilolo, Rafray Maindron, 1878" ( MNHN) ; 1 ♂, " Dodinga , Halmeheira, Doherty VIII/ G.A.K Marshall, Coll., B.M.1950-255." ( BMNH) ; 2 ♂♂, 2 ♀♀, " Halmahera , Telago paca (sw), Tobelo, 27.3.1955, leg. R. Gerstmeier " ( ARC) ; 1 ♂, " Halmahera , Telago paca (sw), Tobelo, 27.3.1955, leg. R. Gerstmeier " ( GPSC) ; 1 ♂, " Museum Paris , Moloque, Gilolo, Rafray Maindron, 1878" ( MNHN) ; 1 ♂, " Ternate / Museum Paris , 1955, coll. A. Clerc " ( MNHN) ; 1 ♂, " Indonesia: Halmahera , Is. , Jailolo Dist., Kampung Pasir Putih, 53'N, 127 41' E / Ruben Tatu, S. Sasamulore, Paul Taylor, 4-17 Dec. 1984 " ( USNM) ; 1 ♀, "[label illegible]" ( MNHN) ; 1 ♀, "Aru Is., Ville Pol / Asytesta definita Faust, Det. K. M. Heller 1912 " ( MSNG) ; 1 ♀, " Halmahera , Bruijn, 75/ 63/ Asytesta humeralis, Pasc " ( MSNG) ; 1 ♀, " Halmahera , Bruijn, 75/ 154/ Asytesta humeralis Pasc. , det. Pascoe 1881" ( MSNG) .

Distribution. Bacan Island, Moluccas. Pascoe (1885: 259) reported additional specimens of A. humeralis from Ternate and Bacan. Faust (1898: 173) reported A. definita from Halmahera, Andai, and the Moluccas but see my comments below.

Remarks. Asytesta humeralis was not seen by Faust at the time he described A. definita . The characters he cites in his diagnosis of the two species are the relatively larger prothorax compared elytra and the deeper elytral punctures of A. humeralis . I can find no appreciable difference in these characters between the two. Several specimens in the type series of A. definita Faust from Andai (West Papua) belong to an undescribed species that poorly agrees with Faust’s description. I have selected a specimen from Faust’s type series that most closely agrees with his description as the lectotype for A. definita syn. n. in order to fix the name to that specimen. Only specimens that are conspecific with the lectotype (and A. humeralis ) are designated as paralectoypes.

Asytesta lugubris Heller

( Figures 165–166, 193)

Asytesta lugubris Heller, 1895: 15 . Lectotype, here designated, in SMTD. Laeled “ lugubris, Heller / K. Wilhelmland, 8577/ Coll. J. Faust, Ankauf 1900/ Syntypus.” Type locality: Papua New Guinea: Bongu.

Asytesta setipes Lea, 1928: 76 , new synonymy. Lectotype, here designated, in SAM. Labeled “Finschhaven, N. E. New Guinea / Asytesta setipes Lea, N. Guinea, type.” Type locality: Wau Creek [possibly in error, see discussion below].

Diagnosis. The mottled, dark brown vestiture that is not marked with distinctly contrasting pale-colored vittae or maculae will separate A. lugubris from most other species except A. allisoni new species and A. morobeana new species. The vestiture in A. lugubris is typically darker than in these two species and has a faint dorsolongitudinal vitta that reaches the base and indistinct, rectangular, maculae of dense brown scales on intervals 5 of the elytra. Neither A. allisoni nor A. morobeana have entire dorsolongitudinal vittae (restricted to the anterior half in A. morobeana ) or humeral maculae.

Redescription. Measurements (n = 9): body length 5.7–7.1 mm (mean 6.4 mm), body width 3.0– 3.5 mm (mean 3.1 mm), pronotal length 2.4–3.1 mm (mean 2.7 mm), elytral length 3.2–4.0 mm (mean 3.7 mm), rostral length 1.8–2.3 mm (mean 1.9 mm). Body oval, 1.9–2.2 X longer than broad; integument dark reddish-brown, antennae and tarsi reddish-brown. Sparsely covered with brown to fuscous scales. Frons, mesepisternum, coxae, and basal portions of femora more densely clothed in yellowish-white to light brown scales; pair of small yellowish-white maculae on temples (as in Figure 104). Femora and tibia with background vestiture interspersed with white, hair-like setae; male with short, suberect, hair-like setae along ventral margin of protibia near apex, setae longer than elsewhere on legs; tarsi clothed with sparse, white, hair-like setae. Dorsum mottled with poorly defined, faint, irregular dark patches; pronotal disk with faint, dorsolongitudinal vitta of dense brown scales from apical margin to base, pair of small dark patches at posterior margin on either side of midline, 4–6 irregular dark, maculae in transverse band slightly in front of midline, reaching lateral margins in most specimens, maculae sometimes fused into transverse band, often very faint (best viewed without magnification), 6–8 small, circular, whitish maculae arranged in two transverse rows of 2–4 maculae each, one row of 2–4 maculae before middle, one row of 2–4 maculae after middle; pair of similarly formed but smaller maculae at apical pronotal margin, sometimes absent, vitta and maculae not in sharp contrast to background squamae, pronotal maculae barely perceptible; anterolateral region of pronotum with poorly defined ring of denser dirty white to light brown scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with darker background scales in center (as in Figure 104); dorsal portion of ring not extending beyond middle of procoxa. Elytral humeri with indistinct, rectangular, maculae of dense brown scales on intervals 5, not in sharp contrast to background squamae. Dorsum lacking other vittae or maculae. Mesosternum and metasternum covered with dense white to tan scales except transverse glabrous line at suture between sclerites, ventrites darker, with sparse, evenly distributed pale scales. Scales course (individually distinct under low magnification), appressed to suberect.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes ( Figure 104), interrupted at back by glabrous V-shaped area where head articulates on pronotum, thin, medial longitudinal carinae within crown from apex of V-shaped area to center of crown. Eyes flat, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.0–1.4 X broader than long, broadest at base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one tan to dirty white, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle slightly cavernous, lateral margins produced, posterior margin slightly open. Scutellum visible diameter subequal to punctures at base of elytra. Elytra 1.1–1.3 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before midline; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, not elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs moderately long, hind femora exceeding elytral apices by 1/3 their length; mesofemora exceeding elytral apices. Profemora unidentate or very rarely bidentate; apical profemoral tooth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire; when present, secondary profemoral tooth minute. Protibia strongly compressed laterally, ventral margin with flange distinct, tapering to apex, male flange terminating before tibial apex, female flange tapering to apex (as in Figures 47, 50); dorsal margin weakly convex. Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with distinct supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on significantly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia. Tergite VII broader than long, anterior margin convex, posterior margin strongly emarginate at middle, wing binding patch distinct, with two rows of 5–6 minute plectral tubercles reaching posterior margin, tubercles minute, distinct only under high magnification.

Female terminalia. Tergite VII longer than broad, anterior and posterior margins broadly convex, wing binding patch distinct, with two rows of 3–5 minute plectral tubercles; plectral rows not reaching posterior margin; tubercles visible only under high magnification. Bursa more or less indistinct from vagina; vagina with walls weakly sclerotized near junction with oviduct; no internal sclerites visible.

