Formicinae

Kiran, Kadrï & Karaman, Celal, 2012, First annotated checklist of the ant fauna of Turkey (Hymenoptera: Formicidae), Zootaxa 3548, pp. 1-38 : 5-15

Subfamily Formicinae Latreille

Camponotus aegaeus Emery, 1915 : [11], [12], [17], [21] (Kiran & Aktaç 2006); [2], [6] ( Karaman et al. 2011); [4] (Lapeva-Gjonova & Kiran 2012).

as Camponotus libanicus var. aegaea : [3] ( Emery 1925a).

as Camponotus aegeus (sic!): Thrace (Radchenko 2007).

as Camponotus robustus Roger, 1863 : Anatolia ( André 1882c); Asia Minor ( Dalla Torre 1893). Comments: See the comments for C. robustus in species excluded from the list of Turkish ants.

Camponotus aethiops (Latreille, 1798) : [12] ( Mayr 1861; Donisthorpe 1950a); [1] ( Forel 1895; Aras & Aktaç 1987); [27] ( Emery 1897; Özdemir et al. 2008); [3], [27] ( Santschi 1921); [10] ( Schkaff 1924; Baroni Urbani 1964); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [61], [67], [71], [73], [74], [76], [80], [81] ( Aktaç 1988); [2] ( Aras & Aktaç 1992;1994); [23], [27], [42], [51] ( Ülgentürk 2001); [11], [12], [16], [17], [21] (Kiran & Aktaç 2006); [27], [67] ( Ionescu-Hirsh 2009); [2], [6] ( Karaman et al. 2011); [1] (Kiran & Karaman 2012); [4] (Lapeva-Gjonova & Kiran 2012). as Formica marginata Latreille, 1798 : Turkey ( Roger 1859).

as Camponotus maculatus subsp. aethiops var. concava Forel, 1888 : [12] ( Emery 1897).

as Camponotus maculatus r. aethiops var. marginata: Sultan Mt. ( Forel 1906) .

as Camponotus maculatus subsp. aethiops : [3], [10], [16] ( Forel 1911).

as Camponotus aethiops var. marginata : [9] ( Emery 1921a).

as Camponotus aethiops var. concava : Asia Minor ( Emery 1925a).

as Componotus (sic!) aethiops : [9], [59], [77] ( Aktaç 1976).

Camponotus aethiops subsp. escherichi Emery, 1925

as Camponotus maculatus subsp. oertzeni var. escherichi : [17] ( Emery 1897); [12] ( Emery 1898). as Camponotus aethiops var. escherichi : Asia Minor ( Emery 1925a).

Camponotus armeniacus Arnol'di, 1967: Turkey ( Radchenko 1996).

Camponotus baldaccii Emery, 1908 : [8] ( Espadaler & Lodos 1983); Thrace ( Agosti & Collingwood 1987; Radchenko 2007); Asia Minor ( Radchenko 1997a); [12] (Kiran & Aktaç 2006); [3] ( Ionescu-Hirsh 2009); [2], [6] ( Karaman et al. 2011).

as Camponotus maculatus subsp. dichrous var. baldaccii : [17] ( Emery 1897). as Camponotus maculatus baldaccii : Asia Minor ( Emery 1908a).

as Camponotus silvaticus subsp. baldaccii : [9] ( Aktaç 1976; 1978).

Camponotus boghossiani Forel, 1911 : [6] ( Forel 1911); [2] ( Karaman et al. 2011).

Camponotus buddhae Forel, 1892 : [74] ( Karavaiev 1926b).

Camponotus candiotes Emery, 1894 : [11], [12] (Kiran & Aktaç 2006); [2], [6] ( Karaman et al. 2011).

Camponotus cilicicus Emery, 1908 : [46] ( Emery 1908a).

Camponotus compressus subsp. cosensis Finzi, 1939: Asia Minor Islands ( Finzi 1939). as Camponotus maculatus sanctus var. cosensis : Asia Minor ( Emery 1908a).

as Camponotus compressus sanctus var. cosensis : [9] ( Emery 1921a).

as Camponotus compressus st. thoracicus var. cosensis : [3], [27] ( Santschi 1921).

Camponotus compressus subsp. symiensis Forel, 1910

as Camponotus compressus st. sanctus var. symiensis : [3] ( Santschi 1934).

Camponotus dalmaticus (Nylander, 1849) : [9] ( Aktaç 1976); Thrace ( Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [2] ( Aras & Aktaç 1992); [11], [12] (Kiran & Aktaç 2006).

as Camponotus lateralis var. dalmatica : [10] ( Schkaff 1924).

Camponotus fallax (Nylander, 1856) : [10] ( Schkaff 1924); Thrace ( Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [11], [21] (Kiran & Aktaç 2006).

as Formica fallax : Turkey ( Roger 1859).

Camponotus fedtschenkoi Mayr, 1877 : [76], [80] ( Aktaç 1988).

Camponotus fellah Dalla Torre, 1893

as Camponotus maculatus subsp. thoracicus var. fellah : [3] ( Forel 1911).

Camponotus festai Emery, 1894 : [36], Taurus Mts. ( Emery 1908a); [77] ( Aktaç 1976); [46], [49] ( Ionescu-Hirsh 2009).

as Camponotus maculatus subsp. festai: Taurus Mts. ( Emery 1898) .

as Camponotus maculatus r. caeciliae Forel, 1906: Taurus Mts. ( Forel 1906). as Camponotus maculatus r. festai : [36], Bolkar Mt., Taurus Mts. ( Forel 1913).

