Veronica aragonensis Stroh (1942: 398)

Rojas-Andrés, Blanca M. & Martínez-Ortega, Montserrat, 2016, Taxonomic revision of Veronica subsection Pentasepalae (Veronica, Plantaginaceae sensu APG III), Phytotaxa 285 (1), pp. 448-450 : 448-450

1. Veronica aragonensis Stroh (1942: 398) [Iconography: Martínez-Ortega et al. 2009, lám. 92]

Stems (5) 10–20 (30) cm long, decumbent to ascending, with a woody and very branched base, covered completely or almost completely by the rocks, the aerial part herbaceous, ± flexuous, glabrous (principally to the base) or with short hairs (0.1) 0.2–0.3 (0.35) mm long, incurvate, appressed and antrorse, more abundant to the apex, where some disperse glandular hairs could appear; apical shoot usually not appreciable. Leaves (2) 7–10 (14) × (1.5) 3–6 (10) mm, the basal and medium ones similar, oblong, elliptic, ovate or very widely ovate, cuneate to attenuate at the base, subentire to crenate, glabrous or with some hairs 0.15–0.4 mm long, sparsely arranged, mainly along the margin. Racemes often pseudoterminal or axillary, opposite, bearing 3–8 (12) flowers, loosely to densely arranged; peduncles from almost not appreciable to up to 3.5 cm long, covered by glandular hairs similar to those of the stem and slightly longer, ± patent, (0.25) 0.3–0.5 (0.6) mm long; bracts (1.5) 3–7 (8) mm long, elliptic, narrowly elliptic, narrowly ovate or lanceolate, entire, indument similar to that of the stem; pedicels (1) 1.5–4 (7) mm long. Calyx (3.5) 4–6 (8) mm long, 4 (5) sepals, narrowly elliptic to oblanceolate, generally as long as or slightly longer than the capsule, sometimes slightly shorter, covered by abundant glandular hairs. Corolla 8–10 mm in diameter, pale

8 • Phytotaxa 285 (1) © 2016 Magnolia Press

ROJAS-ANDRÉS & MARTÍNEZ-ORTEGA blue or rarely pink. Capsule (3) 4.5–5 (6.5) × (3.5) 5–5.5 (7) mm, commonly wider than long, transversely elliptic to very widely obovate, ± rounded at the base, emarginate, sinus up to 1.1 (1.4) mm depth, with abundant glandular hairs, especially along the margin. Style (2) 2.5–3 (3.5) mm long. Seeds 1.6–2.2 × 0.6–1.7 mm, ca. 10–15 per capsule.

Chromosome Number: —2 n = 32.

Habitat: —Screes and stony places, on alpine or subalpine stations, preferentially on limestones; (1,000) 1,400–2,500 m.

Distribution: —Endemic to Spain with disjunct populations in Central Pyrenees and Pre-Pyrenees (Huesca) and Sierra de la Sagra (Granada) ( Fig. 1 ).

Phenology: — Flowering from (May) June to July.

