Coprophilus striatipennis, Eppelsheim, 1894: 291

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1067-1806 : 1067-1806

publication ID

https://doi.org/ 10.1206/0003-0090.265.1.1

persistent identifier

https://treatment.plazi.org/id/03E64505-FE91-5E10-FF6E-A287FD6E96B9

treatment provided by

Felipe

scientific name

Coprophilus striatipennis
status

 

striatipennis Eppelsheim, 1894: 291 View in CoL ( Zonyptilus , cited as Zonoptilus ; Type locality:

Altai Gebirge).

— Fauvel, 1897a: 229 ( Coprophilus ; type locality is Monts Alai not Monts Altai).

— Bernhauer, 1908e: 330 ( Coprophilus ; synonym of pennifer ).

— Bernhauer and Schubert, 1911: 89 ( Coprophilus ; synonym of pennifer ).

— Scheerpeltz, 1933: 1076 ( Coprophilus ; monstrosity of pennifer ).

obscurus Reitter, 1894: 177 ( Zonyptilus , cited as Zonoptilus ; variety of pennifer ; Type locality: Araxes).

— Reitter, 1894a: 292 ( Zonyptilus , cited as Zonoptilus ; characters; Araxes).

— Bernhauer, 1908e: 330 ( Coprophilus ; variety of pennifer ).

— Bernhauer and Schubert, 1911: 89 ( Coprophilus ; variety of pennifer ).

— Scheerpeltz, 1933: 1076 ( Coprophilus ; variety of pennifer ).

araxis Reitter, 1894: 178 ( Zonyptilus , cited as Zonoptilus ; variety of pennifer ; Type locality: Araxes).

— Reitter, 1894a: 292 ( Zonyptilus , cited as Zonoptilus ; characters; Arax valley).

— Bernhauer, 1908e: 330 ( Coprophilus ; synonym of pennifer ).

— Bernhauer and Schubert, 1911: 89 ( Coprophilus ; variety of pennifer ).

— Scheerpeltz, 1933: 1076 ( Coprophilus ; variety of pennifer ).

obsoletus Reitter, 1894a: 292 ( Zonyptilus , cited as Zonoptilus ; variety of sellula ; Type locality: Araxesthal).

— Bernhauer, 1908e: 330 ( Coprophilus ; synonym of pennifer ).

— Bernhauer and Schubert, 1911: 89 ( Coprophilus ; synonym of pennifer ).

marginalis Reitter, 1894a: 292 ( Zonyptilus , cited as Zonoptilus ; variety of sellula ; Type locality: Nördliche Mongolei; Ostsibirien).

— Bernhauer, 1908e: 330 ( Coprophilus ; variety of pennifer ).

— Bernhauer and Schubert, 1911: 89 ( Coprophilus ; variety of pennifer ).

— Scheerpeltz, 1933: 1076 ( Coprophilus ; variety of pennifer ).

eppelsheimi Fauvel, 1897a: 227 , 228 ( Coprophilus ; Type locality: Mongolie, Amdo; Sibérie or., Vladivostok ; Chine, Pekin, Tien-Tsin , Shangai).

— Bernhauer, 1908e: 330 ( Coprophilus ; synonym of pennifer ).

— Bernhauer and Schubert, 1911: 89 ( Coprophilus ; synonym of pennifer ).

dimidiatipennis Fauvel, 1900d: 223 ( Coprophilus ; Type locality: Transcaspienne : Aschabad).

— Bernhauer, 1908e: 330 ( Coprophilus ; variety of pennifer ).

— Bernhauer and Schubert, 1911: 89 ( Coprophilus ; variety of pennifer ).

— Scheerpeltz, 1933: 1076 ( Coprophilus ; variety of pennifer ).

pentatoma Fauvel, 1897a: 227 , 228 ( Coprophilus ; Type locality: Mons Alai Turkestan; [Note: See lectotype designation by Tóth, 1991]. Also cited from Baku; Dobrudja ).

— Bernhauer and Schubert, 1911: 89 ( Coprophilus ; catalog).

— Bernhauer, 1908e: 329, 330 ( Coprophilus ; subgenus Zonyptilus , cited as Zonoptilus ; characters; catalog).

— Scheerpeltz, 1933: 1076 ( Coprophilus ; synonym of longicornis ).

— Scheerpeltz, 1933: 1076 ( Coprophilus ; catalog).

— Gridelli, 1934: 72 ( Coprophilus ; Karakoram region).

— Herman, 1970: 368 ( Coprophilus ).

— Tóth, 1991: 94 ( Coprophilus ; subgenus Zonyptilus , cited as Zonoptilus ; characters; lectotype designation: Mons Alai Turkestan; Turkestan).

— Tóth, 1993: 375 ( Coprophilus ; characters).

— Kashcheev, 1999: 149 ( Coprophilus ; characters).

- DISTRIBUTION: Azerbaijan, Kyrgyzstan.

obscurus Luze, 1904: 79 ( Coprophilus ; aberration of bimaculatus ; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]).

piceus Solsky, 1867: 88 ( Zonyptilus , cited as Zonoptilus ; Type locality: Sarepta).

— Reitter, 1894: 177 ( Zonyptilus , cited as Zonoptilus ; characters; Russia; Austria; Hungary).

— Reitter, 1894a: 292 ( Zonyptilus , cited as Zonoptilus ; characters; Russia; Austria; Hungary).

— Ganglbauer, 1895: 677 ( Coprophilus ; subgenus Zonyptilus , cited as Zonoptilus ; characters; Austria; Hungary; southern Russia).

— Bernhauer, 1908e: 329, 330 ( Coprophilus ; subgenus Zonyptilus , cited as Zonoptilus ; characters; catalog; Austria; Germany; Hungary; southern Russia).

— Bernhauer and Schubert, 1911: 89 ( Coprophilus ; catalog).

— Scheerpeltz, 1933: 1076 ( Coprophilus ; catalog).

— Horion, 1963: 161 ( Coprophilus ; subgenus Zonyptilus , cited as Zonoptilus ; Russia; Bosnia Hercegovina; Hungary; Slovakia; Czech Republic; Austria; Germany).

— Lohse, 1964: 72 ( Coprophilus ; subgenus Zonyptilus , cited as Zonoptilus ; characters; central Europe).

— Smetana, 1964c: 164 ( Coprophilus ; Czechoslovakia).

— Herman, 1970: 368 ( Coprophilus ).

— Ádám, 1987: 138 ( Zonyptilus ; collecting notes; Hungary).

— Lucht, 1987: 91 ( Coprophilus ; checklist; Germany; Czechoslovakia; Austria).

— Tóth, 1988: 199, 202 ( Coprophilus ; characters; Carpathian Basin).

— Boháč, 1993: 42 ( Elonium ; subgenus Zonyptilus ; Slovakia).

— Assing, Frisch, Kahlen, et al., 1998: 127 ( Coprophilus ; Germany).

- DISTRIBUTION: Europe, Russia.

reitteri Luze, 1904a: 69 ( Coprophilus ; subgenus Zonyptilus , cited as Zonoptilus ; Type locality: Aulie-Ata).

— Bernhauer, 1908e: 330 ( Coprophilus ; synonym of schubertii ).

— Bernhauer and Schubert, 1911: 90 ( Coprophilus ; synonym of schubertii ).

— Tóth, 1988: 200 ( Coprophilus ; subgenus Zonyptilus , cited as Zonoptilus ; valid species; characters).

- DISTRIBUTION: Kazakhstan.

rufipennis Reitter, 1894: 178 ( Zonyptilus , cited as Zonoptilus ; Type locality: Margelan).

— Reitter, 1894a: 292 ( Zonyptilus , cited as Zonoptilus ; characters; Margelan; Alai; Chodshent; Persien).

— Bernhauer, 1908e: 329, 330 ( Coprophilus ; subgenus Zonyptilus , cited as Zonoptilus ; characters; catalog; Margelan; Turcmenistan).

— Bernhauer and Schubert, 1911: 90 ( Coprophilus ; catalog).

— Scheerpeltz, 1933: 1076 ( Coprophilus ; catalog).

— Herman, 1970: 368 ( Coprophilus ).

- DISTRIBUTION: Uzbekistan, Turkmenistan, Iran.

rufitarsis Kashcheev, 1999: 149 ( Coprophilus ; subgenus Zonyptilus , cited as Zonoptilus ;

Type locality: Kazakhstan, zapovednik Aksu-Dzhabagly).

- DISTRIBUTION: Kazakhstan.

schubertii Motschulsky, 1860b: 120 ( Zonyptilus , cited as Zonoptilus ; Type locality: Ostrogorst, G Voron [sko?, esh?]; [Note: See lectotype designation by Tóth, 1988]).

— Fauvel, 1875a:VIII [= 1875b: 210] ( Coprophilus ; synonym of piceus ).

— Fauvel, 1897a: 226, 227, 228 ( Coprophilus ; characters; Russia).

— Bernhauer, 1908e: 329, 330 ( Coprophilus ; subgenus Zonyptilus , cited as Zonoptilus ; characters; catalog; Turkestan; southern Russia).

— Bernhauer and Schubert, 1911: 90 ( Coprophilus ; catalog).

— Scheerpeltz, 1933: 1076 ( Coprophilus ; catalog).

— Herman, 1970: 368 ( Coprophilus ).

— Tóth, 1988: 198 ( Coprophilus ; subgenus Zonyptilus , cited as Zonoptilus ; lectotype designation: Ostrogorst, G Voron [sko?, esh?]; characters; notes).