Camponotus gestroi Emery, 1878 : [76], [81] ( Aktaç 1988); Thrace ( Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [12] (Kiran & Aktaç 2006); [2], [6] ( Karaman et al. 2011).

as Camponotus gestroi subsp. creticus Forel, 1886 : [3] ( Forel 1911); [77] ( Aktaç 1976). as Camponotus gestroi r. creticus : [3] ( Forel 1913).

as Camponotus gestroi var. creticus : [3] ( Santschi 1921).

as Camponotus myrmentoma (sic!) gestroi var. creticus : Asia Minor ( Menozzi 1929). as Camponotus creticus : Turkey ( Kugler 1988).

Camponotus herculeanus (Linnaeus, 1758) : [70] ( Forel 1904); [12] ( Donisthorpe 1950a); [59] ( Aktaç 1976). as Formica herculeana : [10] ( Rigler 1852).

as Camponotus herculeanus var. montana Ruzsky, 1904 : [74] ( Karavaiev 1926b).

Camponotus ionius Emery, 1920 : [3], [9], [27] ( Aktaç 1976); [9] ( Aktaç 1978); [6] ( Karaman et al. 2011). as Camponotus sanctus var. ionia : [3] ( Santschi 1921).

as Camponotus samius var. ionia : Asia Minor ( Emery 1925a).

Camponotus kiesenwetteri ( Roger, 1859) : [9] ( Aktaç 1976); [6] ( Karaman et al. 2011).

Camponotus kurdistanicus Emery, 1898 : [44], [69] ( Emery 1920); [1] ( Aras & Aktaç 1987); Thrace (Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Radchenko 2007); [53] ( Ionescu-Hirsh 2009).

as Camponotus gestroi subsp. kurdistanicus : [69] ( Emery 1898).

Camponotus lateralis (Olivier, 1792) : [1], [10] ( Forel 1895); [12] ( Emery 1897); [3], [6], [16] ( Forel 1911); [10] ( Schkaff 1924; Donisthorpe 1950a); [1] ( Aras & Aktaç 1987); Thrace (Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Radchenko 2007); [2] ( Aras & Aktaç 1992;1994); [4], Koru Mt. (Aktaç et al. 1994); [11], [12], [16], [17], [21] (Kiran & Aktaç 2006); Taurus Mt., [46] ( Ionescu-Hirsh 2009); [2], [6] ( Karaman et al. 2011); [4] (Lapeva- Gjonova & Kiran 2012).

as Camponotus kosswigi Donisthorpe, 1950 : [8] ( Donisthorpe 1950a).

Camponotus libanicus André, 1881 : [3] ( Forel 1911); [46] ( Ionescu-Hirsh 2009). as Camponotus libanicus r. sahlbergi Forel, 1913 : Bolkar Mt. ( Forel 1913). as Camponotus libanicus subsp. sahlbergi: Bolkar Mt. ( Emery 1925a) .

Camponotus ligniperda (Latreille, 1802)

as Camponotus higniperda (sic!): [12] ( Donisthorpe 1950a). as Camponotus ligniperdus: Thrace (Radchenko 2007) .

Camponotus maculatus subsp. sylvaticomaculatus Dalla Torre, 1893

as Camponotus maculatus var. silvatico-maculatus: Asia Minor ( Dalla Torre 1893). as Camponotus maculatus subsp. dichrous var. silvatico-maculatas (sic!): [12] ( Emery 1898).

Camponotus oertzeni Forel, 1889 : Asia Minor ( Radchenko 1997a); [6] ( Karaman et al. 2011).

Camponotus piceus (Leach, 1825) : [15] ( Baroni Urbani & Aktaç 1981); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [1] ( Aras & Aktaç 1987); [2] ( Aras & Aktaç 1992;1994); [11], [12], [16], [17], [21] (Kiran & Aktaç 2006); [27] ( Özdemir et al. 2008); [2], [6] ( Karaman et al. 2011); [4] (Lapeva-Gjonova & Kiran 2012).

as Camponotus lateralis var. foveolatus (Mayr, 1853) : [1], [10] ( Forel 1895).

as Camponotus lateralis var. rectus Forel, 1892 : [1] ( Forel 1895).

as Camponotus lateralis var. atricolor (Nylander, 1849) : [70] ( Forel 1904); [3], [10], [16] ( Forel 1911); [67] ( Karavaiev 1926b).

as Camponotus lateralis subsp. picea var. atricolor : [10] ( Schkaff 1924).

as Camponotus piceus var. atricolor : Asia Minor ( Emery 1925b).

as Camponotus piceus subsp. picea var. atricolor : Asia Minor ( Baroni Urbani 1971). as Camponotus atricolor: Thrace ( Agosti & Collingwood 1987) .

Camponotus samius Forel, 1889 : [9], [10] ( Aktaç 1976); [1] ( Aras & Aktaç 1987); Thrace (Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [2] ( Aras & Aktaç 1992;1994); [11], [12], [16], [17], [21] (Kiran & Aktaç 2006); [2], [6] ( Karaman et al. 2011); [4] (Lapeva-Gjonova & Kiran 2012). as Camponotus maculatus samius var. spagnolinii Emery, 1905 : [10] ( Emery 1905).

as Camponotus maculatus r. samius var. bosphoricus Forel, 1906 : [10], [12], [16], Sultan Mt. ( Forel 1906). as Camponotus maculatus subsp. samius : [3], [10], [16] ( Forel 1911).

as Camponotus samius var. spagnolinii : [10] ( Schkaff 1924).