Specimens Examined: — SPAIN. Granada: 15 Km of Puebla de Don Fabrique, Cerro La Sagra, exposed upper slopes of mountain, limestone rocks and screes, 2000 m, 37º57'N, 2º33'W, 6 July 1979, Cannon, Crane, Jury & Moore s.n. (RNG); Huéscar, Sierra de la Sagra, 1750–1800 m, WH3800, 23 April 1997, Martínez Ortega, Rico & Sánchez Agudo MO 236 (SALA93528); Huéscar, Sierra de la Sagra, 1800–2350 m, WH3800–WH3801, 27 July 1996, E. Rico, Romero & Martínez Ortega MO 216 (SALA93529); Huéscar, Sierra de la Sagra, cara S, WG3899, 14 June 1995, Cabezudo, Pérez Latorre, Toro & Navas s.n. (MGC39901, MGC40213); Huescar, Sierra de la Sagra, pedregales móviles proximos a la cumbre, 2 July 1983, Negrillo s.n. (GDA18086); Huesca: Arguis, Bentué de Rasal, peñascos-cueva junto a glera, 1000–1100 m, 30TYM0790, 12 May 1974, P. Montserrat s.n. (JACA52774); Barbaruéns, Cotiella, descenso por senda itinerario hasta la fuente, 1820–2150 m, 31TBH8211, 14 September 1977, P. Montserrat s.n. (JACA0227377); Bisaurri, Montaneta de Gabas, ladera oriental del Monte Baciero, 1900– 2000 m, 31TBH9204, 4 July 1988, J. A. Sese & P. Montserrat s.n. (JACA0665787); Cotiella, Ibón de Plan, 25 & 26 July 1969, Fdz. Casas s.n. (MA408515); Guara, cumbres, June 1947, unknown collector (BC41230); La Estiva, umbría de la Estiva, 1600–1830 m, 31TBH7506, 22 July 1979, P. Montserrat s.n. (JACA256379); Laspuña, Peña Montañesa, 2150–2290 m, 31TBH7107, 27 July 1974, P. Montserrat s.n. (JACA478174); Monte Turbón, in glareosis, glera sobre el muro occidental, 35 SW, 9 July 1952, P. Montserrat s.n. (K); Nocito, Guara, pinares, cantil NW, barranco, crestones y gleras, 1700–1750 m, 30TYM2785, 12 June 1967, P. Montserrat s.n. (JACA311767); Oncins, Peña Montañesa, 1050–1720 m, 31TBH7106, 17 May 1981, P. G. Montserrat & D. Gómez s.n. (JACA34718); Ordesa, sobre las clavijas de Soaso, 2100 m, 31TBH5527, 15 July 1971, P. Montserrat s.n. (JACA473871); Peña Montañesa, subida desde el Collado de Ceresa, glera caliza, 1850 m, 31TBH7109, 21 June 1996, Martínez Ortega & Martín Ballesteros MO147 (SALA93531); Plan, Collado de Sahún, en gleras calizas, 2000 m, 31TBH8616, 16 July 1994, Martínez Ortega & J. X. Soler MO23 (SALA93532); Prepirineos aragoneses, Peña Montañesa (loc. class.), NE of Ainsa, on glera (scree) on high slopes, ca. 2000 m and upwards, 13 July 1956, Sandwith s.n. (K); Revilla, cabecero de Barranco de Angonés, Collado y Barranco Seco, 1900–2450 m, 31TBH6523, Villar, Sese & Perez s.n. (JACA258891); Saravillo, tierra removida, prados con pino negro y abeto, pista de Lavasar, 31TBH7815, 30 July 1975, P. Montserrat & Villar s.n. (JACA364175); Subida a la Peña Montañesa desde el Collado de Ceresa, calizas, 1850 m, 31TBH7109, 9 August 1996, Martínez Ortega, Muñoz Centeno & Gonzalez Rivera MO-220 (SALA93530); Torla, Ordesa, gran glera solana, hacia Cola de Caballo, 1900–1950 m, 31TBH5526, 3 July 1973, P. Montserrat s.n. (JACA277273); Valle de Ordesa, Pirineo Central, circo de Soaso, 1850 m, 3 July 1973, Montserrat s.n. (BC618481); Vilas de Turbón, entre el pueblo y Monte Turbón, 31TBG9699, 20 June 1987, P. & G. Montserrat s.n. (JACA0581987); Yésero-Gavín, cantiles calizos y pedrizas, solana de Tendeñera, 2000–2300 m, 30TYN2628, 7 August 1979, L. Villar s.n. (JACA349179); Bisaurri; Gabás, en el ascenso al Monte Baziero, Gleras calizas, 1823 m, 42º27'45.1''N, 00º27'56.2''E, 1 November 2015, N. Padilla-García, M. Martínez-Ortega, D. Gutierrez Larruscain & Celia Hernandez Galindo NPG70-2015 (SALA157043); Collado de Ceresa; subida a la Peña Montañesa, 31T 270540 4708236, 16 July 2007, A. Abad de Blas, M. Martínez Ortega & A. Tribsch AA14 (SALA121537, SALA121543); Entre Arguis y Bentué de Rasal, gleras calizas, 30TYM 707149 4689902, 17 July 2007, A. Abad de Blas, M. Martínez Ortega & A. Tribsch MO2047 (SALA121540); Nocito; Sierra de Guara; subida al Tozal de Guara, 30 TYM 728340 4685652, 17 July 2007, A. Abad de Blas, M. Martínez Ortega & A. Tribsch AA16 (SALA121538); Plan, Collado de Sahún, glera de rocas ácidas, 1980–2000 m, 31TBH8616, 1 August 1999, M. Martínez Ortega & L. Delgado MO757 (SALA120830); Seira, Barbaruens, sendero que sube hacia el refugio de Armiña, gleras calizas, 1796 m, 42º30'45.3''N, 00º21'35.5''E, 31 October 2015, N. Padilla-García, M. Martínez-Ortega, D. Gutierrez Larruscain & Celia Hernandez Galindo NPG69-2015 (SALA157042); Vilas de Turbón, subida hacia Turbonet, gleras calizas, 31TBG 296372 4698413, 15 July 2007, A.