- DISTRIBUTION: Russia, “Turkestan”.

sellula Kraatz, 1857 , see: pennifer Motschulsky, 1845 .

— Herman, 1970: 368 ( Coprophilus ).

— Campbell, 1979a: 228 ( Coprophilus ; characters).

- DISTRIBUTION: Canada, USA.

sibiricus Bernhauer, 1915 j: 263 ( Coprophilus ; Type locality: Ost-Sibirien, Schipka-Gora).

— Bernhauer, 1923d: 180 ( Coprophilus ; characters; Siberia).

— Scheerpeltz, 1933: 1076 ( Coprophilus ; catalog).

— Herman, 1970: 368 ( Coprophilus ).

— J. Li, 1993: 22 (Coprophils; China).

- DISTRIBUTION: Russia, China.

simplex Sharp, 1889: 415 ( Coprophilus ; Type locality: Wada-toge).

— Bernhauer and Schubert, 1911: 90 ( Coprophilus ; catalog).

— Watanabe and Shibata, 1961a: 43 ( Elonium ; characters; notes; Japan).

— Herman, 1970: 368 ( Coprophilus ).

- DISTRIBUTION: Japan.

solskyi Bernhauer, 1908e: 330 ( Coprophilus ; subgenus Zonyptilus , cited as Zonoptilus ; named for Reitter’s 1894 misidentification of lateralis ; Type locality: Dobrutja).

— Reitter, 1894a: 291 ( Zonyptilus , cited as Zonoptilus ; [Note: Misidentification: Bernhauer and Schubert, 1911: 90 cited lateralis Reitter as a synonym of solskyi ; Reitter did not describe the species as new, he attributed it to Fauvel]; characters; Dobrutscha).

— Bernhauer and Schubert, 1911: 90 ( Coprophilus ; catalog).

— Scheerpeltz, 1933: 1076 ( Coprophilus ; catalog).

— Herman, 1970: 368 ( Coprophilus ).

— Tóth, 1991: 93 ( Coprophilus ; subgenus Zonyptilus , cited as Zonoptilus ; characters).

— Tóth, 1993: 376 ( Coprophilus ; characters).

— Kashcheev, 1999: 150 ( Coprophilus ; characters).

- DISTRIBUTION: Romania.

striatipennis Eppelsheim, 1894 , see: pennifer Motschulsky, 1845 .

striatulus Fabricius, 1793: 525 ( Staphylinus ; Type locality: Germania).

— Olivier, 1795:(42): 30 ( Staphylinus ; [Note: Misidentification: Bernhauer and Schubert, 1911: 90 cited rugosus Olivier as a synonym of striatulus Fabricius ; Olivier did not describe the species as new, he attributed it to Fabricius]; characters; England, France).

— Panzer, 1795: 354 ( Staphylinus ; characters; Germany).

— Fabricius, 1801: 596 ( Staphylinus ; characters; Germany).

— Turton, 1802: 512 ( Staphylinus ; characters; Germany).

— Curtis, 1829: 29 ( Elonium ; catalog; Britain).

— Stephens, 1834: 345 ( Coprophilus ; characters; England).

— Westwood, 1838a: 18 ( Coprophilus ; type species).

— Erichson, 1839a: 609 ( Coprophilus ; characters; Germany).

— Heer, 1839: 199 ( Coprophilus ; characters; Switzerland).

— Stephens, 1839: 423 ( Coprophilus ; characters; England).

— Erichson, 1840: 816 ( Coprophilus ; characters; Europe).

— Redtenbacher, 1849: 742 ( Coprophilus ; characters; Austria).

— Küster, 1854: 21 ( Coprophilus ; characters; Europe).

— Hardy, 1851: 53 ( Coprophilus ; notes; England).

— Fairmaire and Laboulbène, 1856: 620 ( Coprophilus ; characters; France).

— Redtenbacher, 1857: 237 ( Coprophilus ; characters; Austria).

— Kraatz, 1857d: 894 ( Coprophilus ; characters; Germany).

— Thomson, 1859: 45 (type species of Coprophilus ).

— Thomson, 1861: 143 ( Coprophilus ; characters; Scandinavia).

— Fauvel, 1871a: 156 [= 1872: 130] ( Coprophilus ; characters; France; Scandinavia; Britain; Germany; Austria; Italy).

— Redtenbacher, 1874: 261 ( Coprophilus ; characters; Austria).

— LeConte, 1875: 170 ( Coprophilus ; Canada).

— Mulsant and Rey, 1878: 685 ( Coprophilus ; characters; France).

— Fowler, 1888: 394 ( Coprophilus ; characters; Britain).

— Ragusa, 1892: 168 ( Coprophilus ; Sicily).

— Seidlitz, 1891: 352 ( Coprophilus ; characters; Europe).

— Seidlitz, 1891a: 376 ( Coprophilus ; characters; Europe).

— Ganglbauer, 1895: 677 ( Coprophilus ; characters; Europe).

— Fauvel, 1897a: 227, 228 ( Coprophilus ; characters; Italy; Moldavia; Canada).

— Sainte-Claire Deville, 1907: 57 ( Coprophilus ; characters; France).

— Bernhauer, 1908e: 329, 330 ( Coprophilus ; subgenus Coprophilus ; characters; catalog).

— Reitter, 1909: 176 ( Coprophilus ; characters; Germany).

— Bernhauer and Schubert, 1911: 90 ( Coprophilus ; catalog).

— Johansen, 1914: 570 ( Coprophilus ; characters; Denmark).

— Eichelbaum, 1914: 331 ( Coprophilus ; abdominal segmentation; apical segements).

— Portevin, 1929: 426 ( Coprophilus ; cited as striatuius; characters; France).

— Scheerpeltz, 1933: 1076 ( Coprophilus ; catalog).

— Leech, 1939: 261 ( Coprophilus ; characters in key).

— Paulian, 1941: 157 ( Coprophilus ; larval characters).

— Palm, 1961a: 8 ( Coprophilus ; characters; Sweden; Denmark; Norway; Finland).

— Horion, 1963: 161 ( Coprophilus ; Denmark; Sweden; Norway; Britain; Holland; Belgium; France; Switzerland; Italy; Slovenia; Croatia; Bosnia Hercegovina; Romania; Hungary; Czechoslovakia; Poland; Russia; Germany; Austria).

— Lohse, 1964: 72 ( Coprophilus ; subgenus Coprophilus ; characters; central Europe).

— Pototskaia, 1965: 879 ( Coprophilus ; larval characters).

— Pototskaia, 1967: 26 ( Elonium ; larval characters).

— Szujecki, 1968a: 713 ( Coprophilus ; Poland).

— Herman, 1970: 368 ( Coprophilus ).

— Osella and Zanetti, 1975: 88 ( Coprophilus ; collected from nest of Talpa europaea ; notes; Italy).

— Pope, 1977: 24 ( Coprophilus ; Britain).

— Campbell, 1979a: 228 ( Coprophilus ; characters).

— Dettner, 1984 ( Coprophilus ; chemistry of defensive secretions; structure of glands; discussions).

— Segers, 1986: 23 ( Coprophilus ; checklist of species; Belgium).

— Ádám, 1987: 138 ( Coprophilus ; collecting notes; Hungary).

— Lucht, 1987: 91 ( Coprophilus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Tóth, 1988: 202 ( Coprophilus ; Carpathian Basin).

— Nowosad, 1990: 53 ( Elonium ; collected from mole nest; notes; Poland).

— Thayer, 1992a: 25 (requested to place name on Official list of Specific Names in Zoology).

ICZN, 1993a (fixed as type species of Coprophilus ; name placed on Official List of Specific Names in Zoology) .

— Boháč, 1993: 42 ( Elonium ; subgenus Elonium ; Slovakia; Czech Republic).

Hoebeke, 1995 ( Coprophilus ; characters; habitat and collecting notes; presence confirmed in North America ; Ontario; Quebec; New York) .

— Terlutter, 1995: 6 ( Coprophilus ; Germany).

— Ciceroni and Zanetti, 1995: 10 ( Coprophilus ; Italy).

— Owen, 1999a: 15 ( Coprophilus ; subterranean capture; Britain).

— Owen, 2000: 251 ( Coprophilus ; pitfall trap collections from underground at roots of trees; Britain).

- DISTRIBUTION: Europe, Canada, USA.

laesus Germar, 1832: 2 ( Oxytelus ; Type locality: Harcyniae).

— Erichson, 1839a: 609 ( Coprophilus ; synonym of striatulus ).

— Heer, 1839: 199 ( Coprophilus ; synonym of striatulus ).

— Erichson, 1840: 816 ( Coprophilus ; synonym of striatulus ).

— Kraatz, 1857d: 894 ( Coprophilus ; synonym of striatulus ).

— Mulsant and Rey, 1878: 685 ( Coprophilus ; synonym of striatulus ). — Ganglbauer, 1895: 677 ( Coprophilus ; synonym of striatulus ).

— Bernhauer, 1908e: 330 ( Coprophilus ; synonym of striatulus ).

— Bernhauer and Schubert, 1911: 90 ( Coprophilus ; synonym of striatulus ).

subplagiatus Cameron, 1928c: 558 ( Coprophilus ; subgenus Zonyptilus , cited as Zonoptilus ; Type locality: Tibet: Jelap La, 13,000 feet).

— Cameron, 1930: 173 ( Coprophilus ; subgenus Zonyptilus , cited as Zonoptilus ; characters; Tibet). — Scheerpeltz, 1933: 1076 ( Coprophilus ; catalog).

— Herman, 1970: 368 ( Coprophilus ).