Material examined. Lectotype (here designated): ♂, " lugubris, Heller / K. Wilhelmland, 8577/ Coll. J. Faust, Ankauf 1900/ Syntypus " ( SMTD) . Paralectotypes (here designated): 2 ♂♂, "K. Wilhelmland , 8577/ lugubris, Heller / Coll. J. Faust, Ankauf 1900/ Syntypus " ( SMTD) ; 2 ♂♂, " K. Wilhelmland, 8577/ Typus" ( SMTD) ; 1 ♀, "K. Wilhelmland, 8577/ Syntypus " ( SMTD) ; 1 ♀, "K. Wilhelmland , 8577/ Syntypus / Asytesta lugubris, Det. K. M. Heller " ( DEI) 1 ♀, " K. Wilhelmland, 8577/ Cotypus/ Asytesta lugubris, Det. K. M. Heller " ( MSNG) . Non-type material: 1 ♂, " New Guinea: Huon, Peninsula , 1 mi. N., of Finschhafen , 18-11-1969, James E, Tobler, Cal.Acad.Sci.Coll." ( CASC) ; 1 ♀, " Finschhafen, New Guinea, 18-21-44, E. S. Ross " ( CASC) ; 1 ♂, " Fr. Wilhshafen , N. Guinea " ( MNHN) ; 3 ♂♂, 2 ♀♀," Papua New Guinea, Morobe Prov. Surim, Vill. 4-6 JAN.2003, Coll. G. P. Setliff / Finisterre Survey II, Hand collected, 1282m, Huon Penninsula, 146˚ 42' E, 05˚ 52' S" ( GPSC) ; 1 ♀, " Papua New Guinea, Morobe Prov. Surim, Vill. 4-6 JAN.2003, Coll. G. P. Setliff, 1282 m, Huon Penninsula, 146˚ 42' E, 05˚ 52' S" ( UMSP) ; 1 ♂, "D.N.G./ Coll. V. Bennigsen / Asytesta lugubris, Det. K. M. Heller " ( DEI) ; 2 ♂♂, "D.N.G./ Coll. V. Bennigsen " ( DEI) ; 1 ♂, " New Guinea / 543/ Coll. Pape / Asytesta lugubris " (DEI) ; 1 ♂, " New Guinea ( NE), Huon Peninsula, Finschhafen , 80-200m, 13.IV.1963 / J.&M.Sedlacek Collectors, Bishop" ( BPBM) ; 2 ♂♂, " New Guinea: NE, Zenag-Lae Road , 200m, 17.I.1965 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♂, " Sattelberg., Deutsch-Neu Guinea, H. Rolle, Berlin. W. Museum Paris, 1965, Coll. A. CLERC" ( MNHN) ; 1 ♂, "Sattelberg, N. Guinea, Museum Paris, 1949, Col. A. Hustache " ( MNHN) ; 1 ♂, "Lae N. Guinea, July 1944, F. E. Skinner/ Purdue University, Collection" ( BPBM) ; 1 ♀, "N. Guinea [illegible]/ Asytesta lugubris Hell. / Samml. K. F. Hartmann, Ankauf 1941.I" ( SMTD) ; 1 ♀, " Tree #32, Tray #5, Anisoptera thurifera, A. Allison & P. Swift, Colls., Bishop Museum / Papua New Guinea: Morobe Prov.: Oomsis , 530m, 11.IX.1988 / NO. 26569, PNG Canopy, Bishop Museum" ( BPBM) ; 1 ♀, "D.N. Guinea / Coll. Bennigsen " ( DEI) ; 1 ♀, "D.N.G./ Coll. Bennigsen / Asytesta lugubris, Det. K. M. Heller 1913 " ( DEI) ; 1 ♀, " Asytesta lugubris, Det. K. M. Heller " ( DEI) ; 1 ♀, "N. Guinea, Sattelberg , v. Bennigsen/ Asytesta lugubris, Det. K. M. Heller 1913 " ( DEI) ; 1 ♀, "D.N.G/ Coll. v. Bennigsen / Asytesta lugubris, Det. K. M. Heller 1912 " ( DEI) ; 1 ♀, " Sattelberg. , Wahnes. Franklin, Muller/ Heller det." ( DEI) ; 1 ♀, " New Guinea, Lae , 16 III: 1964/ J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♀, " Morobe , AGR. Station, Bubia, N. Guinea, 29-3-1955 / Ex. Coll. Aus., Pt. Moresby, N: 356./ On the DBR., cacao/ Coll. J.J.H., 3 Szent-Ivany" ( BMNH) ; 1 ♀, " Morobe , AGR. Station, Bubia, N. Guinea, 29-3-1955 / Ex. Coll. Aus., Pt. Moresby, N: 356./ Coll. J.J.H., 3 Szent-Ivany/ COM./INST. ENT., Coll. NO. 14276" ( BMNH) . Lectotype for A. setipes Lea (here designated): ♂, "Finschhaven, N. E. New Guinea / Asytesta setipes Lea, N. Guinea, type" ( SAM) .

Distribution. Papua New Guinea: Morobe Province, Huon Peninsula.

Plant associations. A female specimen was collected on Anisoptera thurifera Blume (Dipterocarpaceae) by A. Alison and P. Swift during canopy fogging studies in Papua New Guinea. Two specimens were also collected on cacao ( Theobroma cacao L. Malvaceae ).

Remarks. I select one of Heller’s syntypes as the lectotype for A. lugubris to ensure nomenclatural stability. Lea (1928: 77) reports the type locality for A. setipes as the junction of Wau Creek and the Bulolo River; however the type specimen is labeled as being from Finschhafen. I designate this specimen as the lectotype for A. setipes to fix the name to that specimen.

Asytesta marginalis Setliff , new species

( Figures 167–168, 193)

Diagnosis. This species may be confused with a few other similar species (viz., A. eudyasmoides , A. biakana new species, and A. humeralis ), which have more or less solid, dark (almost black) background coloration with short, white, rectangular humeral maculae. It differs from A. eudyasmoides by its bidentate profemora and shorter body form. Asytesta biakana and A. humeralis are more similar in shape to A. marginalis than the previous species; however, A. biakana lacks a visible scutellum, has unidentate profemora, and has small white maculae that are evenly distributed on the elytra. Asytesta humeralis is the most closely related species to A. marginalis based on the similarity in body shape and vestiture, the visible scutellum, bidentate profemora, and similar protibial shape. Asytesta marginalis differs from A. humeralis in its more densely and uniformly distributed, brown scales on the dorsum and the much more faint and reduced (usually absent) humeral maculae. Several other species of Asytesta also have rectangular humeral maculae (e.g., A. rata , A. signata , and A. thompsoni new species); however, these species all have the dorsal background vestiture strongly mottled with light and dark brown scales.