Camponotus sanctus Forel, 1904 : [2] ( Aras & Aktaç 1992;1994); [12] (Kiran & Aktaç 2006); [27] ( Ülgentürk 2001); [3], [27], [46], [67] ( Ionescu-Hirsh 2009); [2], [6] ( Karaman et al. 2011).

as Camponotus maculatus subsp. thoracicus var. sanctus : [3] ( Forel 1911).

as Camponotus compressus st. sanctus : [3], [27] ( Santschi 1921).

as Camponotus compressus subsp. sancta : Asia Minor ( Emery 1925a).

as Camponotus compressus subsp. sanctus : [9], [77] ( Aktaç 1976); [77] ( Aktaç 1978).

Camponotus shaqualavensis Pisarski, 1971

as Camponotus jaliensis subsp. shaqualavensis : [77] ( Aktaç 1976).

Camponotus sylvaticus (Olivier, 1792) : [10], [27] ( Donisthorpe 1950a); [15] ( Baroni Urbani & Aktaç 1981). as Camponotus maculatus subsp. sylvaticus : [17] ( Emery 1897).

Camponotus tergestinus Müller, 1921 : Thrace ( Agosti & Collingwood 1987; Radchenko 2007).

Camponotus thoracicus (Fabricius, 1804)

as Camponotus compressus subsp. thoracica: [10], [15], [20] ( Donisthorpe 1950a).

Camponotus truncatus (Spinola, 1808) : [3], [6] ( Forel 1911); Thrace (Aktaç et al. 1994; Radchenko 2007); [2], [6] ( Karaman et al. 2011); [1] (Kiran & Karaman 2012); [4] (Lapeva-Gjonova & Kiran 2012). as Colobopsis truncatus : [11], [21] (Kiran & Aktaç 2006).

Camponotus vagus (Scopoli, 1763) : [10], [12], [26], [28], [80] ( Donisthorpe 1950a); [1] ( Aras & Aktaç 1987); Thrace (Çamlıtepe & Aktaç 1987; Radchenko 2007); [4] (Aktaç et al. 1994; Lapeva-Gjonova & Kiran 2012).

Camponotus variegatus (Smith, F., 1858) : [3], [44] ( Mayr 1862). as Camponotus sylvaticus var. variegatus : Anatolia ( André 1882c).

Camponotus vogti Forel, 1906 : Taurus Mts. ( Forel 1906).

as Camponotus kurdistanus (sic!) var. vogti: Taurus Mts. ( Emery 1925a) .

Camponotus xerxes Forel, 1904 : Asia Minor ( Ionescu-Hirsh 2009).

as Camponotus carimatus var. xerxes : East of Turkey ( Emery 1905). as Camponotus maculatus r. thoracicus var. xerxes : [16] ( Forel 1906).

Cataglyphis aenescens (Nylander, 1849) : [1] ( Aras & Aktaç 1987); Thrace (Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Radchenko 2007); [1], [4] (Aktaç et al. 1994); [12] (Kiran & Aktaç 2006); [27] ( Özdemir et al. 2008); [4] (Lapeva-Gjonova & Kiran 2012).

as Myrmecocyctus cursor r. tancrei Forel, 1901: [70] ( Forel 1904).

as Cataglyphis cursor st. aenescens : [27] ( Santschi 1921).

as Cataglyphis cursor subsp. aenescens : [10] ( Schkaff 1924); [3], [10] ( Baroni Urbani 1964). as Cataglyhis (sic!) aenescens : [61], [67], [71], [74], [75], [76], [80], [81] ( Aktaç 1988).

Cataglyphis albicans ( Roger, 1859) : [15] ( Baroni Urbani & Aktaç 1981); [80], [81] ( Aktaç 1988); Thrace (Radchenko 2007).

Cataglyphis bicolor (Fabricius, 1793) : [53], [76] ( Donisthorpe 1950a).

as Cataglyphis bicolor subsp. megalocola (Förster, 1850): [46] ( Donisthorpe 1950a).

Cataglyphis bicolor subsp. rufiventris Emery, 1925: [9] ( Emery 1921a). as Cataglyphis bicolor st. orientalis var. rufiventris : [27] ( Santschi 1921).

Cataglyphis cursor (Fonscolombe, 1846) : [3], [10], [11], [27], [74] ( Donisthorpe 1950a).

as Myrmecocystus cursor: Sultan Mt. ( Forel 1906) .

Comments: The distribution of C. cursor is mentioned mainly as northern Mediterranean (from Spain to Turkey) in the old literature ( Forel 1892; Atanassov 1934; Donisthorpe 1950a; Bernard 1968). Recent studies show that in the northern Mediterranean region five species are possible. Cataglyphis piliscapa (Forel, 1901) and Cataglyphis sp. occur west of the Rhone valley in France. Cataglyphis cursor occurs east of the Rhone valley to Eastern Europe. Cataglyphis hellenica (Forel, 1886) occurs in Greece. Cataglyphis aenescens occurs in steppic parts of Eastern Europe, the higher parts of Anatolia and the Caucasus ( Agosti 1990). Therefore the occurrence of C. cursor in Eastern Europe and Anatolia is improbable and previous identifications of Turkish C. cursor are probably C. aenescens ( Lapeva-Gjonova et al. 2010) .

Cataglyphis frigidus (André, 1881)

as Myrmecocystus cursor var. frigidus : [45] ( Kohl 1905).

Cataglyphis kurdistanicus Pisarski, 1965 : [77] ( Aktaç 1976); Turkey ( Radchenko 1997b).

Cataglyphis laevior Santschi, 1929

as Cataglyphis bicolor var. laevior : [19] ( Donisthorpe 1950a).