— Scheerpeltz, 1976: 17 ( Coprophilus ; subgenus Zonyptilus , cited as Zonoptilus ; characters).

- DISTRIBUTION: China.

vandykei Hatch, 1957 , see: Vicelva .

weiseri Bernhauer, 1927 , see: Homalotrichus .

Coprostygnus

[4 species; Australian Region]

Coprostygnus Sharp, 1886a: 380 (species included: sculptipennis ). TYPE SPECIES: Coprostygnus sculptipennis Sharp , fixed by monotypy.

— Broun, 1893: 1027 (characters; [Note: Cited as Coprostygnus nov. gen. but followed by a citation to Sharp’s description]).

— Bernhauer and Schubert, 1911: 88 (world catalog; 2 species).

— Scheerpeltz, 1933: 1075 (world catalog supplement).

— Blackwelder, 1952: 106 (type species: sculptipennis ).

— Herman, 1970: 365 (characters; distribution; notes; list of species).

— Klimaszewski, Newton, and Thayer, 1996: 150 (endemic to New Zealand).

curvipes Broun, 1921: 484 ( Coprostygnus ; Type locality: west of Mount Algidus ).

— Scheerpeltz, 1933: 1075 ( Coprostygnus ; catalog).

— Herman, 1970: 365 ( Coprostygnus ).

- DISTRIBUTION: New Zealand.

optandus Broun, 1893: 1409 ( Coprostygnus ; Type locality: Capleston, Westland).

— Fauvel, 1900c: 181 ( Coprostygnus ; New Zealand).

— Bernhauer and Schubert, 1911: 88 ( Coprostygnus ; catalog).

— Herman, 1970: 365 ( Coprostygnus ).

- DISTRIBUTION: New Zealand.

picipennis Broun, 1921: 485 ( Coprostygnus ; Type locality: Glenhope).

— Scheerpeltz, 1933: 1075 ( Coprostygnus ; catalog).

— Herman, 1970: 365 ( Coprostygnus ).

- DISTRIBUTION: New Zealand.

sculptipennis Sharp, 1886a: 380 ( Coprostygnus ; Type locality: New Zealand: Greymouth ).

— Broun, 1893: 1027 ( Coprostygnus ; [Note: Cited as “ C. sculptipennis , n.s.” but followed by a citation to Sharp’s description]; characters; New Zealand).

— Fauvel, 1900c: 181 ( Coprostygnus ; New Zealand).

— Bernhauer and Schubert, 1911: 88 ( Coprostygnus ; catalog).

— Herman, 1970: 365 ( Coprostygnus ).

- DISTRIBUTION: New Zealand.

Homalotrichus

[7 species; Neotropical and Australian Regions]

Homalotrichus Solier, 1849: 321 (species included: striatus , impressicollis , obscurus , fuscus , luteipes ). TYPE SPECIES: Homalotrichus striatus Solier , fixed by subsequent designation by Tottenham, 1939d: 229.

— Lacordaire, 1854: 153 (characters; notes; list of species).

— Fairmaire and Germain, 1861: 452 (characters).

— Fauvel, 1867a: 39 [= 1868: 44] (cited as synonym of Coprophilus ; notes; characters; key to Chilean species).

— Gemminger and Harold, 1868: 656 (synonym of Coprophilus ).

— Bernhauer and Schubert, 1911: 89 (synonym of Coprophilus ).

— Tottenham, 1939d: 229 (type species designation: striatus ).

— Blackwelder, 1944: 102 (cited as Coprophilus ; checklist of species from Mexico, Central and South America, and the West Indies).

— Blackwelder, 1952: 190 (type species).

— Coiffait and Saiz, 1968: 415 (characters; key to Chilean species).

— Herman, 1970: 368 (characters; distribution; notes; list of species).

Sharpia Fauvel, 1878 g: 488 (species included; banksi ; preoccupied by Tournier, 1874, and Bocage, 1878). TYPE SPECIES: Sharpia banksi Fauvel , fixed by monotypy.

— Duvivier, 1883: 193 (catalog).

— Blackwelder, 1952: 351 (type species).

— Herman, 1970: 368 (synonym of Homalotrichus ).

Toxoderus Fauvel, 1900c: 189 (replacement name for Sharpia Fauvel ). TYPE SPECIES: Sharpia banksi Fauvel , fixed by objective synonymy with Sharpia .

— Bernhauer and Schubert, 1911: 91 (world catalog; 1 species).

— Blackwelder, 1952: 391 (type species).

— Herman, 1970: 368 (synonym of Homalotrichus ).

banksi Fauvel, 1878 g: 488 ( Sharpia ; Type locality: Australie méridionale, Victoria).

— Bernhauer and Schubert, 1911: 91 ( Toxoderus ; catalog).

— Herman, 1970: 369 ( Homalotrichus ).

- DISTRIBUTION: Australia.

fuscus Solier, 1849 , see: Thinodromus .

impressicollis Solier, 1849: 323 ( Homalotrichus ; Type locality: Chile: San Carlos ).

— Kraatz, 1859a: 6 ( Homalotrichus ; notes).

— Fairmaire and Germain, 1861: 452 ( Homalotrichus ; characters; Chile).

— Fauvel, 1864a: 129 ( Homalotrichus ; notes; Chile).

— Fauvel, 1867a: 41 [= 1868: 46] ( Coprophilus ; characters; Chile).

— Bernhauer and Schubert, 1911: 89 ( Coprophilus ; catalog).

— Coiffait and Saiz, 1968: 419 ( Homalotrichus ; characters; Chile).

— Saiz, 1969a: 10 ( Homalotrichus ; notes).

— Herman, 1970: 369 ( Homalotrichus ).

— Scheerpeltz, 1972: 9 ( Coprophilus ; Argentina).

- DISTRIBUTION: Chile, Argentina.

luteipes Solier, 1849 , see: Thinodromus .

nahuelbutensis Coiffait and Saiz, 1968: 418 ( Homalotrichus ; Type locality: Chili: Nahuelbuta , Malleco).

- DISTRIBUTION: Chile.

obscurus Solier, 1849 , see: Carpelimus .

parvipennis Scheerpeltz, 1972: 64 ( Coprophilus ; subgenus Coprophilus ; Type locality: Argentinien: Rio Negro, El Bolson, 220 m).

— Herman, 2001: 14 ( Homalotrichus ).

- DISTRIBUTION: Argentina.

striatus Solier, 1849: 323 ( Homalotrichus ; Type locality: Chile: provincia de Valdivia).

— Kraatz, 1859a: 6 ( Homalotrichus ; characters).

— Fairmaire and Germain, 1861: 452 ( Homalotrichus ; characters; Chile).

— Fauvel, 1864a: 129 ( Homalotrichus ; notes; Chile).

— Fauvel, 1867a: 42 [= 1868: 47] ( Coprophilus ; characters; Chile).

— Bernhauer and Schubert, 1911: 90 ( Coprophilus ; catalog).

— Coiffait and Saiz, 1968: 417 ( Homalotrichus ; characters; Chile).

— Saiz, 1969a: 10 ( Homalotrichus ; notes).

— Herman, 1970: 369 ( Homalotrichus ).

substriatus Kraatz, 1859a: 16 ( Homalotrichus ; Type locality: Chile).

— Fairmaire and Germain, 1861: 452 ( Homalotrichus ; characters; Chile).

— Fauvel, 1867a: 42 [= 1868: 47] ( Coprophilus ; characters; Chile).

— Bernhauer and Schubert, 1911: 90 ( Coprophilus ; catalog).

— Coiffait and Saiz, 1968: 419 ( Homalotrichus ; characters; Chile).

— Herman, 1970: 369 ( Homalotrichus ).

— Scheerpeltz, 1972: 9 ( Coprophilus ; Argentina).

— Saiz, Solervicens, and Ojeda, 1989: 52 ( Homalotrichus ; collecting notes; habitus; Chile). - DISTRIBUTION: Chile, Argentina.

weiseri Bernhauer, 1927e: 229 ( Coprophilus ; Type locality: Argentinien: Prov. Catamarca, Peñas Azules ).

— Scheerpeltz, 1933: 1076 ( Coprophilus ; catalog).

— Herman, 1970: 369 ( Homalotrichus ).

- DISTRIBUTION: Argentina.

Tribe Deleasterini

Deleasterini Reitter, 1909: 164 (genera included: Syntomium, Deleaster ). TYPE GENUS: Deleaster (see Newton and Thayer, 1992: 60).

— Portevin, 1929: 428 (characters; key to French genera).

— Scheerpeltz, 1933: 1074 (synonym of Coprophili).

— Crowson, 1955: 37 (genera).

— Hatch, 1957: 84 (characters).

— Newton and Thayer, 1992: 60 (synonym of Coprophilini ).

— M. Hansen, 1996: 37, 98 (separate from Coprophilini ; list of Danish species).

— Newton, Thayer, Ashe, and Chandler, 2000: 321, 378 (key to Nearctic genera; characters; notes).

Pholidiens Mulsant and Rey, 1877: 65 (genera included: Deinopsis , Pholidus , Micropeplus ). TYPE GENUS: Pholidus (see Newton and Thayer, 1992: 60).

— Newton and Thayer, 1992: 60 (synonym of Coprophilini ).

Pholidini Acloque, 1896: 145 (genera included: Pholidus ). TYPE GENUS: Pholidus (see Newton and Thayer, 1992: 60).

— Newton and Thayer, 1992: 60 (synonym of Coprophilini ).

Euphaniae Reitter, 1909: 16 (genera included: Euphanias , Pholidus ; [Note: Cited on page 199 as Pholidinae]). TYPE GENUS: Euphanias (see Newton and Thayer, 1992: 60).

