Paracoenia (Paracoenia) wirthi Mathis,

Mathis, Wayne N. & Marinoni, Luciane, 2016, Revision of Ephydrini Zetterstedt (Diptera: Ephydridae) from the Americas south of the United States, Zootaxa 4116 (1), pp. 1-110: 24-27

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Paracoenia (Paracoenia) wirthi Mathis


5. Paracoenia (Paracoenia) wirthi Mathis 

Figs. 27–34View FIGURES 27 – 29View FIGURES 30 – 34, Map 5

Paracoenia (Paracoenia) wirthi Mathis 1975: 78  [ United States. California. Inyo: Tecopa Hot Springs (1.6 km N); HT ♂, CAS (12033)].— Mathis 1980: 9 –11 [revision].— Mathis and Zatwarnicki 1995: 251 [world catalog].

Diagnosis. This species is distinguished from congeners, especially P. bisetosa  , by the following combination of characters: Medium-sized shore flies, body length 3.12–3.65 mm; subshiny dorsally, bluish green to green metallic luster.

Head ( Figs. 27–28View FIGURES 27 – 29): Mesofrons greenish blue, shiny; fronto-orbital areas dark brown, microtomentose to subshiny. Face densely microtomentose, grayish tank, dorsum of facial hump slightly darker than face, grayer; paravertical setae shorter than their distance apart; gena-to-eye ratio 0.30; eye ratio 0.64, height-to-length ratio 0.91; eye-width-to-face-length ratio 0.44.

Thorax: Mesonotum ( Fig. 29View FIGURES 27 – 29) with anterior area slightly microtomentose, becoming subshiny to shiny posteriorly; acrostichal setulae in 3–4 rows anteriorly, becoming irregular, 5–6 rows posteriorly. Mesopleuron with more subdued coloration than mesonotum, mostly olivaceous gray. Wing nearly transparent to faintly brown.

Abdomen: Concolorous with mesonotum; digitiform lateral process of male tergite 5 elongate, acutely pointed apically; male sternite 5 ( Fig. 34View FIGURES 30 – 34) broadly U-shaped with posteriorly extended arms oriented posteriorly, acutely pointed, anterior margin somewhat truncate, posterior margin moderately deeply emarginate. Male terminalia ( Figs. 30–32View FIGURES 30 – 34): Surstylus in posterior view with base robustly developed, width 3 X apical width, thereafter abruptly narrowed with apical extension narrowly digitiform; medial triangular process less than half length of surstylar arm, not cleft apicomedially; gonite more or less triangular, sider basally, apical half tapered, apex digitiform, apex rounded. Female ventral receptacle as in Fig. 33View FIGURES 30 – 34 with large, mushroom-like operculum.

MAP 5. Distribution map for Paracoenia wirthi Mathis. 

Type material. The holotype male is labeled “ USA. CALIF. Inyo Co 1 mi N Tecopa Hot Spg 24 June 1974 [,] Wayne N. Mathis/ HOLOTYPE Paracoenia wirthi Mathis  [red; 12033].” Allotype and 21 paratypes (7 ♂, 14 ♀; CAS, USNM) bear the same label data as the holotype. Other paratypes as listed in Mathis (1975: 78). The holotype male is double mounted (minute nadel), is in excellent condition, and is deposited in the CAS (12033).

Type locality. United States. California. Inyo: Tecopa Hot Springs (1.6 km N; 35 ° 52.7 'N, 116 ° 13.9 'W); HT ♂, CAS (12033).

Other specimens examined. MEXICO. Baja California: Guadalupe Canyon (32 °09.3'N, 115 ° 47.4 'W), 19 May 1957, F. X. Williams (1 ♂, 3 ♀; CAS). Distrito Federal: Mixquic, Chalco Lake (9 km NW; 19 ° 15.5 'N, 98 ° 58.6 'W), 1 Aug 1965, K. R. Valley (4 ♂, 16 ♀; CAS, USNM).

Distribution (Map 5). Nearctic: United States (California). Neotropical: Mexico (Baja California, Distrito Federal).

Remarks. This species is similar to P. bi s pi no s a but is slightly smaller in body length and less setulose. The ventral margin of the male sternite five is broadly U-shaped, and the connecting bridge is wide; the male tergite five is produced into an acutely pointed process; and the surstylus is more robustly developed basally, and the distal surstylar extensions are narrowly digitiform; the gonite is narrowed toward the apex, and the apex is digitiform, apically rounded.


California Academy of Sciences


Smithsonian Institution, National Museum of Natural History














Paracoenia (Paracoenia) wirthi Mathis

Mathis, Wayne N. & Marinoni, Luciane 2016

Paracoenia (Paracoenia) wirthi

Mathis 1995: 251
Mathis 1980: 9
Mathis 1975: 78