Description. Measurements (n = 16): body length 5.9–7.3 mm (mean 6.4 mm), body width 2.9–3.8 mm (mean 3.1 mm), pronotal length 2.4–3.0 mm (mean 2.6 mm), elytral length 3.4–4.3 mm (mean 3.9 mm), rostral length 1.7–2.5 mm (mean 2.1 mm). Body oval, 1.9–2.2 X longer than broad; integument dark reddish-brown to black, antennae and tarsi reddish-brown. Densely and evenly covered with small, tan to brown scales, with more densely distributed dirty white scales in elytral fascia. Frons, pleura, coxae, basal portion of femora, and venter more densely clothed in tan to brown scales; vestiture on femora and tibia interspersed with tan, hair-like setae; male with short, suberect, hair-like setae along ventral margin of protibia near apex, setae longer than elsewhere on legs; tarsi clothed with sparse, white, hair-like setae. Dorsum in dark color form occasionally with small, irregular, maculae of tan scales; maculae less densely distributed on sides. Pronotum occasionally with pair of sublateral vittae of denser tan scales from anterior margin to cervical constriction, sublateral vittae absent in some specimens; anterolateral region of pronotum with poorly defined ring of tan scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with darker background scales in center (as in Figure 104); dorsal portion of ring not extending beyond middle of procoxa. Elytra with thin distinct fascia spanning entire anterior margin, margin between intervals 5 never bare, reaching epipleural margin; humeri rarely with faint, rectangular, maculae of tan scales on intervals 5, usually absent; lacking other vittae. Mesosternum and metasternum covered with dense tan scales except transverse glabrous line at suture between sclerites, ventrites with sparse, evenly distributed tan scales. Scales fine (indistinct under low magnification), appressed to suberect.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes ( Figure 104), interrupted at back by glabrous V-shaped area where head articulates on pronotum, medial longitudinal carinae absent. Eyes flat, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, sides parallel in basal 2/3, lightly broadened in apical 1/3, with interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.1–1.3 X broader than long, broadest subbasally; base nearly truncate, medial area only weakly produced. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one tan, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle slightly cavernous, lateral margins produced, posterior margin slightly open. Scutellum visible, minute, diameter subequal to punctures at base of elytra. Elytra 1.1–1.3 X longer than broad, anterior margins truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before middle of elytra; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, not elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs moderately long, hind femora exceeding elytral apices by 1/3 their length; mesofemora exceeding elytral apices. Profemora bidentate; apical profemoral teeth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange distinct, tapering to apex, male flange terminating before middle of tibia, female flange tapering to apex (as in Figures 45–46); dorsal margin weakly convex. Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on significantly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia. Tergite VII broader than long, anterior margin convex, posterior margin emarginate at middle, wing binding patch distinct, with two rows of 5–8 minute plectral tubercles reaching posterior margin, tubercles greatly reduced, distinct under high magnification.

Female terminalia. Tergite VII longer than broad, anterior and posterior margins broadly convex, wing binding patch distinct, with two rows of 5–8 minute plectral tubercles; plectral rows not reaching posterior margin; tubercles visible, minute, distinct under high magnification. Bursa more or less indistinct from vagina; vagina with walls weakly to moderately sclerotized; no distinct internal sclerites visible.

Material examined. Holotype: ♂ "Marikurubu, 700–1500 m, 29.X.1999 / N. Moluccas, Ternate, leg. A. Riedel " ( MZB) . Paratypes: 1 ♂, 1 ♀, " Marikurubu , 700–1500 m, 29.X.1999 / N. Moluccas, Ternate, leg. A. Riedel " ( MZB) ; 1 ♂, 1 ♀," Marikurubu , 700–1500 m, 29.X.1999 / N. Moluccas, Ternate, leg. A. Riedel " ( SMNK) ; 7 ♂♂, 6 ♀♀, " Marikurubu , 700–1500 m, 29.X.1999 / N. Moluccas, Ternate, leg. A. Riedel " ( ARC) ; 1 ♀," Marikurubu , 700–1500 m, 29.X.1999 / N. Moluccas, Ternate, leg. A. Riedel " ( GPSC) ; 1 ♀," Marikurubu , 700–1500 m, 29.X.1999 / N. Moluccas, Ternate, leg. A. Riedel " ( BMNH) . 1 ♂, " Ternate, Aequi Conora , Beccari 1874. XI/ Asytesta humeralis Psc. , det. Pascoe 1885 " ( MSNG) ; 1 ♂, " Moluche, Ternate, coll. Bruyn 1875/ Asytesta humeralis Psc. , det. Pascoe 1885 " ( MSNG) ; 1 ♀, " Ternate, Aequi Conora , Beccari 1874. XI/ Asytesta humeralis Psc. , det. Pascoe 1885 " ( MSNG) ; 15 ♂♂, 10 ♀♀, " Ternate, coll Bruijn 1877" ( MNHN) ; 1 ♀, " Ternate " ( MNHN) ;

Distribution. Indonesia: Ternate Island.

Etymology. The specific epithet refers to the distinct white fascia that spans the anterior margin of the elytra in this species.

Remarks. Figures 167–168 show the more typical dark color form of this species.

Asytesta morobeana Setliff , new species

( Figures 169–170, 193)

Diagnosis. The solid brown to olive dorsal vestiture that is not marked with distinctly contrasting vittae or maculae and whitish scales on the vertex of the head will separate this species from other Asytesta species except A. allisoni new species and A. lugubris . The vestiture in A. lugubris is typically darker and it has a faint brown dorsolongitudinal vitta that reaches the base and indistinct, rectangular, maculae of dense brown scales on intervals 5 of the elytra. Asytesta allisoni has the anterolateral region of the pronotum in front of the procoxae bearing solid cream colored patch of scales. It also has the small pale maculae on the pronotal disk more distinct and a large and a black medial spot near the base of the elytra.

Description. Measurements (n = 20): body length 4.4–6.9 mm (mean 5.7 mm), body width 2.2–3.6 mm (mean 2.9 mm), pronotal length 2.0– 2.9 mm (mean 2.4 mm), elytral length 2.5–4.0 mm (mean 3.3 mm), rostral length 1.5–2.0 mm (mean 1.8 mm). Body oval, 1.7–2.2 X longer than broad; integument dark reddish-brown, antennae and tarsi reddish-brown. Densely covered with creamy white to light brown to fuscous scales. Frons, pleura, coxae, and basal portions of femora more densely clothed in yellowish-white to light brown scales; vertex of head with white scales; pair of small yellowish-white maculae on temples (as in Figure 104). Legs with alternating bands of light and dark brown scales, background vestiture interspersed with white, hair-like setae; male with short, suberect, hair-like setae along ventral margin of protibia near apex, setae longer than elsewhere on legs; tarsi clothed with sparse, white, hair-like setae. Dorsum not mottled or dappled with poorly defined, irregular yellowishwhite patches; pronotal disk with faint, dorsolongitudinal vitta on cervical constriction from apical margin to about midline; vitta same color as background vestiture; pair of small dark patches at posterior margin on either side of midline, 4–6 irregular dark, maculae in transverse band slightly in front of midline, reaching lateral margins in most specimens, maculae sometimes fused into transverse band, often very faint (best viewed without magnification), 8–12 small, circular, whitish maculae arranged in three transverse rows of 2–4 maculae each, one row of 2–4 maculae before middle, one pair at middle, one row of 2–4 macular after middle; pair of similarly formed but smaller maculae at apical pronotal margin, sometimes absent, vitta and maculae not in sharp contrast to background squamae; anterolateral region of pronotum with poorly defined ring of denser yellowish-white scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with darker background scales in center (as in Figure 104); dorsal portion of ring not extending beyond middle of procoxa. Dorsum lacking other vittae or maculae. Mesosternum and metasternum covered with dense white to tan scales except transverse glabrous line at suture between sclerites, ventrites darker, with sparse, evenly distributed pale scales. Scales course (individually distinct under low magnification), appressed to suberect.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes ( Figure 104), interrupted at back by glabrous V-shaped area where head articulates on pronotum, thin, medial longitudinal carinae within crown from apex of V-shaped area to center of crown. Eyes flat, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate to slightly convex. Antennal club oval.