— Newton and Thayer, 1992: 60 (synonym of Coprophilini ).

Syntomiinae Böving and Craighead, 1931: 28 (genera included: Syntomium ; larval characters). TYPE GENUS: Syntomium (see Newton and Thayer, 1992: 60).

— Bernet Kempers, 1941: 34, 36 (larval characters).

— Crowson, 1955: 37 (genera).

— Newton and Thayer, 1992: 60 (synonym of Coprophilini ).

Syntomiini :

— M. Hansen, 1996: 37, 98 (separate from Coprophilini ; list of Danish species).

Deleasterini Hatch, 1957: 84 , 86 (genera included: Syntomium, Deleaster , Zalobius , Asemobius ). TYPE GENUS: Deleaster (see Newton and Thayer, 1992: 60).

— Newton and Thayer, 1992: 60 (synonym of Coprophilini ).

Deleaster

[11 species (1 doubtful); Nearctic, Palaearctic, and Ethiopian Regions]

Deleaster Erichson, 1839a: 610 (species included: dichrous ). TYPE SPECIES: Anthophagus dichroa Gravenhorst , fixed by monotypy.

— Heer, 1839: 198 (characters).

— Stephens, 1839: 422 (characters).

— Redtenbacher, 1849: 743 (characters).

— Schaum, 1852: 29 (list of species; Europe).

— Lacordaire, 1854: 122 (characters; notes; list of species).

— Fairmaire and Laboulbène, 1856: 621 (characters).

— Gistel, 1856: 389 (list of species).

— Jacquelin du Val, 1857: 60 (characters).

— Redtenbacher, 1857: 238 (characters; Austrian species).

— Kraatz, 1857d: 901 (characters).

— G. Waterhouse, 1858: 30 (catalog of British species).

— Schaum, 1859: 31 (catalog; European species).

— Gredler, 1863: 119 (list of species from Tirol).

— Wencker and Silbermann, 1866: 35 (list of species; collecting notes; France).

— Gemminger and Harold, 1868: 656 (catalog).

— Jacquelin du Val, 1868: 78 (catalog; European species).

— Sharp, 1871d: 14 (list of British species).

— Fauvel, 1871a: 151 [= 1872: 125] (characters).

— Redtenbacher, 1874: 262 (characters).

— Fauvel, 1875a:VIII [= 1875b: 210] (catalog).

— Mulsant and Rey, 1878: 818 (characters).

— Duvivier, 1883: 193 (catalog).

— Fowler, 1888: 396 (notes).

— Heyden, Reitter, and Weise, 1891: 117 (list of species of Europe and Caucasus).

— Ganglbauer, 1895: 677 (characters).

— Heyden, Reitter, and Weise, 1906: 137 (list of species of Europe and Caucasus).

— Reitter, 1909: 177 (characters).

— Bernhauer and Schubert, 1911: 88 (world catalog; 5 species).

— Petri, 1912: 56 (list of species of Siebenbürgen).

— Winkler, 1925: 338 (catalog for Palaearctic region).

— Porta, 1926: 35 (characters; species of Italy).

— Roubal, 1930: 321 (catalog; Slovakia).

— Joy, 1932: 144 (key to British species).

— Scheerpeltz, 1933: 1075 (world catalog supplement).

— Jeannel and Jarrige, 1949: 324 (cave species; type species: dichrous ).

— Kloet and Hincks, 1945: 162 (list of British species).

— Tottenham, 1949: 359, 408 (type species: dichrous Gravenhorst ; list of British species). — Blackwelder, 1952: 118 (type species: dichroa ).

— Tottenham, 1954: 38 (characters; British species).

— Hatch, 1957: 84, 87 (characters; species of Pacific Northwest).

— Kocher, 1958: 95 (checklist of species; Morocco).

— Palm, 1961a: 7 (characters; species of Sweden).

— Horion, 1963: 159 (list of central European species).

— Székessy, 1963: 27 (characters).

— Lohse, 1964: 72 (characters; key to central European species).

— Moore, 1964: 57 (characters).

— Scheerpeltz, 1968a: 20 (catalog of Austrian species).

— Herman, 1970: 363 (characters; distribution; notes; list of species).

— Tikhomirova, 1973a: 142 (checklist of species of USSR).

— Moore and Legner, 1975: 243 (catalog; North America north of Mexico).

— Shibata, 1976: 130 (checklist of species of Japan).

— Pope, 1977: 24 (list of British species).

— Topp, 1978: 310 (larval characters).

— Burakowski, Mroczkowski, and Stefańska, 1979: 80 (catalog; Poland).

— Moore and Legner, 1979: 222 (characters; notes).

— Muona, 1979: 19 (list of Scandinavian species).

— Shibata, 1986a: 118 (supplemental list of species from Taiwan).

— Campbell and Davies, 1991: 92 (checklist of species; Canada).

— Silfverberg, 1992: 23 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Ciceroni and Zanetti, 1995: 10 (list of species of Italy).

— M. Hansen, 1996: 98 (list of Danish species).

— Downie and Arnett, 1996: 441 (characters; species of Northeastern North America).

— Schülke, 1998i: 125 (characters).

— Newton, Thayer, Ashe, and Chandler, 2000: 378 (3 Nearctic species; notes; characters in key). adustus Küster, 1846 , see: dichrous Gravenhorst, 1802 .

bactrianus Semenow, 1900: 684 ( Deleaster ; Type locality: Bucharia orient.: prov. Hissar:

cursus superior fluvii Karatag).

— Bernhauer and Schubert, 1911: 88 ( Deleaster ; catalog).

— Scheerpeltz, 1963: 5 ( Deleaster ; Afghanistan).

— Herman, 1970: 364 ( Deleaster ).

— Coiffait, 1979d: 554 ( Deleaster ; Afghanistan).

— Kashcheev, 1999: 149 ( Deleaster ; characters).

- DISTRIBUTION: Uzbekistan, Afghanistan.

bergi Kashcheev, 1999: 148 ( Deleaster ; Type locality: okr. g. Almaty).

- DISTRIBUTION: Kazakhstan.

brassicae Scopoli, 1763 , see: dichrous Gravenhorst, 1802 .

concolor LeConte, 1866: 375 ( Deleaster ; Type locality: San Francisco , California).

— Fauvel, 1878d: 256 [= 1878e: 92] ( Deleaster ; characters; California).

— Fall and Cockerell, 1907: 168 ( Deleaster ; New Mexico).

— Bernhauer and Schubert, 1911: 88 ( Deleaster ; catalog).

— Hatch, 1957: 87 ( Deleaster ; characters; British Columbia; Washington; Oregon).

— Herman, 1970: 364 ( Deleaster ).

- DISTRIBUTION: Canada, USA.

dichrous Gravenhorst, 1802: 188 ( Anthophagus ; Type locality: Dresdae).

— Latreille, 1804: 369 ( Lesteva ; characters).

— Gravenhorst, 1806: 221 ( Anthophagus ; cited as dichrus).

— Mannerheim, 1830: 55 ( Lesteva ; Germany).

— Mannerheim, 1831: 469 ( Lesteva ; Germany).

— Ménétriés, 1832: 147 ( Lesteva ; Caucasus-Persia region).

— Stephens, 1834: 359 ( Lesteva ; characters; England; Scotland).

— Runde, 1835: 23 ( Lesteva ; characters; Germany).

— Lacordaire, 1835: 479 ( Lesteva ; characters; France).

— Guérin-Méneville, 1829: pl. 9, fig. 10 ( Lesteva ; illustration).

— Erichson, 1839a: 611 ( Deleaster ; characters; Germany).

— Heer, 1839: 198 ( Deleaster ; characters; Switzerland).

— Stephens, 1839: 431 ( Deleaster ; characters; England).

— Erichson, 1840: 819 ( Deleaster ; characters; Germany; France; England).

— Dufour, 1843: 37 ( Anthophagus ; characters; France).

— Kiesenwetter, 1844a: 376 ( Deleaster ; notes; Germany).

— Guérin-Méneville, 1844a: 32 ( Lesteva ; Europe).

— Küster, 1846: 47 ( Deleaster ; characters; Germany; Italy; Switzerland; France; England).

— Hochhuth, 1849: 200 ( Deleaster ; characters).

— Redtenbacher, 1849: 743 ( Deleaster ; characters; Austria).

— Kiesenwetter, 1851a: 433 ( Deleaster ; France).

— Fairmaire and Laboulbène, 1856: 621 ( Deleaster ; characters; France).

— Redtenbacher, 1857: 238 ( Deleaster ; characters; Austria).

— Kraatz, 1857d: 902 ( Deleaster ; characters; Germany).

— Hochhuth, 1862: 101 ( Deleaster ; notes; Russia).

— Fauvel, 1865a: 314 [= 1865b: 64] ( Deleaster ; habitat; France).

— Hochhuth, 1872: 168 ( Deleaster ; notes; Ukraine).

— Redtenbacher, 1874: 263 ( Deleaster ; characters; Austria).

— Seidlitz, 1875: 243 ( Deleaster ; characters; Baltic region).

— Mulsant and Rey, 1878: 820 ( Deleaster ; characters; France).

— Fauvel, 1886: 16 [= 1886a: 8] ( Deleaster ; Morocco).

— Fowler, 1888: 396 ( Deleaster ; characters; Britain).

— Seidlitz, 1891: 349 ( Deleaster ; characters; Europe).

— Seidlitz, 1891a: 373 ( Deleaster ; characters; Europe).

