Zamarada secutaria ( Guenée, 1858 )

Sciarretta, Andrea & Hausmann, Axel, 2020, The Geometridae of Ethiopia III: genus Zamarada (Lepidoptera: Geometridae Ennominae, Cassymini), Zootaxa 4894 (3), pp. 301-328: 308

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Zamarada secutaria ( Guenée, 1858 )


Zamarada secutaria ( Guenée, 1858)  

( Figs 14, 15 View FIGURES 10–18 , 41 View FIGURES 34–41 )

Stegania secutaria Guenée, [1858]   : 45. Locus typicus: Abyssinie [ Ethiopia] (Holotype ♀ in MNHN).

Literature records. Takusa: 6 miles S.W. of Lake Tana ( Fletcher 1974).

Examined material. 1♀: Ethiopie, Sidamo Wondo Genet , 2400 m, 18.11.1978, P.C. Rougeot (Coll. Herbulot pr. n. [1]5828); 1Ƌ   : Ethiopia, West Shewa, Ambo, 2200 m a.s.l., v.2007, leg. V. Kravchenko & G. Müller ( ZSM G 20201 View Materials / Ƌ / BC ZSM Lep 09909); 1Ƌ: S   . ETHIOPIA, Oromia, 13km S Agere Maryam , 1960 m (lux), 5.5149°N 38.2529°E, 7.XI.2010, leg. De Freina, Hacker, Peks, Schreier ( ZSM G 20211 View Materials / Ƌ / LEP-SS-00521 failure) GoogleMaps   ; 1♀: S. ETHIOPIA, Sidamo, 16km SW Kibre Mengist, 1700m, 5.8107°N 38.8880°E, 25.–26.III.2009, lg. R. Beck, M. Dietl ( ZSM G 20355 View Materials / ♀ / BC ZSM Lep 45490); 1Ƌ GoogleMaps   : ÄTHIOPIEN, ( EIP)   Äthiopisches Hochland, Bedele 3km N, 2050m, 08°28.571′N, 36°28.497′O, 10.vii.2015, Beck R. & R. Wanninger leg. ( ZSM G 20316 View Materials / Ƌ / BC ZSM Lep 101616); 1Ƌ: S GoogleMaps   . Ethiopia, Oromia, 6km ESE Jimma , 1850m (lux), 07°39‘34“N 36°53‘20“E, 30.10.2010, leg. de Freina, Hacker, Peks, Schreier ( ZSM G 20532 View Materials / Ƌ) GoogleMaps   ; 1♀: S. Ethiopia, Oromia, Abiyata –Shala-Hayak NP, 1646m (lux), 07°31‘27“N 38°39‘44“E, 16.II.2012, leg. de Freina, Hacker, Peks, Schreier ( ZSM G20533 View Materials / ♀) GoogleMaps   ; 1♀: S. Ethiopia SN, 2,65km ESE Wondo Genet , 1950m (lux), 07°04‘36“N 38°38‘30“E, 5.XI.2010, leg. de Freina, Hacker, Peks, Schreier GoogleMaps   ; 1♀: ETHIOPIA, Oromia, 7km S Kibre Mengist , 1500m, 21.IV.2009, 5°48′46″N 38°57′41″E, S. Naumann, H. Schnitzler ( ZSM G 20804 View Materials / ♀); GoogleMaps   1Ƌ: S. ETHIOPIA—SN Arba Minch, Nechisar NP, 2.75 km sw headquarter, 1170 m (lux) 06°00’13,6’’N 37°33’23,4’’E, 22.II.2012, leg. Hacker & Schreier ( ZSM G 20813 View Materials / Ƌ) GoogleMaps   .

Description. Adult Ƌ ( Fig. 14 View FIGURES 10–18 ): forewing length: 10–12 mm. Hyaline area of wings light yellow, irrorated with brown and black; basal and medial fasciae in fore- and hindwing poorly defined or absent; costa of forewing light brown, irrorated with brown or black, especially in the medial part; discal spot on forewing grey or light brown, black-ringed; discal spot on hindwings punctiform and black. Non-hyaline terminal area of fore- and hindwings light brown, variably irrorated with brown or black. Termen yellowish-brown, interneurally black-spotted, better marked between R 4 and M3 on forewing.

Male genitalia ( Fig. 41 View FIGURES 34–41 ). Uncus digitate, pointed apically. Ventral plate of gnathus as long as uncus, tapered and scobinate with rounded tip. Dorsal process of valva with a narrow, strongly sclerotized projection, apically spined. Dorsal process stout from the projection toward the base, thin and curved from the projection to the apex, which is pointed. Fulcrum extending beyond aedeagus, curved, with the apex bearing 2–4 stout thorn-shaped spines. Aedeagus with a scobinate ridge just below apex and a single cornutus-like lateral projection.

Differential diagnosis. The shape of dorsal process of valva with its projection and fulcrum are the diagnostic characters in male genitalia to discriminate Z. secutaria   from Z. chrysothyra   and Z. amicta   .

Genetic data. BIN: BOLD:AAH8489 (n=3 from Ethiopia). Genetically heterogeneous, one specimen (BC ZSM Lep 45490) diverging by 1.3 %. Nearest species: Z. metallicata Warren, 1914   (4.4%)

Distribution. Algeria, Egypt, Ethiopia, Kenya, Niger, Saudi Arabia, Somalia, Sudan ( Guenée 1858; Warren & Rothschild 1905; Rothschild 1915, 1921; Fawcett 1916; Prout, 1916; Wiltshire 1947).

Remarks. So far, only the female was known, unambiguous attribution of males to females was possible by DNA barcodes.


Royal British Columbia Museum - Herbarium


Bavarian State Collection of Zoology


Departamento de Geologia, Universidad de Chile