Thorax. Pronotum 1.0–1.5 X broader than long, broadest at base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one tan to dirty white, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle slightly cavernous, lateral margins produced, posterior margin slightly open. Scutellum visible diameter subequal to punctures at base of elytra. Elytra 1.1–1.3 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before midline; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, not elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs moderately long, hind femora exceeding elytral apices by 1/3 their length; mesofemora exceeding elytral apices. Profemora unidentate; profemoral tooth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange distinct, tapering to apex, male flange terminating before tibial apex, female flange tapering to apex (as in Figures 47, 50); dorsal margin weakly convex. Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with distinct supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on significantly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia. Paired apodeme-like basal sclerites of endophallus with narrow, curved, projection; originating laterally from about middle of dorsal surface, curved toward base of endophallus. Tergite VII broader than long, anterior margin convex, posterior margin strongly emarginate at middle, wing binding patch distinct, with two rows of 5–8 minute plectral tubercles reaching posterior margin, tubercles minute, distinct only under high magnification.

Female terminalia. Tergite VII longer than broad, anterior and posterior margins broadly convex, wing binding patch distinct, with two rows of 5–6 minute plectral tubercles; plectral rows not reaching posterior margin; tubercles visible only under high magnification. Apodeme of sternite VIII narrow, apex Y-shaped. Bursa more or less indistinct from vagina; vagina with walls moderately sclerotized near junction with oviduct; no internal sclerites visible.

Material examined. Holotype: ♂, " New Guinea : NE, Wau, 21.IV.1962 / J. Sedlacek Collector Bishop " ( BPBM) . Paratypes: 1 ♂, 4 ♀♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 5–6.VII.'61/ J.H. Sedlacek Collector Bishop" ( BPBM) ; 1 ♀, "New Guinea: ( NE), 6km W of Wau, Nami , Creek, 1700 m., 12.VI.1962 / J. Sedlacek Collector Bishop" ( BPBM) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 15–30.IX.1962, J. Sedlacek " ( MNHN) ; 1 ♂, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 15–22.XI.'61/ J. Sedlacek Collector Bishop" ( MSNG) ; 1 ♂, "New Guinea: ( NE), Wau , Morobe Distr., 1200–1300m, 25.II.1962 / J. Sedlacek Collector Bishop " ( BPBM) ; 1 ♂, " New Guinea : NE, Wau, 1150–1600m., 9.II.1968 / J. Sedlacek Collector Bishop" ( SMTD) ; 1 ♀, " New Guinea : NE, Wau, 1700m, 12.III.1969 / J. Sedlacek Collector Bishop" ( BPBM) ; 1 ♂, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 5.VIII.1961 / J. Sedlacek Collector Bishop " ( BPBM) ; 1 ♂, "New Guinea: ( NE), Wau , Morobe Distr., 1200–1300m., 5.VI.1963 / J. Sedlacek Collector Bishop " ( BPBM) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1180–1300m., 11.IX.1964 / J. Sedlacek Collector Bishop " ( BPBM) ; 1 ♂, 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m. 1–15.V.1962 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m., 6–7.XI.1961 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♂, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m. 1–15.V.1966 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m., 19.X.1961 / J. Sedlacek Collector, Bishop" ( GPSC) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 27.X.1961 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♂, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 19.XI.1961 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♂, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 15–22.XI.'61/ J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 15–24.XI.61/ J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♀, " New Guinea : NE, Wau, Hospital Ck., 1200 m, 12.I.1965 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♂, 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 17.IX.1961 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♂, 3 ♀♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 15–22.XI.'61/ J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♀, " New Guinea : NE, Wau, 1200–1300m, 4.IX.1965 / J. Sedlacek Collector, Bishop Mus." ( BPBM) ; 1 ♀, " New Guinea : NE: Wau; 1220m, 9.11.1964, W. A. Steffan. coll., Bishop Museum" ( UMSP) ; 1 ♀, " New Guinea : NE: Wau; 1220m, 9.11.1964, W. A. Steffan. coll., Bishop Museum" ( USNM) ; 1 ♂, 1 ♀, " New Guinea : NE, Wau, 1200–1450m, 18.VI.1968 / N. L. H. Krauss, Collector" ( USNM) ; 1 ♀, " New Guinea : NE, Wau, 1200–1450m, 18.VI.1968 / N. L. H. Krauss, Collector" ( BPBM) ; 1 ♂, "New Guinea: ( NE), Wau , Morobe Distr., 1300 m, 12.VII.1961 / J. Sedlacek Collector, Bishop Mus." ( BPBM) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200–1300m., 14.III.1963 / J. Sedlacek Collector, Bishop Mus." ( BPBM) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 23.VII.1961 / J. Sedlacek Collector, Bishop Mus." ( BPBM) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 15– 30.9.1962, J. Sedlacek " ( BPBM) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 15.VII.1961, J. Sedlacek / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 30.VII.1961 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♂, "New Guinea: ( NE), Wau , Morobe Distr., 1300 m, 22.VII.1961 / J. Sedlacek Collector, Bishop/ J. Sedlacek Collector, Bishop" ( BPBM) ; 2 ♀♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 25.VII.1961 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1300 m, 12.VII.1961 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 10.IX.1961 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♂, 2 ♀♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 15.VII.1961, J. Sedlacek / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1700–1800m., 17.IX.1961 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♀, " New Guinea , Wau 1200 m, 28.XII.1965 / Malaise Trap, J. & M. Sedlacek, Bishop" ( BPBM) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 7.IX.1961 / J. Sedlacek Collector, Bishop" ( BPBM) ; 2 ♀♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 11.IX.1961 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 6–7.XI.1961 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♂, "New Guinea: ( NE), Wau , Morobe Distr., 1270 m, 7.V.1962 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♀, " New Guinea : NE Wau 1200 m, 18.IV.1965 / J. Sedlacek Collector, Bishop Museum" ( BPBM) ; 1 ♂, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 17.IX.1961 / J. Sedlacek Collector, Bishop " ( BPBM) ; 1 ♂, 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 2.VI.1962 / J. Sedlacek Collector, Bishop " ( BPBM) ; 1 ♂, "New Guinea: ( NE), Wau , Morobe Distr., 1200–1300m., 31.III.1963 / J. Sedlacek Collector, Bishop " ( BPBM) ; 1 ♂, " New Guinea : NE, Bulolo R. 680 m, 4.VI.1969 / J. Sedlacek Collector, Bishop Mus." ( BPBM) ; 1 ♂, "New Guinea: ( NE), Wau , Morobe Distr., 1200–1250m., 15–16.VIII.1964 / J. Sedlacek Collector, Bishop " ( BPBM) ; 2 ♀♀, " New Guinea : NE, Wau, Nami Ck., 1700–1850m, 7.II.1966 / J. Sedlacek Collector, Bishop " ( BPBM) ; 1 ♂, " New Guinea : NE, Wau, Hospital Ck., 1200 m, I.1965 / J. Sedlacek Collector, Bishop " ( BPBM) ; 1 ♂, " New Guinea : NE, Wau, 1200 m, 23.II.1966 / J. Sedlacek Collector, Bishop " ( BPBM) ; 1 ♂, " New GUINEA: NE, Wau, 1200 m, 18.4.1965 / J. Sedlacek Collector, Bishop " ( BPBM) ; 1 ♀, " Wau , 14.3.250m./ J. Sedlacek Collector, Bishop " ( BPBM) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1050–1100m., 15.VII.1961 / J., J. & M. Sedlacek, G.Monteith & Native Collectors, Bishop Museum" ( SMTD) ; 1 ♀, "New Guinea: ( NE), Wau , Morobe Distr., 1200 m, 5–6.VII.'61/ J. Sedlacek Collector, Bishop " ( BPBM) ; 1 ♀, " New Guinea : NE, Wau, Nami Ck., 1700–1850m, II.1966 / J. Sedlacek Collector, Bishop Mus." ( BPBM) ; 1 ♂, " New Guinea : NE, Wau, 1700–1800m, 27.IX.1965 / J. Sedlacek Collector, Bishop Mus." ( GPSC) ; 1 ♀, " New Guinea : NE, Wau, 1200–1250 m., 15–16.VIII.1964 / J. Sedlacek, Bishop" ( BPBM) ; 1 ♀, 1 ♂, " New Guinea : NE, Wau, 1790m, 5.II.1966 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♂, "New Guinea: NE, Mt. Missim , 1600–2000m., 21–24.IX.'64/ J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♂, " New Guinea : NE, Mt.Kaindi, 2200–2350m, 25.III.1965 / J. Sedlacek Collector, Bishop" ( BPBM) ; 1 ♂, "Papua N.G., Bulolo , 14.IX.1981 / Prunus sp. billet, H. Roberts coll., C.I.E. A13485 / 1564" ( BMNH) ; 1 ♂, " New Guinea : NE, Wau, 1790m, 5.II.1966 / J. Sedlacek Collector, Bishop" ( BPBM) . 1 ♂, " New Guinea : Morobe , District, Wau, 2–3.X.1969, James E. Tobler, Cal. Acad. Sci. Coll." ( CASC) ; 1 ♂, " Papua New Guinea : Morobe, Wau Ecology Institute, 1200 m, 5.II.2000, 7˚ 20'24'' S 146˚ 42'17'' E, R.S. Anderson, misc. collns, RSA 2000-034X " ( CMNC) .