Cataglyphis lividus (André, 1881) : Asia Minor ( Radchenko 1997b); Thrace (Radchenko 2007). as Cataglyphis livida : [77] ( Aktaç 1976; 1978).

as Myrmecocystus albicans var. mixtus Forel, 1895 : [1] ( Forel 1895). as Myrmecocystus albicans r. lividus: Taurus Mts. ( Forel 1906) .

as Myrmecocystus albicans var. mixta : [3] ( Forel 1911).

as Cataglyphis albicans subsp. viaticoides var. mixta : [1] ( Emery 1925a). as Cataglyphis albicans st. livida var. mixta : Turkey ( Santschi 1929); [3] ( Santschi 1934). as Cataglyphis albicans subsp. livida : [76] ( Donisthorpe 1950a).

as Cataglyphis albicans var. mixtus : [1] ( Agosti 1990).

Cataglyphis lunaticus Baroni Urbani, 1969

as Cataglyphis lunatica : [23] ( Baroni Urbani 1969).

Cataglyphis nodus (Brullé, 1833) : [9], [77] ( Aktaç 1976); [77] ( Aktaç 1978); [15] ( Baroni Urbani & Aktaç 1981); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [1] ( Aras & Aktaç 1987); [2] ( Aras & Aktaç 1992;1994); [11], [12], [16], [21] (Kiran & Aktaç 2006); [3], [20] ( Lush 2009); [4] (Lapeva-Gjonova & Kiran 2012).

as Myrmecocystus viaticus var. orientalis Forel, 1895 : [1] ( Forel 1895).

as Myrmecocystus bicolor var. orientalis : [12], [16], Sultan Mt. ( Forel 1906).

as Myrmecocystus viaticus subsp. orientalis : [3], [16] ( Forel 1911).

as Cataglyphis bicolor st. orientalis : [3] ( Santschi 1921).

as Cataglyphis bicolor var. nodus : [10] ( Schkaff 1924); [53] ( Donisthorpe 1950a). as Cataglyphis bicolor st. nodus : [3], [36] ( Santschi 1934).

as Cataglyphis bicolor subsp. nodus : Asia Minor ( Finzi 1939).

Cataglyphis ruber (Forel, 1903) : Asia Minor (Radchenko 1998). Comments: See the comments for Cataglyphis viaticoides .

Cataglyphis setipes (Forel, 1894) : [81] ( Aktaç 1988).

Comments: Cataglyphis setipes is distributed in Daghestan, Central Asia, Kazakhstan, Eastern Iran, Afghanistan and India ( Radchenko 1997b). According to its distribution, the Turkish record is doubtful and needs confirmation. Most probably, Turkish records are Cataglyphis nigriceps Arnol’di, 1964 (Radchenko, pers. comm.), distributed in Transcaucasus and Western Iran. These two species are very similar. The erect hairs on the body of C. setipes are fine and whitish, while on C. nigripes they are more coarse and blackish or brownish.

Cataglyphis viaticoides (André, 1881) : [9], [77] ( Aktaç 1976); [77] ( Aktaç 1978); [15] ( Baroni Urbani & Aktaç 1981); [1] ( Aras & Aktaç 1987; Aktaç et al. 1994); [11], [12], [16], [21] (Kiran & Aktaç 2006); Thrace (Radchenko 2007).

as Myrmecocystus albicans var. viaticoides : [3] ( Forel 1911).

as Catagliphis (sic!) albicans st. viaticoides : [3] ( Santschi 1934).

Comments: Radchenko (1997b) indicated that previous records of C. viaticoides from Turkey ( Aktaç 1976; Baroni Urbani & Aktaç 1981; Aras & Aktaç 1987) are probably misidentifications of C. ruber .

Cataglyphis viaticus (Fabricius, 1787) : [44] (Roger 1863).

as Formica viatica : Turkey ( Roger 1859).

as Myrmecocystus viaticus : Turkey ( André 1882b); [12] ( Emery 1897). Comments: This species has mainly a western Mediterranean distribution ( Spain, Tunisia, Algeria and Morocco; Emery 1908c) and its existence in Turkey is doubtful.

Formica anatolica Seifert & Schultz, 2009 : [20], [28], [42], [45] ( Seifert & Schultz 2009).

Formica cinerea Mayr, 1853 : [70] ( Forel 1904); [1] ( Aras & Aktaç 1987); Thrace (Çamlıtepe & Aktaç 1987; Radchenko 2007); [67], [71], [74] ( Aktaç 1988); [1], [4] (Aktaç et al. 1994).

as Formica cinerea var. armeniaca Ruzsky, 1905 : [59] ( Aktaç 1976).

as Formica imitans Ruzsky, 1902 : Thrace ( Agosti & Collingwood 1987; Radchenko 2007). as Formica balcanina Petrov & Collingwood, 1993 : [25], [75] ( Petrov & Collingwood 1993); [16] (Kiran & Aktaç 2006).

as Formica fuscocinerea Forel, 1874 : Thrace (Radchenko 2007 – misidentification). Comments: See the comments for F. fuscocinerea in species excluded from the list of Turkish ants.

Formica cinereofusca Karavaiev, 1929 : [59] ( Aktaç 1976); Thrace ( Agosti & Collingwood 1987; Radchenko 2007).

Formica clara Forel, 1886 : [3], [4], [8], [9], [20], [27], [28], [34], [36], [37], [51], [66], [67], [70], [73], [74], [76], [80] ( Seifert & Schultz 2009); [4] (Lapeva-Gjonova & Kiran 2012).

