Magnificus roseus ( Oberthür, 1911 ) Grehan & Mielke & Minet & Ignatev & Buchsbaum & Xue, 2021

Grehan, John R., Mielke, Carlos G. C., Minet, Joël, Ignatev, Nikolai, Buchsbaum, Ulf & Xue, Dayong, 2021, Taxonomic composition and monophyly of the genus Magnificus (Lepidoptera Hepialoidea: Hepialidae), Zootaxa 4920 (3), pp. 339-358 : 346

Magnificus roseus ( Oberthür, 1911) comb. n.

( Figs 5a, 5b , 13a, 13b , 22c , 23g )

Hepialus (Phassus) roseus Oberthür (1911: 325 ; Fig. 652)

Phassus roseus: Pfitzner (1912: 438)

Phassus regius roseus: Bang-Haas (1939: 58)

Sthenopis roseus: Nielsen et al. (2000: 850)

Examined material: Lectotype ♂, MNHN. Verbatim label: Frontière orientale du Thibet, Chasseurs indigènes du P. Déjean, 1905 / ESN slide no. 2871 ( Fig. 5a ); 1 ♂, [ China, Xizang], Tâ-tsien-lou [ Kangding ], 1898. local collector, ZSBS ( Fig. 5b ).

Original description ( Oberthür 1911): Illustration only, plate 5, Fig. 11 , no written description. “Placed close to regius Staudinger. ”

Female: unknown.

Etymology. Presumably named for the rose-coloured tint of the hindwings.

Flight period. Unknown.

Remarks. Elongate dark shaded patch present between outer anterior discal cell and basal Rs4-M 1 and M 1 -M 2 cells ( Fig. 30f ) similar to M. miniatus ( Fig. 30d ). Originally described and illustrated for a single male specimen which is recognized here as the lectotype. Sympatric with M. bouvieri ( Fig. 31 ).

Figures

FIGURE 23. Magnificus abdominal sclerites: (a) M. regius tergite II, (MWM), photo by Nikolai Ignatev, (b) M. bouvieri holotype SII (photo by Joël Minet), (c) Magnificus sp. sternites VII-VIII, photo by Nikolai Ignatev), (d) M. bouvieri holotype sternites VII-VIII, photo by Joël Minet), (e) M. regius sternites VII-VIII, photo by Nikolai Ignatev, (f) M. regius and (g) M. roseus, photos by Ulf Buchsbaum, (h) M. bouvieri holotype abdomen, photo by Joël Minet, (i) Magnificus sp. tergosternal sclerite, (j) Magnificus regius tergosternal sclerite, photos by Nikolai Ignatev, (k) Hepialus humuli tsclerite, (l) Zenophassus schamyl tergosternal sclerite, photos by John Grehan.

FIGURE 30. Magnificus FW major banding patterns (coloured outlines). Wing veins—blue lines. FW illustrations 30g, h of limited quality. Higher resolution unavailable.

FIGURES 1–9. Adult moths: 1, Magnificus bouvieri male holotype, dorsal (MNHN); 2, M. dirschi male holotype, dorsal (MfN). Photo by Théo Léger; 3, M. miniatus female holotype, (a) dorsal, (b) ventral (IZAS). Photo by Dayong Xu; 4, M. regius: (a) female syntype (from Staudinger 1896, pl. 5, Fig. 11), (b) M. regius rubellus, male lectotype, dorsal (MfN). Photo by Théo Léger, (c) M. regius rubellus (ZSBS), Photo by Ulf Buchsbaum, (d) M. regius rubellus (MWM). Photo Nikolai Ignatev; 5, M. roseus: (a) holotype male, dorsal (MNHN). Photo by Joël Minet, (b) male, dorsal (ZSBS), Photo by Ulf Buchsbaum; 6, M. zhidu- oensis female holotype, dorsal [cropped image available only] (GAAH). Photo by Dayong Xue; 7, Magnificus sp. female, dorsal (MWM). Photo by Nikolai Ignatev; 8, Hepialus humuli, female, dorsal (Moscow, Russia SKC). Photo by Svyatoslav Knyazev; 9. Zenophassus schamyl, male, dorsal (Apsheronsk environs, Caucasus, Russia SKC). Photo by Svyatoslav Knyazev.

FIGURES 10–20. Genitalia. 10, Magnificus bouvieri male holotype genitalia, ventral view. Photo by Joël Minet; 11, Magnificus dirschi male holotype genitalia, ventral view. Photo by Théo Léger; 12, Magnificus regius genitalia: a, male syntype, ventral view, dissection ESN 2845(MfN). Photo by Théo Léger; b, male, ventral view (ZSBS). Photo by Ulf Buchsbaum; c, male, ventral (left) and lateral (right) (MWM). Photo by Nikolai Ignatev; 13, Magnificus roseus male genitalia, ventral view, (a) holotype (MNHN). Photo by Joël Minet, (b) (ZSBS). Photo by Ulf Buchsbaum; Fig. 16. Magnificus jiuzhiensis genitalia: (a) holotype male, ventral view (from Yan 2000); 15, ‘Hepialiscus sylvina Linnaeus, 1761’, male, ventral view (from Chu & Wang 1985b: fig. 31 and Zhu et al. 2004: fig. 120); 16, Magnificus jiuzhiensis, female external posterior view (from Yan 2000); 17, Magnificus miniatus female holotype external genitalia, posterior view. Photo by Dayong Xu; 18, Magnificus regius female external posterior and lateral interior view (MWM). Photo by Nikolai Ignatev; 19, Magnificus zhiduoensis female holotype external posterior view (from Yan 2000); 20, Magnificus sp. female external posterior view (MWM). Photo by Nikolai Ignatev.

FIGURES 21–22. Magnificus thoracic appendages: 21, wing venation a, Magnificus sp. female, b, M. regius male. Diagrams by John Grehan; 22, legs: left to right – pro. meso, meta (a) Magnificus sp. female (MWM), photo by Nikolai Ignatev, (b) Magnificus regius male (ZSBS), (c) Magnificus roseus male, photos by Ulf Buchsbaum.

FIGURES 31–32. Distribution: 31, Magnificus species distribution records. Offset symbols represent different species at the same locality; 32, Disjunct allopatry of Magnificus (red outline with blue dots) and the northern Eurasian sister group comprising Hepialus humuli—blue outline, Pharmacis—crimson outline, Triodia—yellow outline, Zenophassus schamyl—dark green outline. Distribution data for northern Eurasian genera from Grehan & Knyazev (2019).