Distribution. Papua New Guinea: Morobe Province, Wau area.

Plant associations. This species has been collected on Prunus (Rosaceae) and Theobroma cacao L. ( Malvaceae ).

Etymology. This species is named for the Morobe Province of Papua New Guinea where the species occurs.

Asytesta rata Heller

( Figures 47, 50, 67, 104, 173–174, 194)

Asytesta rata Heller, 1910: 29 . Lectotype, here designated, in SMTD. Labeled “Kais Wilhelmsland, Paup, Dr. Schlaginhaufen / 1900/ type. Type locality: Papua New Guinea: Bongu.

Asytesta circulifera Lea, 1928: 75 , new synonymy. Lectotype, here designated, in SAM. Labeled “ Asytesta circulifera , N. Guinea/ TY [handwritten on card beneath specimen].” Type locality: Papua New Guinea: Manumbo.

Diagnosis. This species has a strongly mottled pattern of dark scales against a yellowish-white background. It also has a somewhat variable arrangement of small white maculae on the elytral humeri. It is closely allied with A. thompsoni new species, from which it can be differentiated by its lighter scales on the vertex of the head, base of pronotal disk, and legs and the more mottled appearance from the light yellowish-white to tan colored scales nearly covering its dorsum. Also, the dorsomedian longitudinal vitta on the pronotum is more distinct, usually exceeding the midline or reaching the posterior margin.

Redescription. Measurements (n = 10): body length 4.8–6.9 mm (mean 5.6 mm), body width 2.5–3.5 mm (mean 3.0 mm), pronotal length 2.0–3.0 mm (mean 2.5 mm), elytral length 2.8–3.9 mm (mean 3.2 mm), rostral length 1.4–2.0 mm (mean 1.9 mm). Body oval, 1.8–2.1 X longer than broad; integument dark reddish-brown to black, antennae and tarsi reddish-brown. Densely covered with yellowish-white to fuscous scales. Frons, pleura, and coxae similarly clothed as dorsum; pair of small white maculae on temples ( Figure 104). Femora with alternating bands of yellowish-white, tan, and brown scales, tibia more or less uniformly light brown; legs with background vestiture interspersed with white, hair-like setae; male with short, suberect, hair-like setae along ventral margin of protibia near apex, setae longer than elsewhere on legs; tarsi clothed with sparse, white, hair-like setae. Dorsum mottled with poorly to well defined, irregular dark patches; pronotal disk with faint to well defined, dorsolongitudinal vitta of dense white scales from apical margin to base, pair of small dark patches at posterior margin on either side of midline, 4–6 irregular dark, maculae in transverse band slightly in front of midline, reaching lateral margins in most specimens, maculae sometimes fused into transverse band (best viewed without magnification), 6–8 small, circular, whitish maculae arranged in two transverse rows of 2–4 maculae each, one row of 2–4 maculae before middle, one row of 2–4 maculae after middle; pair of similarly formed but smaller maculae at apical pronotal margin, sometimes absent, vitta and maculae in sharp contrast to background squamae, anterolateral region of pronotum with well defined ring of denser dirty white scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with darker background scales in center ( Figure 104); dorsal portion of ring not extending beyond middle of procoxa. Elytra usually with thin distinct fascia of yellowishwhite scales spanning entire anterior margin, fascia sometimes absent (as in Figure 174), usually reaching epipleural margin; humeri with distinct, rectangular, maculae of dense white scales on intervals 5 and 6; macula on intervals 5 longer than macula on intervals 6; maculae in sharp contrast to background squamae; maculae often reduced to thin rectangular maculae on fifth intervals only. Dorsum lacking other vittae or maculae. Mesosternum and metasternum covered with dense yellowish-white scales except transverse glabrous line at suture between sclerites, ventrites darker, with sparse, evenly distributed pale scales ( Figure 104). Scales course (individually distinct under low magnification), appressed to suberect.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes ( Figure 104), interrupted at back by glabrous V-shaped area where head articulates on pronotum, thin, medial longitudinal carinae within crown from apex of V-shaped area to center of crown. Eyes flat, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.1–1.3 X broader than long, broadest at base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one white to brown, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle slightly cavernous, lateral margins produced, posterior margin slightly open. Scutellum visible, minute, diameter smaller than punctures at base of elytra. Elytra 1.0–1.2 X longer than broad, anterior margin truncate, declivity gradually declining to apex ( Figure 67); punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated near base; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, not elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs moderately long, hind femora exceeding elytral apices by 1/3 their length; mesofemora exceeding elytral apices. Profemora unidentate; profemoral tooth slightly larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange distinct, tapering to apex, male flange terminating before tibial apex, female flange tapering to apex ( Figures 47, 50); dorsal margin weakly convex. Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with distinct supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on significantly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male and female terminalia. As in generic description.