Formica cunicularia Latreille, 1798 : [59] ( Aktaç 1976; 1978); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [6] (Aktaç & Çamlıtepe 1987); [1] ( Aras & Aktaç 1987); [61], [67], [71], [73], [74], [75] ( Aktaç 1988); [2] (Aras & Aktaç 1994); [27] ( Ülgentürk 2001; Özdemir et al. 2008); [11], [12], [16], [21] (Kiran & Aktaç 2006); [3], [9], [12], [37], [70], [74], [75], Istranca Mt. ( Seifert & Schultz 2009); [4] (Lapeva-Gjonova & Kiran 2012).

as Formica fusca var. fusco-rufibarbis Forel, 1874: [70] ( Forel 1904); [10] ( Forel 1911).

as Formica fusco-rufibarbis: Sultan Mt. ( Forel 1906).

as Formica fusca subsp. glebaria var. rubescens Forel, 1904 : [10] ( Schkaff 1924).

as Formica fusca var. rubescens : [70] ( Donisthorpe 1950a).

as Formica glauca Ruzsky, 1895 : Thrace ( Agosti & Collingwood 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [11], [16] (Kiran & Aktaç 2006); [27] ( Özdemir et al. 2008); Kaz Mt. ( Aktaç & Karaman 2012).

Formica exsecta Nylander, 1846 : [73], [75] ( Aktaç 1988); Turkey (Seifert 2000b); Thrace (Radchenko 2007).

Formica foreli Bondroit, 1918 : [67], [73], [74] ( Aktaç 1988); Turkey (Seifert 2000b).

Formica fusca Linnaeus, 1758 : [12] ( Forel 1906; Donisthorpe 1950a); [10] ( Schkaff 1924); [3] ( Santschi 1934); [59] ( Aktaç 1976; 1978); Thrace (Çamlıtepe & Aktaç 1987; Radchenko 2007); [61], [67], [73], [74] ( Aktaç 1988); Istranca Mt. (Aktaç et al. 1994); [11], [16], [21] (Kiran & Aktaç 2006); [4] (Lapeva-Gjonova & Kiran 2012).

Formica gagates Latreille, 1798 : Asia Minor ( André 1882b); [12] ( Emery 1897); [10] ( Schkaff 1924); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [1] ( Aras & Aktaç 1987); [2] (Aras & Aktaç 1994); [11], [12], [16], [21] (Kiran & Aktaç 2006); [4] (Lapeva- Gjonova & Kiran 2012).

as Formica fusca gagates : Asia Minor ( Emery 1909b).

as Formica gagatoides Ruzsky, 1904 : Thrace ( Aras & Aktaç 1990 – misidentification). Comments: See the comments for Formica gagatoides in species excluded from the list of Turkish ants.

Formica georgica Seifert, 2002 : [66] Madenli-Buzlupınar Village (400m.), N40˚ 59ʹ, E40˚ 47ʹ, 12.VIII.2000, 8 workers; [66] Ïkizdere-Rüzgarlı Village (600m.), N40˚ 45ʹ, E40˚ 33ʹ, 13.VIII.2000, 6 workers; [73] Çamgeçidi (2150m.), N41˚ 0 4ʹ, E42˚ 30ʹ, 14.VIII.2000, 8 workers; [70] Şavşat (1550m.), N41˚ 14ʹ, E42˚ 21ʹ, workers; [70] Borçka (640m.), N41˚ 21ʹ, E41˚ 40ʹ, 17.VIII.2000, workers; [59] Uzungöl (1800m.), N40˚ 38ʹ, E40˚ 16ʹ, 17.VIII.2000, workers.

Comments: New record for Turkey.

Formica lemani Bondroit, 1917 : Thrace ( Agosti & Collingwood 1987); [61], [67], [71], [73], [74], [75] ( Aktaç 1988).

Formica lugubris Zetterstedt, 1838 : [61] Yeşilçat Village (1384m.), N39° 48', E39° 21', 25.VII.2004, 24 workers. Comments: New record for Turkey.

Formica pratensis Retzius, 1783 : [12] ( Forel 1906); [12], [76] ( Donisthorpe 1950a); [4] (Aktaç 1987; Aktaç et al. 1994); Thrace ( Aras & Aktaç 1990; Aktaç et al. 1998; Radchenko 2007); [1], [4] (Lapeva-Gjonova & Kiran 2012). as Formica rufa Linnaeus, 1761 : [4] (Lapeva-Gjonova & Kiran 2012 – misidentification).

Comments: Material collected in the European part of Turkey was misidentified as F. r u f a (Lapeva-Gjonova & Kiran 2012) and belongs to F. pratensis .

Formica rufa Linnaeus, 1761 : [10] ( Rigler 1852); [12] ( Donisthorpe 1950a); [12], [13], [22], [25], [27], [30], [31], [37], [43], [55], [70], [73] ( Baş 1973); Thrace ( Agosti & Collingwood 1987; Radchenko 2007); [12], [22], [25], [26], [31], [37], [44], [61], [67], [73], [74] (Aktaç 1987); [61], [67], [73], [74] ( Aktaç 1988); [23] ( Avcı et al. 2000).

Formica rufibarbis Fabricius, 1793 : [1] ( Forel 1895); [27] ( Santschi 1921; Özdemir et al. 2008); [10], [12], [27], [28], [53] ( Donisthorpe 1950a); [3], [10] ( Baroni Urbani 1964); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [79] ( Seifert & Schultz 2009). as Formica fusca subsp. rufibarbis : [10] ( Schkaff 1924).

Formica sanguinea Latreille, 1798 : [59] ( Aktaç 1976); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aktaç et al. 1994; Radchenko 2007); [73] ( Aktaç 1988); [12], [16] (Kiran & Aktaç 2006); [4] (Lapeva- Gjonova & Kiran 2012).

Formica truncorum Fabricius, 1804 : Asia Minor ( Czechowski et al. 2002).