— Ganglbauer, 1895: 678 ( Deleaster ; characters; middle Europe; eastern Mediterranean region).

— Fauvel, 1897d: 249 ( Deleaster ; Morocco).

— Fauvel, 1902b: 57 ( Deleaster ; Algeria; Tunisia).

— Sainte-Claire Deville, 1906: 72 ( Deleaster ; Corsica).

— Sainte-Claire Deville, 1907: 56 ( Deleaster ; characters; France).

— Reitter, 1909: 178 ( Deleaster ; characters; Germany).

— Bernhauer and Schubert, 1911: 88 ( Deleaster ; catalog).

— Portevin, 1929: 428 ( Deleaster ; characters; France).

— Bernhauer, 1931e: 9 ( Deleaster ; Morocco).

— Scheerpeltz, 1933: 1075 ( Deleaster ; catalog).

— Olsson, 1934: 67 ( Deleaster ; Sweden).

— Chagnon, 1934 ( Deleaster ; characters; Canada).

— Jules, 1935 ( Deleaster ; characters; Canada).

— C. Koch, 1941: 54 ( Deleaster ; Morocco).

— MacNay, 1946: 213 ( Deleaster ; Ontario).

— Tottenham, 1949: 408 ( Deleaster ; checklist; Britain).

— Jeannel and Jarrige, 1949: 324 ( Deleaster ; notes; cave species; Spain; Serbia; Romania).

— Palm, 1953: 9 ( Deleaster ; ecological notes; Sweden).

— Kocher, 1958: 95 ( Deleaster ; Morocco).

— Palm, 1961a: 7 ( Deleaster ; characters; Sweden; Denmark; Finland).

— Horion, 1963: 159 ( Deleaster ; Morocco; Iran; Britain; Holland; Belgium; France; Switzerland; Spain; Corsica; Sardinia; Sicily; Italy; Croatia; Bosnia Hercegovina; Albania; Macedonia; Greece; Romania; Hungary; Czechoslovakia; Poland; Russia; Denmark; Sweden; Finland; Germany; Austria).

— Lohse, 1964: 72 ( Deleaster ; characters; central Europe).

— Smetana, 1967e: 307 ( Deleaster ; Turkey).

— Szujecki, 1968a: 712 ( Deleaster ; Poland).

— Scheerpeltz, 1968a: 20 ( Deleaster ; catalog; Austria).

— Bolov, 1969: 513 ( Deleaster ; collecting notes; Russia).

— Bolov, 1969a: 328 ( Deleaster ; collecting notes; Russia).

— Herman, 1970: 365 ( Deleaster ).

— Pope, 1977: 24 ( Deleaster ; Britain).

— Bordoni, 1978a: 55 ( Deleaster ; cave collection; Turkey).

— Muona, 1979: 19 ( Deleaster ; Finland; Russia; Sweden; Denmark).

— Burakowski, Mroczkowski, and Stefańska, 1979: 80 ( Deleaster ; catalog; Poland).

— Frank, 1982: 13 ( Deleaster ; list of parasites).

— Dettner, Schwinger and Wunderle, 1985 ( Deleaster ; defensive gland morphology, secretions, function).

— Segers, 1986: 23 ( Deleaster ; checklist of species; Belgium).

— Lucht, 1987: 91 ( Deleaster ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— M. Hansen, Kristensen, Mahler, and Pedersen, 1991: 106 ( Deleaster ; Denmark).

— Silfverberg, 1992: 23 ( Deleaster ; Finland; Russia; Sweden; Norway; Denmark; Estonia; Lithuania).

— Gusarov, 1992a: 776 [= 1993a: 63] ( Deleaster ; synonymic notes).

— Boháč, 1993: 42 ( Deleaster ; Czech Republic; Slovakia).

— Dearborn and Donahue, 1994: 14 ( Deleaster ; Maine).

— Terlutter, 1995: 5 ( Deleaster ; Germany).

— Sparacio, 1995: 144 ( Deleaster ; characters; summary of distribution; notes; Sicily).

— Ciceroni and Zanetti, 1995: 10 ( Deleaster ; Italy).

— Downie and Arnett, 1996: 441 ( Deleaster ; characters; Canada).

— Franzen, 1996: 206 ( Deleaster ; collecting notes; Germany).

— M. Hansen, 1996: 98 ( Deleaster ; checklist; Denmark).

— Sterrenburg and Schülke, 1997: 16 ( Deleaster ; Netherlands).

— Czarniawski and Staniec, 1997: 38 ( Deleaster ; structure of opening of abdominal defensive reservoirs).

— Wenzel, 1997: 345 ( Deleaster ; collecting notes; Germany).

— Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 45 ( Deleaster ; Latvia).

— Anderson, Nash, and O’Connor, 1997: 16 ( Deleaster ; Ireland).

— Hansen, Jørum, Palm, and Pedersen, 1997: 125 ( Deleaster ; checklist; Denmark).

— Outerelo, Gamarra, and Salgado, 1998: 124 ( Deleaster ; collected from cave; Spain).

— Kashcheev, 1999: 149 ( Deleaster ; characters).

— Eyre, Luff, and Lott, 2000: 33 ( Deleaster ; Britain).

— Herman, 2001: 23 ( Deleaster ; nomen protectum).

- DISTRIBUTION: Morocco, Algeria, Tunisia, Europe, Russia, Armenia, Turkey, Iran, USA, Canada.

brassicae Scopoli, 1763: 102 ( Staphylinus ; [Note: This name is older than dichrous . Bernhauer and Schubert (1911: 88) and other authors cited Schrank, 1798 as the author of brassicae but Schrank attributed the species to Scopoli]; Type locality: Carniola).

— Goeze, 1777: 725 ( Staphylinus ; characters).

— Villers, 1789: 424 ( Staphylinus ; characters).

— Gmelin, 1790: 2032 ( Staphylinus ; page misnumbered as 1032; characters).

— Schrank, 1798: 644 ( Staphylinus ; characters; notes).

— Turton, 1802: 516 ( Staphylinus ; characters; Carniola).

— Erichson, 1840: 819 ( Deleaster ; synonym of dichrous ).

— Redtenbacher, 1849: 743 ( Deleaster ; synonym of dichrous ).

— Kraatz, 1857d: 902 ( Deleaster ; synonym of dichrous ).

— Ganglbauer, 1895: 678 ( Deleaster ; synonym of dichrous ).

— Bernhauer and Schubert, 1911: 88 ( Deleaster ; synonym of dichrous ).

— Herman, 2001: 23 ( Deleaster ; nomen oblitum).

leachii Curtis, 1830 : pl. 303 ( Lesteva ; Type locality: England: Cumberland , near Holme Gate, banks of river Eden).

— Stephens, 1834: 359 ( Lesteva ; synonym of dichrous Gravenhorst ).

— Erichson, 1839a: 611 ( Deleaster ; synonym of dichrous ).

— Stephens, 1839: 431 ( Deleaster ; synonym of dichroa ).

— Erichson, 1840: 819 ( Deleaster ; synonym of dichrous ).

— Kraatz, 1857d: 902 ( Deleaster ; synonym of dichrous ).

— Fauvel, 1871a: 152 [= 1872: 126] ( Deleaster ; synonym of dichrous ).

— Mulsant and Rey, 1878: 821 ( Deleaster ; synonym of dichrous ).

— Fowler, 1888: 397 ( Deleaster ; variety of dichrous ; characters; Britain).

— Ganglbauer, 1895: 678 ( Deleaster ; variety of dichrous ).

— Bernhauer and Schubert, 1911: 88 ( Deleaster ; variety of dichrous ).

— Scheerpeltz, 1933: 1075 ( Deleaster ; aberration of dichrous ).

— Smetana, 1967e: 307 ( Deleaster ; aberration of dichrous ; Turkey).

adustus Küster, 1846: 48 ( Deleaster ; Type locality: Rothenthurmpass in Siebenbürgen ).

— Redtenbacher, 1849: 743 ( Syntomium ; characters).

— Kraatz, 1857d: 902 ( Deleaster ; synonym of dichrous ).

— Fauvel, 1869: 493 ( Deleaster ; synonym of dichrous ).

— Fauvel, 1871a: 152 [= 1872: 126] ( Deleaster ; synonym of dichrous ).

— Ganglbauer, 1895: 678 ( Deleaster ; synonym of dichrous ).

— Bernhauer and Schubert, 1911: 88 ( Deleaster ; synonym of dichrous ).

erichsoni Hochhuth, 1851: 57 ( Deleaster ; Type locality: Gebirge Armeniens; [Note: See lectotype designation by Gusarov, 1992a]).

— Fauvel, 1871a: 152 [= 1872: 126] ( Deleaster ; synonym of dichrous ).

— Ganglbauer, 1895: 678 ( Deleaster ; synonym of dichrous ).

— Bernhauer and Schubert, 1911: 88 ( Deleaster ; variety of dichrous ).

— Scheerpeltz, 1933: 1075 ( Deleaster ; variety of dichrous ).

— Gusarov, 1992a: 776 [= 1993a: 63] ( Deleaster ; synonym of dichrous ; lectotype designation). testaceicolor Roubal, 1941: 10 ( Deleaster ; aberration of dichrous ; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Ross. subcarp.: Rachov-Mencul; Slov.: Trencin). dryophilus Gistel, 1857: 11 ( Deleaster ; [Note: Nomen dubium]; Type locality: Styriae).

- DISTRIBUTION: Austria.

erichsoni Hochhuth, 1851 , see: dichrous Gravenhorst, 1802 .

grandiceps † Wickham, 1912: 12 ( Deleaster ; Miocene fossil; Type locality: Florissant , Colorado) .