Material examined. Lectotype (here designated): 1 ♂, "Kais Wilhelmsland, Paup, Dr. Schlaginhaufen / 1900/ type" ( SMTD) . Paralectotypes (here designated): 1 ♂, "N. Guinea, [illegible]/ Coll. J. Faust, Ankauf 1900/ Syntypus " ( SMTD) ; 1 ♀, "N. Guinea, [illegible]/ Coll. J. Faust, Ankauf 1900/ Syntypus / Asytesta Heller , det." ( SMTD) ; 1 ♀, " Neu Guinea 76593/ Syntypus " ( SMTD) . Non-type material: 1 ♂, "27400/ Sorovi, Northern Dist. , Papua: 25.9.73, B. Keoro " ( BMNH) ; 1 ♂, "Bongu, Moluc./ [illegible]/ Asytesta Heller / Samml. K. F. Hartmann, Ankauf 1941.1" ( SMTD) ; 1 ♂, "D. Neu-Guinea, Wahnes Franklin-Muller/ C. F. Baker, collection, 1927" ( USNM) ; 1 ♂, "D. Neu-Guinea, Wahnes Franklin-Muller/ C. F. Baker, collection, 1927/ Asytesta Heller / Asytesta rata, DEM Heller " ( USNM) ; 1 ♂, "D.N.G/ Coll. Bennigsen / Asytesta rata 10., Det. K. M. Heller " ( DEI) ; 3 ♂♂, "D. Neu- Guinea, Wahnes Franklin-Muller/ Heller det." ( DEI) ; 1 ♂, "D.N. Guinea, Rattalberg/ Coll. V. Bennigsen " ( DEI) ; 2 ♂♂, "D.N.G/ Coll. V. Bennigsen " ( DEI) ; 1 ♂, "D.N.G/ Coll. V. Bennigsen / Asytesta rata 10., Det. K. M. Heller " ( DEI) ; 1 ♂, " Papua New Guinea, Madang Prov. Ohu Vill. , 200m 11-Oct-2002, Coll. G. P. Setliff / 145˚ 41' E 5˚ 14' S" ( GPSC) ; 1 ♂, " Papua New Guinea, Madang Prov. Ohu Vill. , 200m 07-NOV-2002, Coll. J. Hulcr / on dead, Myristica sp. , 145˚ 41' E 5˚ 14' S" ( GPSC) ; 1 ♂, " Papua New Guinea, Madang Prov. Ohu Vill. , 200m 30-Jan-2001, Coll. M.Janda / hand collected, 145˚ 41' E 5˚ 14' S" ( GPSC) ; 3 ♂♂, 6 ♀♀ " Papua New Guinea, Madang Prov. Ohu Vill. , exposed 29-Jan- 2002, 200m. 145˚ 41' E 5˚ 14' S/ L.Cizek, B.Isua, & J.Auga coll., reared from wood of Ficus wassa " (GPSC) ; 1 ♂, " Papua New Guinea, Madang Prov. Ohu Vill. , exposed 10 Jan 2003, 200m. 145˚ 41' E 5˚ 14' S/ L.Cizek, B.Isua, & J.Auga coll., reared from wood of Ficus wassa WAS " ( GPSC) ; 3 ♂♂, 1 ♀, " Papua New Guinea, Madang Prov. Ohu Vill. , exposed 20 March 2003, 200m. 145˚ 41' E 5˚ 14' S/ L.Cizek, B.Isua, & J.Auga coll., reared from wood of Syzigium SY 2" ( GPSC) ; 1 ♀, " Papua New Guinea, Madang Prov. Ohu Vill. 12 vi. 2003. G.P. Setliff, on dead Ficus nodosa " (GPSC) ; 1 ♀, " Papua New Guinea, Madang Prov. Ohu Vill. , 200m. 09 May 2003, Coll. G. P. Setliff / hand collected, prim & sec. forest, 145˚ 41' E 5˚ 14' S" ( GPSC) ; 7 ♀♀, " Papua New Guinea, Madang Prov. Ohu Vill., exposed. 29-JAN- 2002, 200m. 145˚ 41' E 5˚ 14' S/ G.P.Setliff, M.Mogia & M.Andress coll., reared from branches of Mystica sp. MYR" ( GPSC) ; 1 ♀, " Papua New Guinea, Madang Prov. Ohu Vill. , exposed 07 May 2003 200m. 145˚ 41' E 5˚ 14' S/ L.Cizek, B.Isua, & J.Auga coll., reared from wood of Ficus wassa WAS " ( GPSC) ; 2 ♀♀, " Papua New Guinea, Madang Prov. Ohu Vill. , 200m. 09 May 2003, Coll. G. P. Setliff, 145˚ 41' E 5˚ 14' S" ( GPSC) ; 1 ♀, " Papua New Guinea, Madang Prov. Ohu Vill., exposed 17 Feb. 2003, 200m. 145˚ 41' E 5˚ 14' S/ G. P. Setliff, M. Mogia & M. Andress coll., reared from branches of Mystica sp. ( MYR)" ( MSNG) ; 1 ♂, " Papua New Guinea, Madang Prov. Ohu Vill. , exposed 21 April 2002 200m. 145˚ 41' E 5˚ 14' S/ L.Cizek, B.Isua, & J.Auga coll., reared from wood of Sterculia schumanniana ” (GPSC) ; 1 ♂, 1 ♀, " Papua New Guinea, Madang Prov. Ohu Vill. , exposed 29-Jan- 2002, 200m. 145˚ 41' E 5˚ 14' S/ L.Cizek, B.Isua, & J.Auga coll., reared from wood of Ficus wassa " (UMSP) ; 4 ♂♂, " Papua New Guinea: Madang, Baitabag , Madang (8km N.W) 100m, 31.1.2000, 5 08'19"S 145˚ 46'34"E, R.S.Anderson rainforest, RSA2000-028X" ( CMNC) ; 1 ♂, "Gauli m/ J. & M. Sedlacek Collectors, Bishop" ( BPBM) ; 1 ♀, "New Guinea: NE, Madang District, Wanuma 600–720m, VIII.1968 / N.L.H. Krauss, Collector, Bishop Museum" ( BPBM); 4 ♀♀, " Coll. R. I. Sc. N. B., Papua New Guinea, Madang Prov., Baiteta 11-V-1994, leg. Olivier Missa " ( ISNB); 1 ♀, "FOG XL-A2, CCL-3824" ( ISNB); 1 ♀, "FOG M8-7, SP40A, CCL-8582" ( ISNB); 1 ♀, "FOG M8-6, CCL-3622" ( ISNB) ; 2 ♀♀, "D.N.G/ Coll. V. Bennigsen " ( DEI) ; 1 ♀, "D.N.G/ Asytesta rata , det. K. M. Heller 1912" ( DEI) ; 1 ♀, "D. Neu-Guinea, Wahnes Franklin-Muller/ Asytesta rata , det. K. M. Heller 1912" ( DEI) ; 4 ♀♀, "D.N.G/ Coll. v. Bennigsen " ( DEI) ; 1 ♀, "D. Neu-Guinea, Wahnes Franklin-Muller/ Heller det." ( DEI) ; 1 ♀, "D.N.G/ Coll. n. Bennigsen/ Asytesta rata , det. K. M. Heller 1912" ( DEI) ; 1 ♀, "Sattelberg, D.N. Guinea / Coll. v. Bennigsen / Asytesta rata det. K. M. Heller 1913" ( DEI) ; 1 ♀, " Coll. V. Bennigsen " ( DEI) ; 1 ♀, "D.N.G/ Coll. v. Bennigsen / Asytesta rata m.'10, det. K. M. Heller 1912" ( DEI) ; 1 ♀, "New Guinea: NE, Finistere Range, Saidor: Gabumi, Vill. VI-24-30-'58/ W. W. Brandt, Collector, Bishop" ( BPBM) ; 1 ♀, " New Guinea ( NE), Huon Peninsula, Finschhafen , 200m, 10.IV.1963 / W. W. Brandt, Collector, Bishop" ( BPBM) ; 1 ♂, 1 ♀, " Papua New Guinea, Madang, Province, Sapi Forest Reserve, Sapi River at confluence with Gogol River , 50 m, 15 March 1989 Stop# 89-26A/ D.H. Kavanaugh, G.E. Ball, N.D. Penny, & P.A. Meyer collectors, Cal. Acad. Sci. Coll./ Papua New Guinea Expedition, 1989" ( CASC) ; 1 ♀, " Papua New Guinea, Madang, Province, Baiteta Road , 2.5 km W of North Coast, Road, 60 m, 2 March 1989, Stop#89-7/ D.H. Kavanaugh, G.E. Ball, N.D. Penny, & P.A. Meyer collectors, Cal. Acad. Sci. Coll./ Papua New Guinea Expedition, 1989" ( CASC); 3 ♂♂, " Coll. R. I. Sc. N. B., Papua New Guinea, Madang Prov., Baiteta 23-VI-1994, leg. Olivier Missa " ( ISNB); 2 ♂♂, "FOG AR 57-6, CCL-A5264" ( ISNB) . Lectotype for A. circulifera Lea (here designated): 1 ♂, " Asytesta circulifera , N. Guinea / TY [handwritten on card beneath specimen]" ( SAM) . Paralectotypes for A. circulifera : 2 ♀♀, [mounted on either side of Lectotype on same card] ( SAM) .