Lasius alienus (Förster, 1850) : Asia Minor ( André 1882b); [12], [17], [27], [28] ( Emery 1897); [10], [27], [74] ( Wilson 1955); [10] ( Baroni Urbani 1964); [9], [59], [77] ( Aktaç 1976); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [1] ( Aras & Aktaç 1987); [61], [67], [71], [73], [74], [75], [76], [80], [81] ( Aktaç 1988); [2] ( Aras & Aktaç 1992;1994); [28] ( Seifert 1992); [23], [27] ( Ülgentürk 2001); [11], [12], [16], [21] (Kiran & Aktaç 2006); [27] ( Özdemir et al. 2008); [4], [10] (Lapeva- Gjonova & Kiran 2012); Eastern Black Sea ( Ocak & Aktaç 2012).

as Lasius niger r. alienus : [16], Sultan Mt. ( Forel 1906).

as Lasius niger subsp. alienus : [3] ( Forel 1911); [10] ( Schkaff 1924).

as Acanthomyops alienus : [10], [11], [12], [27], [70], [74] ( Donisthorpe 1950a).

Comments: After Seifert's (1992) separation of former L. alienus into L. alienus , L. paralienus and L. psammophilus , which are also found in Turkey, all old material identified as L. alienus should be reexamined.

Lasius austriacus Schlick-Steiner, 2003 : Turkey ( Seifert 2007).

Lasius balcanicus Seifert, 1988 : [12] Iznik (110m.), N40° 25', E29° 43', 30.VIII.1998, workers; [12] Ïznik-Sarısu Village (720m.), N40° 34', E29° 47', 07.VIII.1998, workers; [34] Ortaköy (7 km NE) (1140m.), N38° 44' E34° 02', 26.VIII.2004, 1 worker, leg and det. K.Kiran.

Comments: New record for Turkey.

Lasius bicornis (Förster, 1850) : Taurus Mts. ( Wilson 1955). as Acanthomyops bicornis : [10], [12] ( Donisthorpe 1950a).

Lasius brunneus (Latreille, 1798) : [44] (Roger 1863); Asia Minor ( André 1882b); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [6] (Aktaç & Çamlıtepe 1987); [1] ( Aras & Aktaç 1987); [61], [67] ( Aktaç 1988); [2] ( Aras & Aktaç 1992;1994); Anatolia ( Seifert 1992); [11], [12], [16], [21] (Kiran & Aktaç 2006); [4] (Lapeva-Gjonova & Kiran 2012); Eastern Black Sea ( Ocak & Aktaç 2012).

as Lasius niger subsp. brunneus : [10] ( Schkaff 1924).

as Acanthomyops brunnens (sic!): [10] ( Donisthorpe 1950a).

Lasius carniolicus Mayr, 1861 : Turkey ( Buschinger & Seifert 1997). New localities: [18] Sandıklı (1739m.), N 38° 28', E 30° 22', 04.IX.2008, 3 females, 5 males, 30 workers, leg. and det. K.Kiran.

Lasius distinguendus ( Emery, 1916) : Thrace ( Agosti & Collingwood 1987; Radchenko 2007). as Lasius umbratus var. nuda Bondroit, 1917 : [10] ( Schkaff 1924).

Lasius emarginatus (Olivier, 1792) : [3], [10] ( Santschi 1934); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aktaç et al. 1994; Radchenko 2007); Turkey ( Seifert 1992); [12] (Kiran & Aktaç 2006); [4] (Lapeva- Gjonova & Kiran 2012).

as Acanthomyops emarginatus : [12], [26] ( Donisthorpe 1950a).

Lasius flavoniger Seifert, 1992 : [36] ( Seifert 1992).

Lasius flavus (Fabricius, 1782) : [12], [17], [27] ( Emery 1897); [59] ( Aktaç 1976; 1978); Thrace ( Agosti & Collingwood 1987; Radchenko 2007); [61], [67], [73], [74] ( Aktaç 1988); [21] (Kiran & Aktaç 2006). as Acanthomyops flavus : [12] ( Donisthorpe 1950a).

Lasius fuliginosus (Latreille, 1798) : [17] ( Emery 1897); [10] ( Schkaff 1924); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [1] ( Aras & Aktaç 1987); [73] ( Aktaç 1988); [12], [21] (Kiran & Aktaç 2006); [4] (Lapeva-Gjonova & Kiran 2012).

Lasius lasioides (Emery, 1869) : Turkey ( Seifert 1992); Thrace (Radchenko 2007).

New localities: [20] Kaş (30km N), 26.V.2000, 4 workers; [36] Erdemli-Kızılen Village, 23.V-17. VI.2005, 4 workers; [36] Erdemli-Devrent, 04-23. V.2005, 3 workers; [36] Gülnar-Taşdüştü Village, N36˚ 30ʹ, E33˚ 0 7ʹ, 24.VI., 26.VII., 26.VIII.2006, 10 workers, leg. N.Jansson, det. K.Kiran.

Lasius meridionalis (Bondroit, 1920) : Thrace ( Agosti & Collingwood 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007).

Lasius mixtus (Nylander, 1846)

as Acanthomyops mixtus: Black Sea coast ( Donisthorpe 1950a).

Lasius myops Forel, 1894 : Thrace ( Agosti & Collingwood 1987; Radchenko 2007).

