Gastrotheca helenae
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Barrio-Amorós, César L., Rojas-Runjaic, Fernando J. M. & Señaris, J. Celsa, 2019, Catalogue of the amphibians of Venezuela: Illustrated and annotated species list, distribution, and conservation, Amphibian & Reptile Conservation (e 180) 13 (1), pp. 1-198
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Gastrotheca helenae
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Cruz-Rodríguez, Cristian, Lynch, John D., Cárdenas, Julieth S. & Caicedo-Portilla, José Rancés, 2021, Type specimens housed in the herpetological collections at Museo de La Salle Bogotá and their historical importance for the consolidation of herpetology in Colombia, Zootaxa 4965 (2), pp. 261-292
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