Distribution. Papua New Guinea: Madang Province: Bongu [type locality], Madang area; Morobe Province. I examined two specimens of this species in the SMTD collection that were collected on New Britain. Additional material from the Bismarck Archipelago is needed to confirm this locality.

Plant associations. See Table 1 for list of putative adult and larval host plants.

Remarks. I have selected the specimen labeled “type” from Heller’s collection to be the lectotype for the A. rata to fix the name to that single specimen. The remaining types were labeled as syntypes and become paralectotypes. I also designate a lectotype for Lea’s A. circulifera as identified in the material examined section above to ensure the stability of nomenclature.

Asytesta signata Faust

( Figures 177–178, 194)

Asytesta signata Faust, 1898: 165 [key], 171 [description]. Lectotype, here designated, in SMTD. Labeled “Kiriwini, Trobriand Is., III. IV. X. 95, (A.S. Meek)/ signata, Faust / Coll. J. Faust, Ankauf 1900/ Type.” Type localites: Fergusson Islands; Trobriand Islands.

Diagnosis. This is one of the smallest species (4.1–4.8 mm) in the genus aside from the two species in the A. doriae species group. The vestiture is more or less uniformly brown to fuscous in color and weakly mottled with poorly defined, irregular patches of dark brown scales. There is a pair of larger dark patches at posterior margin of pronotal disk on either side of midline, 4 irregular dark maculae in transverse band slightly in front of midline, and 10–14 small, circular, whitish maculae arranged in three transverse rows of 2–4 maculae each. The elytral humeri bear distinct, rectangular, maculae of dense white scales on intervals 5 and 6. The scutellum is extremely minute but visible (under high magnification) in all specimens examined. It is most similar to A. brevipennis , which is somewhat larger, has fewer maculae on the pronotal disk (6–8) and entirely lacks humeral maculae.

Redescription. Measurements (n = 5): body length 4.1–4.8 mm (mean 4.5 mm), body width 2.0– 2.5 mm (mean 2.3 mm), pronotal length 1.8–2.0 mm (mean 2.0 mm), elytral length 2.3–2.8 mm (mean 2.5 mm), rostral length 1.3–1.6 mm (mean 1.4 mm). Body oval, 1.8–2.1 X longer than broad; integument dark reddish-brown, antennae and tarsi reddish-brown. Densely covered with brown to fuscous scales. Frons, pleura, coxae, basal portions of femora, and venter more densely clothed in dirty white to light brown scales; pair of small yellowishwhite maculae on temples (as in Figure 104). Legs with background vestiture interspersed with white, hair-like setae; male with short, suberect, hair-like setae along ventral margin of protibia near apex, setae longer than elsewhere on legs; tarsi clothed with sparse, white, hair-like setae. Dorsum mottled with faint, dark brown areas; pronotal disk with pair of faint, dark patches at posterior margin on either side of midline, 4 irregular dark maculae in transverse band slightly in front of midline not reaching sides in most specimens (best viewed without magnification), 10–14 small, circular, whitish maculae arranged in three transverse rows of 2–4 maculae each, one row of 2–4 maculae before middle, one pair at middle, one row of 2–4 macular after middle; 2–4 similarly formed but smaller maculae at apical pronotal margin, sometimes absent; maculae in sharp contrast to background squamae; anterolateral region of pronotum with poorly defined ring of denser dirty white to tan scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with darker background scales in center, absent in some specimens. Elytral humeri with distinct, rectangular, maculae of dense white scales on intervals 5 and 6; macula on intervals 5 longer than macula on intervals 6. Dorsum lacking other vittae or maculae. Mesosternum and metasternum covered with sparse brown scales except transverse glabrous line at suture between sclerites, ventrites with sparse, evenly distributed paler scales. Scales course (individually distinct under low magnification), appressed to suberect.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes ( Figure 104), interrupted at back by glabrous V-shaped area where head articulates on pronotum, thin, lacking medial longitudinal carinae within crown from apex of V-shaped area to center of crown. Eyes flat, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.1–1.3 X broader than long, broadest at base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one tan to dirty white, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle slightly cavernous, lateral margins produced, posterior margin slightly open. Scutellum visible, very minute, diameter smaller than punctures at base of elytra. Elytra 1.0–1.2 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before midline; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, not elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs moderately long, hind femora exceeding elytral apices by 1/3 their length; mesofemora exceeding elytral apices. Profemora unidentate; profemoral tooth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange distinct, tapering to apex, male flange terminating before tibial apex, female flange tapering to apex (as in Figures 47, 50); dorsal margin weakly convex. Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with distinct supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on significantly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia. Tergite VII broader than long, anterior margin convex, posterior margin emarginate at middle, wing binding patch distinct, with two rows of 6–8 very minute plectral tubercles reaching posterior margin, tubercles greatly reduced, distinct only under high magnification.

Female terminalia. Tergite VII longer than broad, anterior and posterior margins broadly convex, wing binding patch distinct, with two rows of 8 very minute plectral tubercles; plectral rows not reaching posterior margin; tubercles greatly reduced, distinct only under high magnification. Bursa more or less indistinct from vagina; vagina with walls weakly sclerotized; no distinct internal sclerites visible.

Material examined. Lectotype (here designated): ♂, "Kiriwini, Trobriand Is., III. IV. X. 95, (A.S. Meek)/ signata, Faust / Coll. J. Faust, Ankauf 1900/ Type" ( SMTD) . Paralectotypes (here designated): 1 ♀, "Kiriwini, Trobriand Is., III. IV. X. 95, (A.S. Meek)/ signata, Faust / Coll. J. Faust, Ankauf 1900/ Type" ( SMTD) ; 1 ♂, "Fergusson I., X.X.XI.XI.96, (A.S. Meek)/ signata, Faust / Coll. J. Faust, Ankauf 1900/ Type" ( SMTD) . Non-type material: 1 ♂, "New Guinea SE, Popondetta, Buka-Bara, 23.IX.1963 / P.S." ( BPBM) ; 1 ♂, "Kiriwini, Trobriand Is., III. IV. X. 95, (A.S. Meek)" ( MNHN) ; 1 ♀, "Trobriand Is./ Samml. K. F. Hartmann, Ankauf 1941.1/ Asytesta humeralis?, Pasc. " ( SMTD) ; 4 ♀♀, "Kiriwini, Trobriand Is., A.S. Meek 1984" ( MNHN) .