- DISTRIBUTION: USA.

leachii Curtis, 1830 , see: dichrous Gravenhorst, 1802 .

pectinatus Fauvel, 1882b: 129 ( Deleaster ; Type locality: Abyssinie).

— Bernhauer and Schubert, 1911: 88 ( Deleaster ; catalog).

— Herman, 1970: 365 ( Deleaster ).

- DISTRIBUTION: Ethiopia.

pekinensis Fairmaire, 1893a : ccxii ( Deleaster ; Type locality: Peking).

— Bernhauer and Schubert, 1911: 88 ( Deleaster ; catalog).

— Herman, 1970: 365 ( Deleaster ).

— J. Li, 1993: 23 ( Deleaster ; China).

- DISTRIBUTION: China.

taiwanensis Hayashi, 1984: 91 ( Deleaster ; Type locality: Mt. Lishan , alt. about 2500 m, Nantou Hsien, Taiwan).

- DISTRIBUTION: Taiwan.

testaceicolor Roubal, 1941 [Note: Nomen nudum], see: dichrous Gravenhorst, 1802 . trimaculatus Fall, 1910: 115 ( Deleaster ; Type locality: Cañon City, Colorado).

— Scheerpeltz, 1933: 1075 ( Deleaster ; catalog).

— Herman, 1970: 365 ( Deleaster ).

— Moore, 1975d: 174 ( Deleaster ; Arizona; California).

- DISTRIBUTION: USA.

yokoyamai Adachi, 1935: 126 ( Deleaster ; Type locality: Shimajima Valley , Shinano Province).

— Nakane, 1963a: 83 ( Deleaster ; characters; habitus photograph; Japan).

— Herman, 1970: 365 ( Deleaster ).

— Frank, 1982: 13 ( Deleaster ; list of parasites).

- DISTRIBUTION: Japan.

Euphanias

[5 species; Neotropical and Palaearctic Regions]

Euphanias Fairmaire and Laboulbène, 1856: 657 (species included: insignicornis ). TYPE SPECIES: Euphanias insignicornis Fairmaire and Laboulbène , fixed by monotypy. — Kraatz, 1857a: 51 (characters).

— Redtenbacher, 1857: 991 (characters).

— Jacquelin du Val, 1857: 65 (characters).

— Kraatz, 1857d: 806 (characters; notes).

— Schaum, 1859: 31 (catalog; European species).

— Jacquelin du Val, 1868: 79 (catalog; European species).

— Fauvel, 1869: 493 (synonym of Pholidus ).

— Fauvel, 1871a: 158 [= 1872: 132] (synonym of Pholidus ).

— Redtenbacher, 1874: 260 (characters).

— Fauvel, 1875a:VIII [= 1875b: 210] (synonym of Pholidus ).

— Ganglbauer, 1895: 689 (synonym of Pholidus ).

— Bernhauer and Schubert, 1910: 8 (world catalog; Piestini ).

— Winkler, 1925: 323 (catalog for Palaearctic region).

— Scheerpeltz, 1933: 994 (world catalog supplement).

— Blackwelder, 1944: 101 (checklist of species from Mexico, Central and South America, and the West Indies).

— Blackwelder, 1952: 157 (type species: insignicornis ).

— Tikhomirova, 1973a: 142 (checklist of species of USSR).

— Blinshtein, 1978: 80 (key to species of Mediterranean region).

— Newton, 1982a: 6, 8 (characters; transferred to Oxytelinae ).

— Ciceroni and Zanetti, 1995: 10 (list of species of Italy).

Pholidus Mulsant and Rey, 1856a: 7 [= 1856b: 7] (species included: insignis ; preoccupied by Rafinesque, 1815, and Gray, 1840). TYPE SPECIES: Pholidus insignis Mulsant and Rey , fixed by monotypy.

— Redtenbacher, 1857: 991 (synonym of Euphanias ).

— Schaum, 1859: 31 (synonym of Euphanias ).

— Gemminger and Harold, 1868: 655 (synonym of Euphanias ).

— Fauvel, 1871a: 158 [= 1872: 132] (characters).

— Fauvel, 1875a:VIII [= 1875b: 210] (catalog).

— Mulsant and Rey, 1880: 408 (characters).

— Heyden, Reitter, and Weise, 1891: 117 (list of species of Europe and Caucasus).

— Ganglbauer, 1895: 689 (characters).

— Heyden, Reitter, and Weise, 1906: 125 (list of species of Europe and Caucasus).

— Bernhauer and Schubert, 1910: 8 (synonym of Euphanias ).

— Porta, 1926: 6 (characters; species of Italy).

— Scheerpeltz, 1933: 994 (synonym of Euphanias ).

— Blackwelder, 1952: 306 (type species: insignis ).

boiteli Normand, 1936: 163 ( Euphanias ; Type locality: Tunisie: Bords du lac de Bizerte).

— Blinshtein, 1978: 80 ( Euphanias ; characters).

- DISTRIBUTION: Tunisia.

ganglbaueri Bernhauer, 1908d: 286 ( Pholidus ; Type locality: Brasilien: Blumenau ).

— Bernhauer and Schubert, 1910: 8 ( Euphanias ; catalog).

- DISTRIBUTION: Brazil.

insignicornis Fairmaire and Laboulbène, 1856 , see: insignis Mulsant and Rey, 1856 .

insignis Mulsant and Rey, 1856a: 9 [= 1856b: 9] ( Pholidus ; Type locality: Hyères; [Note:

See lectotype designation by Gusarov, 1997]).

— Fauvel, 1871a: 158 [= 1872: 132] ( Pholidus ; characters; France).

— Mulsant and Rey, 1880: 409 ( Pholidus ; characters; France).

— Ganglbauer, 1895: 690 ( Pholidus ; characters; France).

— Fauvel, 1897d: 250 ( Pholidus ; Algeria).

— Fauvel, 1902b: 58 ( Pholidus ; Algeria; Tunisia; Egypt).

— Bernhauer and Schubert, 1910: 8 ( Euphanias ; catalog).

— Portevin, 1929: 458 ( Pholidus ; characters; France).

— Scheerpeltz, 1933: 994 ( Euphanias ; catalog).

— C. Koch, 1934: 33 ( Euphanias ; characters; Egypt).

— C. Koch, 1936b: 126 ( Euphanias ; Egypt).

— Blinshtein, 1978: 80 ( Euphanias ; characters).

— Haghebaert, 1989a: 277 ( Euphanias ; notes; Camargue).

— Sparacio, 1995: 144 ( Euphanias ; characters; notes; Sicily).

— Adorno, Ciceroni, and Zanetti, 1996: 51 ( Euphanias ; notes; Sicily).

Gusarov, 1997: 278 ( Euphanias ; lectotype designation; Saudi Arabia) .

- DISTRIBUTION: Algeria, Tunisia, Egypt, Saudi Arabia, Syria, France.

insignicornis Fairmaire and Laboulbène, 1856: 657 ( Euphanias ; Type locality: Hyères).

— Redtenbacher, 1857: 991 ( Euphanias ; characters; Austria).

— Fauvel, 1869: 493 ( Euphanias ; synonym of insignis ).

— Fauvel, 1871a: 158 [= 1872: 132] ( Pholidus ; synonym of insignis ).

— Redtenbacher, 1874: 260 ( Euphanias ; characters).

— Mulsant and Rey, 1880: 410 ( Pholidus ; synonym of insignis ).

— Ganglbauer, 1895: 690 ( Pholidus ; synonym of insignis ).

— Bernhauer and Schubert, 1910: 8 ( Euphanias ; synonym of insignis ).

— Scheerpeltz, 1933: 994 ( Euphanias ; synonym of insignis ).

insularis C. Koch, 1934: 33 ( Euphanias ; subspecies of insignis ; Type locality: Sardinien: Cagliari).

— Ciceroni and Zanetti, 1995: 10 ( Euphanias ; subspecies of insignis ; Italy).

insularis C. Koch, 1934 , see: insignis Mulsant and Rey, 1856 .

pliginskii Bernhauer, 1912: 170 ( Euphanias ; Type locality: Sebastopol in der Krim).

— Scheerpeltz, 1933: 995 ( Euphanias ; catalog).

— Blinshtein, 1978: 80 ( Euphanias ; characters).

- DISTRIBUTION: Ukraine.

pusanovi Blinshtein, 1976: 78 ( Euphanias ; Type locality: Cheromorskii zapovednik, Potievskii uchastok).

- DISTRIBUTION: Russia.

Mitosynum

[1 species; Nearctic Region]

Mitosynum Campbell, 1982a: 687 (species included: vockerothi ; habitus). TYPE SPECIES:

Mitosynum vockerothi Campbell , fixed by original designation and monotypy.

— Downie and Arnett, 1996: 441 (characters; species of Northeastern North America).

— Newton, Thayer, Ashe, and Chandler, 2000: 379 (1 Nearctic species; notes; characters in key). vockerothi Campbell, 1982a: 690 ( Mitosynum ; Type locality: New Brunswick: Kouchi-

bouguac N. P.).

— Campbell and Davies, 1991: 93 ( Mitosynum ; checklist of species; Canada).

— Downie and Arnett, 1996: 441 ( Mitosynum ; characters; USA).

- DISTRIBUTION: Canada.

Oxypius

[1 species; Australian Region]

Oxypius Newton, 1982a: 2 (species included: peckorum ; systematic position). TYPE SPECIES: Oxypius peckorum Newton , fixed by original designation. peckorum Newton, 1982a: 10 ( Oxypius ; Type locality: Australia: Western Australia: Walpole N.P., Tingle Tree; larval description also included).

