Plantago atrata carpatica (Pilg.) Soo , Acta Geobot. Hung. 3: 61 (1940)

Novikov, Andriy, 2023, An annotated nomenclatural checklist of endemic vascular plants distributed in the Ukrainian Carpathians, Biodiversity Data Journal 11, pp. 103921-103921 : 103921

Plantago atrata carpatica (Pilg.) Soo, Acta Geobot. Hung. 3: 61 (1940)

Plantago atrata ≡ Plantago atrata subsp. atrata var. carpathica (Pilg.) Pilg., Pflanzenr. (Engler) 102: 296 (1937); GBIF: https://www.gbif.org/species/8397828; BHL: https://www.biodiversitylibrary.org/item/71651#page/304

Plantago montana ≡ Plantago montana subsp. atrata var. carpathica Pilg., Repert. Spec. Nov. Regni Veg. 23: 256 (1926-1927); GBIF: https://www.gbif.org/species/7564352; POWO: https://powo.science.kew.org/taxon/3007639-4

Plantago montana ≡ Plantago montana subsp. carpatica (Pilg.) Soó ex Balázs, Acta Geobot. Hung. 2: 40 (1938-1939); POWO: https://powo.science.kew.org/taxon/3007638-4

Plantago atrata = Plantago atrata subsp. atrata var. carpathica subvar. rigidior (Pilg.) Pilg., Pflanzenr. (Engler): 296 (1937); BHL: https://www.biodiversitylibrary.org/item/71651#page/304

Plantago atrata = Plantago atrata subsp. atrata var. carpathica subvar. vestita (Pilg.) Pilg., Pflanzenr. (Engler): 296 (1937); BHL: https://www.biodiversitylibrary.org/item/71651#page/304

Plantago atrata = Plantago atrata var. carpathica f. vestita (Pilg.) Borza, Consp. Fl. Rom.: 255 (1949)

Plantago atrata = Plantago atrata subsp. carpathica f. vestita (Pilg.) Soó, Acta Geobot. Hung. 3: 61 (1940)

Plantago lanceolata = Plantago lanceolata [unranked] β Plantago alpestris Wahlenb., Fl. Carpat. Princip.: 44 (1814)

Plantago montana = Plantago montana [unranked] Thlaspi alpestre Wahlenb., Fl. Carpat. Princip.: 44 (1814)

Plantago montana = Plantago montana subsp. atrata var. carpathica subvar. rigidior Pilg., Repert. Spec. Nov. Regni Veg. 23: 257 (1926-1927)

Plantago montana = Plantago montana subsp. atrata var. carpathica subvar. vestita Pilg., Repert. Spec. Nov. Regni Veg. 23: 257 (1926-1927)

Plantago montana = Plantago montana subsp. carpatica subvar. rigidior (Pilg.) Balázs, Acta Geobot. Hung. 2: 40 (1938-1939)

Plantago montana = Plantago montana subsp. carpatica subvar. vestita (Pilg.) Balázs, Acta Geobot. Hung. 2: 40 (1938-1939)

Plantago alpina - Plantago alpina Vill. sensu Rochel, Pl. Banat. Rar.: 32; Nr. 4, Tab. 1, fig. 4 (1828)

Plantago alpina - Plantago alpina Vill. sensu Schur, Enum. Pl. Transsilv.: 564 (1866), non alior; BHL: https://www.biodiversitylibrary.org/item/7364#page/584

Plantago atrata - Plantago atrata Hoppe, Bot. Taschenb. 1799: 85 (1799) [p. p., tantum quod plantas ucrain. carpat.] *; GBIF: https://www.gbif.org/species/8083357; IPNI: https://www.ipni.org/n/684903-1; WFO: http://www.worldfloraonline.org/taxon/wfo-0000487646; POWO: powo.science.kew.org/taxon/684903-1; BHL: https://www.biodiversitylibrary.org/item/22931#page/95

Plantago montana - Plantago montana Lam. sensu Schur, Enum. Pl. Transsilv.: 564 (1866), non alior *; BHL: https://www.biodiversitylibrary.org/item/7364#page/584

Plantago saxatilis - Plantago saxatilis M.Bieb., Fl. Taur.-Caucasus 1: 109 (1808) [p. p.]; GBIF: https://www.gbif.org/species/4156187; IPNI: https://www.ipni.org/n/685623-1; WFO: http://www.worldfloraonline.org/taxon/wfo-0000487134; POWO: https://powo.science.kew.org/taxon/685623-1; BHL: https://www.biodiversitylibrary.org/page/11268189#page/117

Conservation status

In Ukraine - LC ( Onyshchenko et al. 2022).

Distribution

Pancarpathian endemic.

Notes

There are nine subspecies of P. atrata Hoppe, from which only P. atrata subsp. carpatica is usually reported for the Ukrainian Carpathians. However, Chrtek (2000) also delimited P. atrata subsp. ucrainica Chrtek that, as he indicated, mainly occurs in the Svydovets Mts. Besides this, he mentioned the presence of this subspecies in Romania (Slănic Moldova and Retezat). Plantago atrata subsp. ucrainica differs by erect ascending (vs. decumbent to prostrate in P. atrata subsp. carpatica ), longer (up to 17 cm long vs. 14 cm in P. atrata subsp. carpatica ) and more narrow (up to 8 mm wide vs. 16 mm P. atrata subsp. carpatica ) leaves. Distribution and phylogenetic position of P. atrata subsp. ucrainica still requires clarifications since, after Chrtek (2000), there were no further corresponding investigations on this subspecies.

The combination P. montana subsp. carpatica Soó, Acta Geobot. Hung. 2: 40 (1938-1939) and consequent recombination P. atrata subsp. carpatica ( Soó) Soó, Acta Geobot. Hung. 3: 61 (1940), commonly circulated in the checklists, seem to be incorrect because it was Pilger who first applied the epithet Amphigenes carpathica in the name P. montana subsp. atrata var. carpathica Pilg. in 1926 ( Pilger 1926). Later, in 1937, Pilger introduced a new combination P. atrata subsp. atrata var. carpathica (Pilg.) Pilg. ( Pilger 1937). It looks like Soó made further taxonomic recombinations, based on these two Pilger’s names, but, unfortunately, I could not locate the original works of Soó regarding P. atrata to check.

GBIF (https://www.gbif.org/species/11030083, accessed on 06.06.2023) incorrectly provides the name P. atrata subsp. carpathica (Pilg.) Pilg. It should be either P. atrata var. carpathica (Pilg.) Pilg. (incorrect taxonomic rank is indicated) or P. atrata subsp. carpatica (Pilg.) Soó (the incorrect authorship is provided). In both cases, the entry is duplicating other existing records.