Magnificus dirschi ( Bang-Haas, 1939 ) Grehan & Mielke & Minet & Ignatev & Buchsbaum & Xue, 2021

Grehan, John R., Mielke, Carlos G. C., Minet, Joël, Ignatev, Nikolai, Buchsbaum, Ulf & Xue, Dayong, 2021, Taxonomic composition and monophyly of the genus Magnificus (Lepidoptera Hepialoidea: Hepialidae), Zootaxa 4920 (3), pp. 339-358 : 344

Magnificus dirschi ( Bang-Haas, 1939) comb. n.

( Figs 3 , 11 )

Phassus dirschi Bang-Haas (1939: 59)

Sthenopis dirschi: Nielsen et al. (2000: 850)

Material examined. Holotype ♂ MfN: Verbatim label: [ China], Kansu [recte Gansu] occ., Liang tschou, Richthofen mont. mar. / Slide ESN 2486 ♂ / Zool. Mus. Berlin / One more red label.

Original description of Phassus dirschi ( Bang-Haas 1939) . “ Upper side of FW dark brown, on the front edge are four brown, white edged spots. A white band runs from the 2nd spot to the middle of the forewings. From the 2nd spot a white band runs to the middle of FW, from the 4th spot a brown band with 2 metal-gold spots, then a further, larger golden spot towards the base. A straight white submarginal band extends just before the apex to the inner edge, a 6 mm long oblique band of the same colour ends in the tip of FW. At the base of the inner margin is a carmine-red spot. Underside of FW brownish; the markings of HW evenly carmine red. Both wings with blackbrown fringes. Underside of HW dirty red. Head, thorax brownish red, abdomen carmine red, the last segments brownish. Antennae yellowish.”

Female. Unknown.

Etymology. Named in honour of P. Mansuetus Dirsch ( Bang-Haas 1939).

Flight period. Late spring-early summer. Holotype collected June 20 th, 1938 ( Bang-Haas 1939)

Remarks. Postdiscal longitudinal band extending as a narrow sub-straight white line between costal margin near junction of Rs1 and Rs2 and CuA 2 ( Fig. 30b ). Slide mounted image of male genitalia ( Fig. 11 ) indicate that the lateral flange on the posterior saccus margin is more broadly extended laterally than in other Magnificus species. Sympatric within distribution range of S. regius ( Fig. 31 ).

Figures

FIGURE 30. Magnificus FW major banding patterns (coloured outlines). Wing veins—blue lines. FW illustrations 30g, h of limited quality. Higher resolution unavailable.

FIGURES 1–9. Adult moths: 1, Magnificus bouvieri male holotype, dorsal (MNHN); 2, M. dirschi male holotype, dorsal (MfN). Photo by Théo Léger; 3, M. miniatus female holotype, (a) dorsal, (b) ventral (IZAS). Photo by Dayong Xu; 4, M. regius: (a) female syntype (from Staudinger 1896, pl. 5, Fig. 11), (b) M. regius rubellus, male lectotype, dorsal (MfN). Photo by Théo Léger, (c) M. regius rubellus (ZSBS), Photo by Ulf Buchsbaum, (d) M. regius rubellus (MWM). Photo Nikolai Ignatev; 5, M. roseus: (a) holotype male, dorsal (MNHN). Photo by Joël Minet, (b) male, dorsal (ZSBS), Photo by Ulf Buchsbaum; 6, M. zhidu- oensis female holotype, dorsal [cropped image available only] (GAAH). Photo by Dayong Xue; 7, Magnificus sp. female, dorsal (MWM). Photo by Nikolai Ignatev; 8, Hepialus humuli, female, dorsal (Moscow, Russia SKC). Photo by Svyatoslav Knyazev; 9. Zenophassus schamyl, male, dorsal (Apsheronsk environs, Caucasus, Russia SKC). Photo by Svyatoslav Knyazev.

FIGURES 10–20. Genitalia. 10, Magnificus bouvieri male holotype genitalia, ventral view. Photo by Joël Minet; 11, Magnificus dirschi male holotype genitalia, ventral view. Photo by Théo Léger; 12, Magnificus regius genitalia: a, male syntype, ventral view, dissection ESN 2845(MfN). Photo by Théo Léger; b, male, ventral view (ZSBS). Photo by Ulf Buchsbaum; c, male, ventral (left) and lateral (right) (MWM). Photo by Nikolai Ignatev; 13, Magnificus roseus male genitalia, ventral view, (a) holotype (MNHN). Photo by Joël Minet, (b) (ZSBS). Photo by Ulf Buchsbaum; Fig. 16. Magnificus jiuzhiensis genitalia: (a) holotype male, ventral view (from Yan 2000); 15, ‘Hepialiscus sylvina Linnaeus, 1761’, male, ventral view (from Chu & Wang 1985b: fig. 31 and Zhu et al. 2004: fig. 120); 16, Magnificus jiuzhiensis, female external posterior view (from Yan 2000); 17, Magnificus miniatus female holotype external genitalia, posterior view. Photo by Dayong Xu; 18, Magnificus regius female external posterior and lateral interior view (MWM). Photo by Nikolai Ignatev; 19, Magnificus zhiduoensis female holotype external posterior view (from Yan 2000); 20, Magnificus sp. female external posterior view (MWM). Photo by Nikolai Ignatev.

FIGURES 31–32. Distribution: 31, Magnificus species distribution records. Offset symbols represent different species at the same locality; 32, Disjunct allopatry of Magnificus (red outline with blue dots) and the northern Eurasian sister group comprising Hepialus humuli—blue outline, Pharmacis—crimson outline, Triodia—yellow outline, Zenophassus schamyl—dark green outline. Distribution data for northern Eurasian genera from Grehan & Knyazev (2019).