New localities: [70] Borçka (110m.), N 41˚ 21ʹ, E 41˚ 40ʹ, 14.VIII.2000, 2 workers; [70] Tırtıklı-Kafkasör upland (1200m.), N 41˚ 11ʹ, E 41˚ 41ʹ, 1 female, 3 workers; [4] Pehlivanköy-Kuştepe Village, 14.IV.2002, 4 workers; [11] Akdaġmadeni (1410m.), N 39˚ 39ʹ, E 35˚ 53ʹ, 24.VIII.2004, 28 males, workers; [11] Çayıralan (1600m.), N 39˚ 18ʹ, E 35˚ 38ʹ, 26.VIII.2004, 7 workers; [30] Safranbolu-Bulak Village (622m.), N 41˚ 15ʹ, E 32˚ 39ʹ, 14.VIII.2005, 14 workers; [21] Pamukova-Eskiyayla Village (1400m.), N 40° 33', E 30° 07', 20.VIII.1998, 7 workers, leg. and det. K. Kiran.

Lasius neglectus Van Loon, Boomsma & Andrasfalvy, 1990: [3], [9], [11], [19], [20], [28], [37], [55], [56], [79], [80] ( Seifert 2000a); [28], [80] ( Schultz & Seifert 2005); [1], [2], [10] (Cremer et al. 2008); [1], [2] ( Ulgevig et al. 2008); [1] (Kiran & Karaman 2012).

Lasius niger (Linnaeus, 1758) : Asia Minor ( André 1882b); [17] ( Emery 1897); [70] ( Forel 1904); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994); [1] ( Aras & Aktaç 1987); [75], [76] ( Aktaç 1988); [12], [21] (Kiran & Aktaç 2006).

as Formica nigra : [10] ( Rigler 1852).

as Acanthomyops niger : [53] ( Donisthorpe 1950a).

Comments: Seifert (1992) separated former L. niger into L. niger and L. platythorax according to their morphology and ecological preferences. After separation, both species were also recorded in Turkey. Therefore, former L. niger material should be reexamined.

Lasius paralienus Seifert, 1992 : Southern coasts of Anatolia ( Seifert 1992); [12] (Kiran & Aktaç 2006); Thrace (Radchenko 2007); [27] ( Özdemir et al. 2008); [4] (Lapeva-Gjonova & Kiran 2012). Comments: See the comments for L. alienus .

Lasius platythorax Seifert, 1991 : [21] (Kiran & Aktaç 2006); [4] (Lapeva-Gjonova & Kiran 2012). Comments: See the comments for L. niger .

Lasius psammophilus Seifert, 1992 : [16] (Kiran & Aktaç 2006); [4] (Lapeva-Gjonova & Kiran 2012). Comments: See the comments for L. alienus .

Lasius rabaudi (Bondroit, 1917) : Thrace ( Agosti & Collingwood 1987; Radchenko 2007).

Lasius schulzi Seifert, 1992 : [20], [27] ( Seifert 1992).

Lasius turcicus Santschi, 1921 : [31] ( Seifert 1992); [3] ( Steiner et al. 2004); [11], [12], [16], [21] (Kiran & Aktaç 2006); Thrace (Radchenko 2007); [2], [3], [4], [6], [10], [13], [14], [17], [18], [21], [22], [23], [28] (Cremer et al. 2008); [27] ( Özdemir et al. 2008).

as Lasius niger st. turcica : [27] ( Santschi 1921).

as Lasius niger st. turcicus : [10] ( Santschi 1934).

Lasius umbratus (Nylander, 1846) : [1] ( Aras & Aktaç 1987); Thrace (Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994); [61], [67], [73], [75] ( Aktaç 1988); [12] (Kiran & Aktaç 2006).

Lepisiota dolabellae ( Forel, 1911)

as Acantholepis frauenfeldi var. dolabellae : [3] ( Forel 1911). as Acantholepis frauenfeldi st. dolabellae: Menderes River ( Forel 1913) . as Acantholepis frauenfeldi var. dolabella (sic!): [3] ( Santschi 1934).

Lepisiota frauenfeldi (Mayr, 1855) : [11], [12], [17], [21] (Kiran & Aktaç 2006); Thrace (Radchenko 2007). as Acantholepis frauenfeldi : [1] ( Forel 1895); [3], [16] ( Forel 1911); [10] ( Schkaff 1924); [10], [11], [36], [49] ( Donisthorpe 1950a); [3] ( Baroni Urbani 1964); [9], [77] ( Aktaç 1976; 1978); [15] ( Baroni Urbani & Aktaç 1981); Thrace ( Aras & Aktaç 1990; Aktaç et al. 1994); [2] ( Aras & Aktaç 1992;1994).

as Acantholepis fraunfeldi (sic!): [6] (Aktaç & Çamlıtepe 1987).

Lepisiota melas ( Emery, 1915)

as Acantholepis frauenfeldi st. melas : Asia Minor ( Santschi 1917). as Acantholepis frauenfeldi subsp. melas : [9] ( Emery 1921a). as Acantholepis frauenfeldi var. melas : Asia Minor ( Finzi 1939).

Lepisiota syriaca (André, 1881)

as Acantholepis frauenfeldi var. syriaca : [46] ( Viehmeyer 1922). as Acantholepis syriaca : Turkey ( Kugler 1988).

Nylanderia jaegerskioeldi (Mayr, 1904) : [36], [46], [58] (Karaman & Kiran 2012).

Nylanderia vividula (Nylander, 1846)

as Paratrechina vividula : [10], [56] ( Donisthorpe 1950a); Thrace (Radchenko 2007).

Plagiolepis ampeloni (Faber, 1969) : Turkey ( Trontti et al. 2006). as Aporomyrmex cf. ampeloni : [31] ( Heinze & Kauffmann 1993).

Plagiolepis ancyrensis Santschi, 1920 : Turkey ( Kugler 1988).

as Plagiolepis maura var. ancyrensis : [27] ( Santschi 1920); [9] ( Emery 1921a). as Plagiolepis maura var. angyrensis (sic!): [27] ( Santschi 1921).

as Plagiolepis pallescens var. ancyrensis : Asia Minor ( Emery 1925a).