TAXONOMIC REVISION OF VERONICA SUBSECTION PENTASEPALAE

Phytotaxa 285 (1) © 2016 Magnolia Press • 9 Abad de Blas , M . Martínez Ortega & A . Tribsch MO2042 ( SALA121536 ); Vilas del Turbón ; subida al monte Turbón, 31 TBG 296300 4698554, 15 July 2007, A . Abad de Blas , M . Martínez Ortega & A . Tribsch AA13 (ind 2- 20) ( SALA121542 ) ; Vilas del Turbón ; subida al monte Turbón, 31 TBG 296417 4698523, 15 July 2007, A . Abad de Blas , M . Martínez Ortega & A . Tribsch AA13 (ind 1) ( SALA121535 ) ; Bisaurri, Gabás, subida a la Montañeta de Gabás , en el ascenso al Monte Baziero , gleras calizas situadas más al este, 1830 m, 42º27'45.00"N, 0º27'56.20"E, 09 July 2014, N . Padilla-García & G . Sgarlata NPG13 ( SALA154272 ); Nerín, ruta desde Nerín hacia “cuello Arenas” al llegar a la zona umbría (orinetación Norte), montaña "La Estiba", gleras calizas a un lado y otro de la pista, 1755 m, 42º35'54.10"N, 0º 0'30.70"E, 12 July 2014, N . Padilla-García & N . Machon & G . Sgarlata NPG18 ( SALA154410 ); Plan, pista desde Plan a Chía para el collado de Sahún, gleras calizas, 1722 m, 42º33'13.50"N, 0º26'9.80"E, 08 July 2014, N . Padilla-García , N . Machon , G . Sgarlata & M . Daoud NPG12 ( SALA154268 ); Seira, Barbaruens , Pista que sube al refugio de Armiña, gleras calizas de alta montaña, 1806 m, 42º30'44.8''N, 0º21'37.0''E, 10 July 2014, N . Padilla-García , N . Machon , G . Sgarlata & M . Daoud NPG15 ( SALA154362 ); Yésero, barranco del Puerto (Sierra Tendereña), gleras calizas de alta montaña, 1971 m, 42º40'12.10"N, 0º12'25.70"O, 14 July 2014, N . Padilla-García , G . Sgarlata & M . Daoud NPG22 ( SALA155054 ); unknown province: unknown collector ( BCF41411 ) .

Figures

FIGURE 1. Distribution map of V. aragonensis based on herbarium specimens (black circles) and collected material (open circles).