- DISTRIBUTION: Australia.

Syntomium

[8 species; Nearctic and Palaearctic Regions]

Syntomium Curtis, 1828 : pl. 228 (species included: nigroaeneum ). TYPE SPECIES: Syntomium nigroaeneum Curtis , fixed by original designation and monotypy.

— Stephens, 1829a: 24 (catalog; Britain).

— Stephens, 1834: 329 (characters; habitat).

— Westwood, 1838a: 18 (characters; type species).

— Heer, 1839: 199 (characters).

— Erichson, 1840: 821 (characters).

— Redtenbacher, 1849: 743 (characters).

— Schaum, 1852: 29 (list of species; Europe).

— Lacordaire, 1854: 123 (characters; notes; list of species).

— Fairmaire and Laboulbène, 1856: 622 (characters).

— Jacquelin du Val, 1857: 60 (characters).

— Redtenbacher, 1857: 239 (characters; Austrian species).

— Kraatz, 1857d: 890 (characters).

— Thomson, 1858: 39 (characters).

— G. Waterhouse, 1858: 30 (catalog of British species).

— Thomson, 1859: 44 (characters; type species: aeneum ).

— Schaum, 1859: 31 (catalog; European species).

— LeConte, 1861a: 69 (characters).

— Thomson, 1861: 140 (characters).

— LeConte, 1863: 26 (list of North American species).

— Wencker and Silbermann, 1866: 35 (list of species; collecting notes; France).

— Gemminger and Harold, 1868: 655 (catalog).

— Jacquelin du Val, 1868: 78 (catalog; European species).

— Fauvel, 1871a: 156 [= 1872: 130] (characters).

— Sharp, 1871d: 14 (list of British species).

— Redtenbacher, 1874: 261 (characters).

— Fauvel, 1875a:VIII [= 1875b: 210] (catalog).

— Mulsant and Rey, 1878: 823 (characters).

— Fowler, 1888: 393 (notes; larval characters).

— Heyden, Reitter, and Weise, 1891: 117 (list of species of Europe and Caucasus).

— Ganglbauer, 1895: 679 (adult and larval characters).

— Heyden, Reitter, and Weise, 1906: 137 (list of species of Europe and Caucasus).

— Reitter, 1909: 178 (characters).

— Bernhauer and Schubert, 1911: 84 (world catalog; 2 species).

— Petri, 1912: 56 (list of species of Siebenbürgen).

— Johansen, 1914: 580 (characters).

— Leng, 1920: 95 (catalog of North American species).

— Winkler, 1925: 337 (catalog for Palaearctic region).

— Porta, 1926: 35 (characters; species of Italy).

— Portevin, 1929: 394 (larval illustration).

— Roubal, 1930: 321 (catalog; Slovakia).

— Joy, 1932: 140 (key to British species).

— Scheerpeltz, 1933: 1074 (world catalog supplement).

— Kloet and Hincks, 1945: 162 (list of British species).

— Tottenham, 1949: 359, 408 (type species: aeneum P. W. J. Müller = nigroaeneum Curtis ; list of British species).

— Blackwelder, 1952: 367 (type species: nigroaeneum ).

— Hatch, 1957: 84, 86 (characters; key to species of Pacific Northwest).

— Watanabe and Shibata, 1960: 103 (list of species of world).

— Palm, 1961a: 6 (characters; species of Sweden).

— Horion, 1963: 157 (list of central European species).

— Székessy, 1963: 27 (characters).

— Lohse, 1964: 71 (characters; key to central European species).

— Moore, 1964: 57 (characters).

— Pototskaia, 1967: 24 (larval characters).

— Kasule, 1968: 118 (larval characters).

— Scheerpeltz, 1968a: 20 (catalog of Austrian species).

— Herman, 1970: 372 (characters; distribution; notes; list of species).

— Tikhomirova, 1973a: 142 (checklist of species of USSR).

— Moore and Legner, 1975: 252 (catalog; North America north of Mexico).

— Pope, 1977: 24 (list of British species).

— Topp, 1978: 309 (larval characters).

— Burakowski, Mroczkowski, and Stefańska, 1979: 80 (catalog; Poland).

— Moore and Legner, 1979: 232 (characters; notes).

— Muona, 1979: 19 (list of Scandinavian species).

— Uhlig, Vogel, and Sieber, 1980: 241 (some species collected in Germany).

— Uhlig and Vogler, 1981: 88 (list of some species collected in Germany; habitat and collecting notes).

— Muona and Viramo, 1986: 16 (list of species of northeastern Finland).

— Dettner, 1987: 31 (defensive glands).

— Campbell and Davies, 1991: 93 (checklist of species; Alaska and Canada).

— Silfverberg, 1992: 23 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Siitonen, 1993: 230 (list of Finnish species).

— M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 27 (additional records to Danish list of species).

— Ciceroni and Zanetti, 1995: 10 (list of species of Italy).

— M. Hansen, 1996: 98 (list of Danish species).

— M. Hansen, Mahler, Palm, and Pedersen, 1996: 241 (additions to list of Danish species).

— Downie and Arnett, 1996: 441 (characters; species of Northeastern North America).

— Schülke, 1998i: 125 (characters).

— Newton, Thayer, Ashe, and Chandler, 2000: 379 (4 Nearctic species; notes; characters in key).

Eunonia Casey, 1904: 313 (species included: keeniana ). TYPE SPECIES: Eunonia keeniana Casey , fixed by original designation and monotypy.

— Bernhauer and Schubert, 1910: 50 (world catalog; 1 species).

— Leng, 1920: 94 (catalog of North American species).

— Blackwelder, 1952: 157 (type species: keeniana ).

— Hatch, 1957: 51, 59 (characters; species of Pacific Northwest).

— Moore, 1966: 56 (synonym of Syntomium ).

— Herman, 1970: 372 (synonym of Syntomium ).

aeneum P. W. J. Müller, 1821: 216 ( Omalium ; Type locality: Odenbach).

— Gyllenhal, 1827: 466 ( Omalium ; characters; Sweden).

— Heer, 1839: 199 ( Syntomium ; characters; Switzerland).

— Erichson, 1840: 821 ( Syntomium ; characters; Germany; Sweden; England).

— Redtenbacher, 1849: 743 ( Syntomium ; characters; Austria).

— Küster, 1854: 23 ( Syntomium ; characters; Europe).

— Hardy, 1851: 53 ( Syntomium ; notes; England).

— Fairmaire and Laboulbène, 1856: 622 ( Syntomium ; characters; France).

— Redtenbacher, 1857: 239 ( Syntomium ; characters; Austria).

— Kraatz, 1857d: 892 ( Syntomium ; characters; Germany).

— Thomson, 1859: 44 (type species of Syntomium ).

— Thomson, 1861: 141 ( Syntomium ; characters; Scandinavia).

— Fauvel, 1871a: 157 [= 1872: 131] ( Syntomium ; characters; France; Lapland; Scandinavia; Britain; Germany; Switzerland; Austria; Italy).

— Schiødte, 1873: 559 ( Syntomium ; larval characters).

— Fauvel, 1874: 325 [= 1874b: 31] ( Syntomium ; Russia).

— Redtenbacher, 1874: 260 (Synotomium; characters; Austria).

— Seidlitz, 1875: 245 ( Syntomium ; characters; Baltic region).

— J. Sahlberg, 1876: 182 ( Syntomium ; Finland).

— Mulsant and Rey, 1878: 825 ( Syntomium ; characters; France).

— Fowler, 1888: 393 ( Syntomium ; characters; Britain).

— Seidlitz, 1891: 349 ( Syntomium ; characters; Europe).

— Seidlitz, 1891a: 373 ( Syntomium ; characters; Europe).

— Ganglbauer, 1895: 681 ( Syntomium ; characters; north and middle Europe).

— Sainte-Claire Deville, 1907: 55 ( Syntomium ; characters; France).

— Bernhauer and Schubert, 1911: 87 ( Syntomium ; catalog).

— Johansen, 1914: 581 ( Syntomium ; characters; Denmark).

— Portevin, 1929: 429 ( Syntomium ; characters; France).

— Paulian, 1941: 156 ( Syntomium ; larval characters; France).

— Jarrige, 1946: 96 ( Syntomium ; cited as aencum; natural history notes).

— Palm, 1961a: 6 ( Syntomium ; characters; Sweden; Denmark; Norway; Finland).

— Horion, 1963: 157 ( Syntomium ; Russia; Finland; Sweden; Norway; Denmark; Britain; Holland; Belgium; France; Switzerland; Italy; Croatia; Bosnia; Romania; Czechoslovakia; Poland; Germany; Austria).

— Lohse, 1964: 71 ( Syntomium ; characters; central Europe).

— Pototskaia, 1967: 24 ( Syntomium ; larval characters).

— Herman, 1970: 373 ( Syntomium ).

— Pope, 1977: 24 ( Syntomium ; Britain).

— Segers, 1986: 23 ( Syntomium ; checklist of species; Belgium).

— Lucht, 1987: 91 ( Syntomium ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Boháč, 1993: 42 ( Syntomium ; Slovakia; Czech Republic).

— Terlutter, 1995: 5 ( Syntomium ; Germany).

— Ciceroni and Zanetti, 1995: 10 ( Syntomium ; Italy).

- DISTRIBUTION: Europe, Russia.

nigroaeneum Curtis, 1828 : pl. 228 ( Syntomium ; Type locality: England, Middlesex , Southgate).

