Panjange togutil Huber, 2019

Huber, Bernhard A. & Carvalho, Leonardo S., 2019, Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae), Zootaxa 4546 (1), pp. 1-96 : 82-84

Panjange togutil Huber sp. n.

Figs 304–307 , 312–315 , 319–321

Panjange Ind 109: Eberle et al. 2018 (molecular data); Huber et al. 2018: fig. 10. Type material. INDONESIA: ♂ holotype, ZFMK (Ar 20663), Halmahera, degraded forest along river near Sofifi (0.715°N, 127.586°E), 130–180 m a.s.l., 11.xi.2009 (S. Sutono).

Other material examined. INDONESIA: 3♂ 5♀ 7 juvs, ZFMK (Ar 20664–65), and 7 juvs in pure ethanol, ZFMK (Ind203), same data as holotype .

Etymology. The species is named for the Togutil people in the interiors of north and central Halmahera; noun in apposition.

Diagnosis. Males are easily distinguished from other known representatives of the cavicola group by strongly widening distal part of procursus ( Figs 312–313 ; without dorsal process as in P. cavicola ) and by color pattern on carapace ( Fig. 304 ; anterior half pale whitish, posterior half dark brown); females differ (except from P. cavicola ) by short epigynal scape ( Figs 315 , 319–320 ); both males and females also distinguished by posteriorly widened abdomen (in dorsal view; Figs 304–307 ).

Description. Male (holotype). MEASUREMENTS. Total length 3.9, carapace width 1.0. Distance PME-PME 390 µm; diameter PME 90 µm; distance PME-ALE 30 µm; AME absent. Leg 1: 36.9 (8.5 + 0.4 + 8.7 + 17.4 + 1.9), tibia 2: 5.5, tibia 3: 3.0, tibia 4: 4.5; tibia 1 L/d: 109.

COLOR (in ethanol). Carapace anterior half pale whitish, posterior half dark brown; ocular area and clypeus also brown; sternum whitish; legs pale ochre-yellow, brown rings in patella area and at tibia-metatarsus joint. Abdomen pale ochre-yellow, with dark marks dorsally and laterally.

BODY. Habitus as in Fig. 304 . Eye triads on stalks ( Fig. 314 ), each stalk with process ending in two tips each. Thoracic furrow absent. Clypeus unmodified. Sternum wider than long (0.62/0.50), unmodified. Abdomen cylindrical, widened posteriorly, pointed dorso-posteriorly.

CHELICERAE. As in Fig. 314 , with pair of simple weakly sclerotized processes proximally.

PALPS. As in Figs 312–313 , symmetric; coxa with small but distinct retrolateral-ventral apophysis, trochanter with retrolateral-dorsal process, femur with rounded dorsal process proximally; femur-patella joints strongly shifted towards prolateral side; tarsus with long whitish dorsal process with tarsal organ near tip; procursus complex, distal part apparently hinged, proximal part with two ventral processes, without ventral parallel ridges; brush of hairs dorsally between proximal and distal parts; distal part wide and flat; genital bulb oval, with two processes lying parallel to each other: semitransparent embolus and slightly sclerotized long bulbal apophysis (‘appendix’; ba in Fig. 312 ).

LEGS. Without spines and curved hairs, few vertical hairs; retrolateral trichobothrium of tibia 1 at 3%; tibia 1 without prolateral trichobothrium (present on other tibiae); tarsus 1 with ~30 pseudosegments, only distally a few fairly distinct.

Male (variation). Tibia 1 in 3 other males: 9.0, 9.5, 9.9.

Female. In general similar to male ( Figs 306–307 ) but eye triads on low humps, closer together (distance PME-PME 210 µm) and clypeus not darkened (only dark band between eye triads). Carapace pattern in most females like in males (but with light median line dividing dark mark), in one female different ( Fig. 307 ). Tibia 1 in 5 females: 6.7–8.0 (mean 7.5). Epigynum weakly sclerotized ( Fig. 319 ), with short folded scape. Internal genitalia with small median pocket and pair of large pore plates ( Figs 315 , 321 ).

Distribution. Known from type locality only ( Fig. 350 ).

Natural history. The spiders were found in a disturbed forest ( Fig. 341 ), on the undersides of green leaves about 1–2 m above the ground. The indistinct domed webs were connected to the undersides of the leaves at their apex, with a few silk lines extending beyond the leaves. Most specimens were found in a dryer patch of forest higher up on the hill.

Figures

FIGURE 350. Known distribution of Panjange in eastern Indonesia (not shown: record of P. alba from Ambon; see Huber 2011, fig. 435).

FIGURES 300–307. Panjange spp. live specimens. 300–303. P. thomi sp. n., adult and penultimate instar males, and female with egg sac, from Gunung Meja. 304–307. P. togutil sp. n., adult and penultimate instar males, females with and without eggsac, from near Sofifi.

FIGURES 312–313. Panjange togutil sp. n. (ZFMK, Ar 20664), left male palp, prolateral and retrolateral views. ba, bulbal apophysis; em, embolus; gb, genital bulb; pr, procursus; tp, tarsal process. Scale line: 0.5.

FIGURES 314–315. Panjange togutil sp. n. (ZFMK, Ar 20664–65), male prosoma and chelicerae, oblique frontal view, and cleared female genitalia, dorsal view. Scale lines: 0.3.

FIGURES 316–321. Panjange spp., epigyna in ventral views, and cleared female genitalia in ventral and dorsal views. 316– 318. P. thomi sp. n. (ZFMK, Ar 20662). 319–321. P. togutil sp. n. (ZFMK, Ar 20665). Scale lines: 0.3.

FIGURES 336–341. Selection of collecting sites and habitats. 336. Forest above Mokwam (West Papua), type locality of Arnapa arfak sp. n. and Aetana mokwam sp. n. 337. Tinoor Fall near Tomohon (North Sulawesi), type locality of Arnapa tinoor sp. n. 338. Entrance to Caverna Porco do Mato II (Rio Grande do Norte), type locality of Pinocchio barauna sp. n. 339. Desert between Sur and Kamil (Oman), type locality of Magana velox sp. n. 340. Bromeliad at Reserva Particular do Patrimônio Nacional Serra Bonita (Bahia), type locality of Psilochorus bromelicolus sp. n. (note pair of spiders among leaves). 341. Disturbed forest near Sofifi (Halmahera), type locality of Giloloa sofifi sp. n. and Panjange togutil sp. n.