Plagiolepis pallescens Forel, 1889 : [9] ( Aktaç 1976; 1978); Thrace ( Agosti & Collingwood 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [1] ( Aras & Aktaç 1987); [61], [67], [71], [74], [81] ( Aktaç 1988); [2] ( Aras & Aktaç 1992); [11], [12], [16], [17] (Kiran & Aktaç 2006); [27] ( Özdemir et al. 2008).

as Plagiolepis pygmaea var. pallescens : [3] ( Forel 1911).

Plagiolepis pallescens subsp. maura Santschi, 1920

as Plagiolepis pallescens var. maura : Asia Minor ( Emery 1921b); Turkey ( Sharaf et al. 2011).

Comments: Recently Sharaf et al. (2011) studied the Plagiolepis of the Arabian Peninsula. They gave the distribution of P. pallescens subsp. maura as North Africa, Egypt, Oman, Saudi Arabia, Turkey and Yemen. On the other hand, they did not study Turkish material and probably their record is based on Emery’s data. This species inhabits mainly desert habitats and is doubtful for Turkey. This record needs confirmation.

Plagiolepis pygmaea (Latreille, 1798) : [1] ( Forel 1895; Aras & Aktaç 1987); [12] ( Emery 1897); [16] ( Forel 1906); [3], [16] ( Forel 1911); [10] ( Schkaff 1924); [59] ( Aktaç 1976); [76], [80], [81] ( Aktaç 1988); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [2] ( Aras & Aktaç 1992;1994); [11], [12], [16], [21] (Kiran & Aktaç 2006); [1], [4] (Lapeva-Gjonova & Kiran 2012). as Plagiolepis pigmaea (sic!): [6] (Aktaç & Çamlıtepe 1987).

as Plagliolepis (sic!) pygmaea : [10], [52], [53] ( Donisthorpe 1950a).

Plagiolepis pygmaea subsp. minu Forel, 1911

as Plagiolepis pygmaea var. minu : [6] ( Forel 1911).

Plagiolepis taurica Santschi, 1920 : [11], [12], [16], [21] (Kiran & Aktaç 2006); [4] (Lapeva-Gjonova & Kiran 2012).

as Plagiolepis vindobonensis Lomnicki, 1925 : Thrace ( Agosti & Collingwood 1987); [27] ( Ülgentürk 2001; Özdemir et al. 2008).

as Plagiolepis tauricus: Thrace (Radchenko 2007) .

Plagiolepis xene Stärcke, 1936 : [2] ( Kiran & Aktaç 2007); Thrace (Radchenko 2007).

Polyergus rufescens (Latreille, 1798) : Thrace (Çamlıtepe & Aktaç 1987; Radchenko 2007); [4] (Aktaç et al. 1994); [34], [40] ( Schulz & Sanetra 2002).

as Formica rufescens : [10] ( Rigler 1852).

Prenolepis nitens (Mayr, 1853) : Turkey ( André 1882b); [10] ( Forel 1911; Schkaff 1924); Bolkar Mt. ( Forel 1913); [74] ( Donisthorpe 1950a); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [23], [28] ( Ülgentürk 2001); [16] (Kiran & Aktaç 2006); [4] (Lapeva- Gjonova & Kiran 2012).

as Formica crepusculascens Roger, 1859 : North of Turkey ( Roger 1859).

as Lasius crepusculascens : North of Turkey ( Mayr 1861).

as Prenolepis imparis r. nitens : [10] ( Forel 1895).

as Prenolepis imparis subsp. nitens : Asia Minor ( Emery 1910).

Proformica kobachidzei Arnol'di, 1968: [59] ( Aktaç 1976); [15] ( Baroni Urbani & Aktaç 1981); Thrace (Radchenko 2007); [4] (Lapeva-Gjonova & Kiran 2012).

Proformica korbi (Emery, 1909) : [27] ( Dlussky 1969); [1] ( Aras & Aktaç 1987); Thrace (Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990); [61], [67], [73], [74], [75], [80], [81] ( Aktaç 1988); [2] (Aras & Aktaç 1994); [28] ( Tinaut et al. 2010).

as Formica korbi: Sultan Mt. ( Emery 1909b) .

as Proformica corbi (sic!): [1], [4] (Aktaç et al. 1994).

Proformica kosswigi (Donisthorpe, 1950)

as Acanthomyops kosswigi : [74] ( Donisthorpe 1950d).

Proformica nasuta (Nylander, 1856)

as Formica nasuta : [1] ( Forel 1895); [45] ( Kohl 1905); [16] ( Forel 1906).

Comments: Although Forel and Kohl recorded this species from various localities in Turkey, our Proformica material from Edirne, one of the areas Forel collected, does not match the characters of P. n a s u t a. The material is very close to P. striaticeps with some small differences (Galkowski, pers. comm.). Here we decided to leave this species in the list of Turkish ant fauna, but it needs further investigation.

Proformica pilosiscapa Dlussky, 1969

as Proformica pilosiscapus : [61], [67], [71], [74], [75], [76], [80] ( Aktaç 1988).

Proformica striaticeps ( Forel, 1911) : Turkey ( Collingwood & Heatwole 2000).

Rossomyrmex anatolicus Tinaut, 2007 : [28] ( Tinaut 2007); [28], [51] ( Tinaut et al. 2010).

as Rossomyrmex minuchae Tinaut, 1981 : [51] ( Schulz & Sanetra 2002 – misidentification).

Comments: See the comments Rossomyrmex minuchae in species excluded from the list of Turkish ants.