— Curtis, 1829: 28 ( Bolitobius ; catalog; Britain).

— Stephens, 1834: 330 ( Syntomium ; characters; England).

— Westwood, 1838a: 18 ( Syntomium ; type species).

— Stephens, 1839: 424 ( Syntomium ; characters; England).

— Erichson, 1840: 821 ( Syntomium ; synonym of aeneum ).

— Redtenbacher, 1849: 743 ( Syntomium ; synonym of aeneum ).

— Hardy, 1851: 53 ( Syntomium ; synonym of aeneum ).

— G. Waterhouse, 1858: 30 ( Syntomium ; synonym of aeneum ).

— Kraatz, 1857d: 892 ( Syntomium ; synonym of aeneum ).

— Mulsant and Rey, 1878: 825 ( Syntomium ; synonym of aeneum ).

— Ganglbauer, 1895: 681 ( Syntomium ; synonym of aeneum ).

— Bernhauer and Schubert, 1911: 86 ( Syntomium ; synonym of aeneum ).

confragosum Mäklin, 1852: 319 ( Syntomium ; Type locality: insula Sitkha).

— Fauvel, 1878d: 258 [= 1878e: 94] ( Syntomium ; characters; Alaska; Lake Superior).

— Bernhauer and Schubert, 1911: 87 ( Syntomium ; catalog).

— Hatch, 1957: 86 ( Syntomium ; characters; British Columbia).

— Herman, 1970: 373 ( Syntomium ).

— Downie and Arnett, 1996: 441 ( Syntomium ; characters; Canada; USA).

- DISTRIBUTION: Canada, USA.

grahami Hatch, 1957: 86 ( Syntomium ; Type locality: British Columbia: Stanley ).

— Herman, 1970: 373 ( Syntomium ).

- DISTRIBUTION: Canada.

japonicum Watanabe and Shibata, 1960: 103 ( Syntomium ; Type locality: Itsukaichi , Nishitama, Tokyo, Japan).

— Herman, 1970: 373 ( Syntomium ).

- DISTRIBUTION: Japan.

keenianum Casey, 1904: 314 ( Eunonia ; Type locality: British Columbia, Metlakatla ).

— Bernhauer and Schubert, 1910: 50 ( Eunonia ; catalog).

— Hatch, 1957: 69 ( Eunonia ; characters; British Columbia; Washington; Oregon).

— Moore, 1966: 56 ( Syntomium ).

— Herman, 1970: 373 ( Syntomium ).

longicorne Peyerimhoff, 1913: 253 ( Syntomium ; Type locality: Azerou Tidjer, vers 1300 m). — Scheerpeltz, 1933: 1074 ( Syntomium ; catalog).

— Herman, 1970: 373 ( Syntomium ).

- DISTRIBUTION: Algeria.

malkini Hatch, 1957: 86 ( Syntomium ; Type locality: Oregon: Coos Co.: Charleston ).

— Herman, 1970: 373 ( Syntomium ).

- DISTRIBUTION: USA.

marusiki Ryabukhin, 1992a: 147 ( Syntomium ; Type locality: Maradenskaia obl., 35 km severnee Maradana poc. Snezhnaia Dolina).

— Ryabukhin, 1993: 162 ( Syntomium ; translation of Ryabukhin, 1992a; Magadan Province).

— Ryabukhin, 1999: 35 ( Syntomium ; catalog; habitat; distribution; northeastern Russia).

- DISTRIBUTION: Russia.

nigroaeneum Curtis, 1828 , see: aeneum P. W. J. Müller, 1821 .

Tribe Oxytelini

Oxytelini :

— Thomson, 1858: 39 (characters; key to genera).

— Eichelbaum, 1909: 117 (list of genera with references; generic synonyms; number of genera; number of species in each genus; general distribution of genera).

— Reitter, 1909: 163 ( Bledius , Platystethus , Haploderus , Oxytelus ).

— Blatchley, 1910: 462 (characters; included Megalops , Platystethus , Oxytelus , Trogophloeus, Apocellus ; key to Indiana genera).

— Bernhauer and Schubert, 1911: 87 (world catalog; Coprophili, Actochari, Toxoderi, Oxyteli, Thinobii).

— Johansen, 1914: 530 (characters).

— Porta, 1926: 34 (characters; species of Italy).

— Cameron, 1930: 169 (characters; key to genera of British India).

— Scheerpeltz, 1933: 1073 (world catalog supplement).

— Scheerpeltz, 1940: 22 (key to Palaearctic genera).

— Crowson, 1955: 37 (genera).

— Arnett, 1963: 255 (characters; list of North American genera).

— Pototskaia, 1967: 17, 21 (larval characters; key to some genera).

— Tikhomirova, 1968: 146 (fossil species).

— Herman, 1970: 398 (characters; notes).

— Hammond, 1975: 145 (notes; characters; key to species of Ceylon).

— Muona, 1979: 19 (list of Scandinavian species).

— Outerelo and Gamarra, 1985: 100 (key to genera of Iberian Peninsula).

— Newton and Thayer, 1992: 60 (tribe of Oxytelinae ).

— Klimaszewski, Newton, and Thayer, 1996: 150 (list of New Zealand genera; number of

New Zealand species).

— M. Hansen, 1996: 37, 98 (not clearly defined; list of Danish species).

— Newton, Thayer, Ashe, and Chandler, 2000: 322, 380 (key to Nearctic genera; characters; notes).

Oxytelida:

— Heer, 1839a: 14 (cited as family).

Oxytelitae:

— Jeannel and Jarrige, 1949: 323 (notes).

Oxytelina:

— Jakobson, 1908: 446, 466 (characters; key to genera; list of species of Russia and western Europe).

— Johansen, 1914: 533 (characters; key to Danish genera).

— Arnett, 1963: 255 (characters; list of North American genera).

Oxyteli:

— Bernhauer and Schubert, 1911: 91 (world catalog; Planeustomus , Ancyrophorus , Trogophloeus , Xerophygus , Blediotrogus , Trogactus , Parosus, Apocellus , Oxytelopsis , Delopsis , Oxytelodes , Hoplitodes , Aploderus , Anisopsis , Oxytelus , Platystethus , Bledius , Typhlobledius ).

— Scheerpeltz, 1933: 1078 (world catalog supplement).

Apocellaria Lynch, 1884: 373 (genera included: Apocellus ). TYPE GENUS: Apocellus (see Newton and Thayer, 1992: 60).

— Newton and Thayer, 1992: 60 (synonym of Oxytelini ).

Ecitoclimacini Borgmeier, 1934: 452 (tribe of Aleocharinae ; genera included: Ecitoclimax ). TYPE GENUS: Ecitoclimax (see Newton and Thayer, 1992: 60).

— Newton and Thayer, 1992: 60 (synonym of Oxytelini ).

Apocellini Hatch, 1957: 85 (genera included: Apocellus ). TYPE GENUS: Apocellus (see Newton and Thayer, 1992: 60).

— Newton and Thayer, 1992: 60 (synonym of Oxytelini ).

Anisopsidius

[1 species; Ethiopian Region]

Anisopsidius Fagel, 1960: 8 (species included: quadricollis ). TYPE SPECIES: Anisopsis quadricollis Bernhauer , fixed by original designation and monotypy.

— Herman, 1970: 407 (characters; distribution; notes; list of species).

quadricollis Bernhauer, 1932a: 76 ( Anisopsis ; Type locality: Haut-Uele: Moto).

— Scheerpeltz, 1933: 1093 ( Anisopsis ; catalog).

— Fagel, 1957a: 35 ( Anisopsidius ; Ruanda).

— Fagel, 1960: 10 ( Anisopsidius ; characters; Belgian Congo; Ruanda).

— Herman, 1970: 408 ( Anisopsidius ).

- DISTRIBUTION: Congo, Rwanda.

Anisopsis

[1 species; Ethiopian Region]

Anisopsis Fauvel, 1904b: 108 (species included: flexuosa , carinata ). TYPE SPECIES: Anisopsis flexuosa Fauvel , fixed by subsequent designation by R. Lucas, 1920: 98.

— Bernhauer and Schubert, 1911: 109 (world catalog; 2 species).

— Scheerpeltz, 1933: 1093 (world catalog supplement).

— Blackwelder, 1952: 53 (type species).

— Fagel, 1960: 4 (characters).

— Herman, 1970: 406 (characters; distribution; notes; list of species).

carinata Fauvel, 1904 , see: Paroxytelopsis .

eichelbaumi Bernhauer, 1927 , see: Paroxytelopsis .

flexuosa Fauvel, 1904b: 108 ( Anisopsis ; Type locality: Zambese).

— Bernhauer and Schubert, 1911: 109 ( Anisopsis ; catalog).

— Scheerpeltz, 1933: 1093 ( Anisopsis ; catalog).

— Fagel, 1960: 6 ( Anisopsis ; characters; Zanzibar; Mashonaland).

— Herman, 1970: 407 ( Anisopsis ).

- DISTRIBUTION: Zimbabwe, “Zambezi”, Zanzibar, Congo, Cameroon.

rugosa Fagel, 1960: 7 ( Anisopsis ; subspecies of flexuosa ; Type locality: Congo Belge: Bassin Lukuga. Also cited from Cameroon).

guineensis Bernhauer, 1932 , see: Paroxytelopsis .

longicornis Cameron, 1928 , see: Paroxytelopsis .

minima Bernhauer, 1927 , see: Paroxytelopsis .

quadricollis Bernhauer, 1932 , see: Anisopsidius .

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Coprophilus

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