Distribution. Papua New Guinea: D’Entrecasteaux Islands: Fergusson Islands; Trobriand Islands.

Remarks. A lectotype is designated from the type series to ensure stability of nomenclature and fix the name to a single specimen. I selected the specimen in the best condition from those specimens that were labeled with Faust’s “type” label.

Asytesta thompsoni Setliff , new species

( Figures 179–180, 194)

Diagnosis. This species is very close in appearance to A. rata , from which it can be differentiated by the darker scales on the vertex of the head, base of pronotal disk, and legs and the less mottled appearance of its dorsum. Legs are not banded in this species and the transverse band of small white maculae on the pronotal disk behind the midline is absent or indistinct. Also, the dorsomedian longitudinal vitta on the pronotum is fainter behind the midline and is most often restricted to the area before the cervical constriction.

Description. Measurements (n = 16): body length 5.4–7.7 mm (mean 6.0 mm), body width 2.8–4.0 mm (mean 3.3 mm), pronotal length 2.2–3.3 mm (mean 2.5 mm), elytral length 3.2–4.4 mm (mean 3.7 mm), rostral length 1.8–2.5 mm (mean 2.1 mm). Body oval, 1.9–2.1 X longer than broad; integument dark reddish-brown to black, antennae and tarsi reddish-brown. Densely covered with brown to fuscous scales. Frons, pleura, and coxae similarly clothed as dorsum; pair of small yellowish-white maculae on temples (as in Figure 104). Legs uniformly dark brown, with background vestiture interspersed with white, hair-like setae; male with short, suberect, hair-like setae along ventral margin of protibia near apex, setae longer than elsewhere on legs; tarsi clothed with sparse, white, hair-like setae. Dorsum mottled with poorly to well defined, irregular yellowish patches on apical portion of pronotum and after base of elytra; pronotal disk with dorsolongitudinal vitta of dense yellowish-white scales from apical margin to base; vitta well defined apically, faint to absent from middle to base. Pair of small dark patches at posterior pronotal margin on either side of midline, 4–6 irregular dark, maculae in transverse band slightly in front of midline, reaching lateral margins in most specimens, maculae sometimes fused into transverse band (best viewed without magnification), 4 small, circular, whitish maculae arranged in transverse row before middle; maculae after middle indistinct; 2 pairs of similarly formed but smaller maculae at apical pronotal margin, apical vitta and maculae in sharp contrast to background squamae, anterolateral region of pronotum with well defined ring of denser yellowish-white scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with darker background scales in center ( Figure 104); dorsal portion of ring not extending beyond middle of procoxa. Elytra without thin distinct fascia on anterior margin; humeri with distinct, rectangular, maculae of dense white scales on intervals 5 and 6; macula on intervals 5 longer than macula on intervals 6; maculae in sharp contrast to background squamae. Dorsum lacking other vittae or maculae. Mesosternum and metasternum covered with dense brown scales except transverse glabrous line at suture between sclerites, ventrites similarly clothed as metasternum. Scales course (individually distinct under low magnification), appressed to suberect.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes ( Figure 104), interrupted at back by glabrous V-shaped area where head articulates on pronotum, thin, medial longitudinal carinae within crown from apex of V-shaped area to center of crown. Eyes flat, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.2–1.3 X broader than long, broadest at base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one dirty white to brown, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle slightly cavernous, lateral margins produced, posterior margin slightly open. Scutellum visible, minute, diameter smaller than punctures at base of elytra. Elytra 1.1–1.3 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated near base; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, not elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs moderately long, hind femora exceeding elytral apices by 1/3 their length; mesofemora exceeding elytral apices. Profemora unidentate; profemoral tooth slightly larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange distinct, tapering to apex, male flange terminating before tibial apex, female flange tapering to apex (as in Figures 47, 50); dorsal margin weakly convex. Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with distinct suprauncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on significantly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male and female terminalia. As in generic description.

Material examined. ♂ Holotype: "Solomon Is., Bougainville, Kokure nr. Crown Prince Ra. 900 m., VI-11- 1956 / E. J. Ford, Jr., Collector" ( BPBM) . Paratypes: 2 ♂♂, "Solomon Is., Bougainville, Kokure nr. Crown, Prince Ra. 900 m., VI-11-1956 / E. J. Ford Jr., Collector" ( BPBM) ; 2 ♂♂, 3 ♀♀, " Solomon Is., Bougainville, Guata , 720 m., VI-19–21-1956 / E. J. Ford, Jr., Collector" ( BPBM) ; 5 ♂♂, 5 ♀♀, "Solomon Is., Bougainville, Kokure nr. Crown Prince Ra. 900 m., VI-10-1956 / E. J. Ford, Jr., Collector" ( BPBM) ; 2 ♀♀, "Solomon Is., Bougainville, Kokure nr. Crown Prince Ra. 900 m., VI-11-1956 / E. J. Ford, Jr., Collector" ( BPBM) ; 1 ♂, " Bougainville: NE, Mutahi. 700 m, 18km S.E. Tinputz / 8–14.III.1968 / Tawi, Collector, Bishop" ( MNHN) ; 1 ♀, " Bougainville: NE, Mutahi. 700 m, 18km S.E. Tinputz / 8–14.III.1968 / Tawi, Collector, Bishop" ( UMSP) ; 1 ♂, " Bougainville, Togerao 600 m, 15–21. 4, 1968/ R. Straatman, collectors, Bishop" ( MSNG) ; 1 ♂, 1 ♀, "Solomon Is., Bougainville, Kokure nr. Crown, Prince Ra. 900 m., VI-10-1956 / E. J. Ford, Jr., Collector" ( BPBM) ; 2 ♂♂, "Solomon Is., Bougainville, Kokure nr. Crown, Prince Ra. 900 m., VI-9-1956 / E. J. Ford, Jr., Collector" ( BPBM) ; 1 ♂, " Bougainville: NE, Mutahi. 700 m, 18 km S.E. Tinputz / 15–21.III.1968 / & R. Straatman, Collectors, Bishop Museum" ( BPBM) ; 1 ♂, " Bougainville: NE, Mutahi. 700 m, 18 km S.E. Tinputz / 1–7.III.1968 / Tawi, Collector, Bishop" ( SMTD) ; 1 ♀, " Solomon Is., Bougainville, Kukugai vill., 150 m, X.1960 / W. W. Brandt, Collector, Bishop" ( BPBM) ; 1 ♂, 1 ♀, "Solomon Is., Bougainville, Kokure nr. Crown, Prince Ra. 900 m., VI-10-1956 / E. J. Ford, Jr., Collector" ( USNM) ; 1 ♂, 1 ♀, " Solomon Is., Bougainville, Kokure , 690 m., VI-8–13-1956 / E. J. Ford, Jr., Collector" ( GPSC) ; 1 ♂, " Solomon Is., Bougainville, Kokure , 690 m., VI-8–13-1956 / E. J. Ford, Jr., Collector/ Asytesta not in keys det. R. T. Thompson, 1993/ Crypt. 314" ( BMNH) ; 1 ♀, "Solomon Is., Bougainville, Kokure nr. Crown, Prince Ra. 900m., VI-8-1956 / E. J. Ford, Jr., Collector" ( BMNH) .

Distribution. North Solomon Islands: Bouganville Island (= Papua New Guinea: North Solomon Province).

Etymology. This species is dedicated to R. T. Thompson, who first recognized this species as belonging to Asytesta and as new to science.