Burmochlamys poongee (Theobald, 1859) Pholyotha & Sutcharit & Lin & Panha, 2022

Pholyotha, Arthit, Sutcharit, Chirasak, Lin, Aung & Panha, Somsak, 2022, Uncovering local endemism from southeastern Myanmar: description of the new karst-associated terrestrial snail genus Burmochlamys (Eupulmonata, Helicarionidae), ZooKeys 1110, pp. 1-37 : 1

publication ID	https://dx.doi.org/10.3897/zookeys.1110.82461
publication LSID	lsid:zoobank.org:pub:01302157-EE40-4B99-99DD-3EC3377D929C
persistent identifier	https://treatment.plazi.org/id/63E44D84-EC38-58B3-B53A-7CE60D550B83
treatment provided by	ZooKeys by Pensoft
scientific name	Burmochlamys poongee (Theobald, 1859)
status	comb. nov.

Treatment

Burmochlamys poongee (Theobald, 1859) comb. nov.

Figs 1 View Figure 1 , 2B View Figure 2 , 3B View Figure 3 , 5C View Figure 5 , 6E, F View Figure 6 , 8D-F View Figure 8 , 9B View Figure 9 , 11B View Figure 11

Helix poongee Theobald, 1859: 307. Type locality: prope Moulmein [Mawlamyine, Mon State, Myanmar].

Helix poongee - Pfeiffer, 1868: 134; Hanley and Theobald 1876: 8, pl. 16, fig. 9.

Helix poongi Theobald, 1876: 19 [incorrect subsequent spelling].

Macrochlamys poongee - Godwin-Austen, 1882: 90, pl.14, fig. 1; Pholyotha et al. 2020b: 190, 191, figs 3e, 3f.

Nanina poongee - Tryon, 1886: 98, pl. 33, fig. 70.

Macrochlamys pungi - Blanford and Godwin-Austen, 1908: 122. [unjustified emendation; ICZN 1999: Art.19.1 and 33.2.3].

Material examined.

Type material. The type series could not be located.

Other material.

Moulmein: NHMUK 1869.06.1.1 ex. Blanford Coll. (one shell; specimen figured in Pholyotha et al. 2020b: fig. 3e), NHMUK 1888.12.4.23- 26 ex. Blanford Coll. (four shells; Fig. 6E View Figure 6 ; specimen figured in Pholyotha et al. 2020b: fig. 3f), NHMUK ex. Blanford Coll. (two shells). Kaw Ka Thaung Cave , Hpa-An, Kayin State, Myanmar (16°49'42.0"N, 97°42'22.9"E): CUMZ 14210 (Fig. 6F View Figure 6 ) GoogleMaps .

Diagnosis.

Shell globose and pale to dark brownish. Animal pale to dark greyish with a brown or yellow tinge and five mantle extensions. Genitalia with slender epiphallus, rather short epiphallic caecum attached by a thin penial retractor muscle, and very long and slender at the base of dart apparatus.

Description.

Shell (Figs 6E, F View Figure 6 , 8D-F View Figure 8 ). Shell globose, small (width up to 6.5 mm, height up to 5.2 mm), rather thin, semi-translucent. Colour pale to dark brownish. Protoconch and teleoconch surface with similar sculpture. Surface of body whorl with distinct spiral furrows at regular intervals, crossed by distinctly undulating radial lines (Fig. 8D-F View Figure 8 ). Whorls 6-7, increasing regularly; suture shallowly impressed; spire much elevated; last whorl well-rounded. Aperture obliquely crescent-shaped; peristome simple. Columellar margin simple, slightly reflected near umbilicus. Umbilicus open, narrow, and deep.

Genital organs (Fig. 9B View Figure 9 ). Atrium (at) rather short. Penis (p) moderate, cylindrical, and with a penial sheath. Epiphallus (e1+e2), slender, ca. four-fifths of penis length; e1 cylindrical and its diameter smaller than penis and e2; e2 bulbous and ca. one-third of e1 length. Epiphallic caecum (ec) rather short, cylindrical, and with a thin penial retractor muscle (prm) attached at tip. Vas deferens (vd) thin tube. Dart apparatus large, very long cylindrical with at the base very long, small, convoluted, and located on atrium near genital orifice. Vagina (v) rather short, and cylindrical. Gametolytic sac (gs) bulbous; gametolytic duct (gd) cylindrical, moderate, ca. three-fifths of penis length. Free oviduct (fo) ca. three-fifths of penis length, cylindrical, and encircled with thick tissue near vagina.

Radula (Fig. 11B View Figure 11 ). Resembles B. cassidula . Teeth arranged in wide U-shaped rows with each row consisting of ~ 60 teeth. Central tooth monocuspid, large and ovate spatulate shape; with two smaller triangular-shaped plates. Laterals and marginals not differentiated, monocuspid, large spatulate shape and then gradually become narrower, elongate, and rather more pointed cusps, and with one smaller triangular-shaped plate at base. Some outermost teeth with small and pointed cusp at inner side.

External appearance (Fig. 3B View Figure 3 , 5C View Figure 5 ). Living animal with reticulated skin, pale to dark greyish with a brown or yellow tinge, slightly lighter on foot sole and darker colour on caudal horn. Mantle extension well developed with three dorsal lobes and two shell lobes, similar colour to body.

Distribution.

Burmochlamys poongee can be found only from Kaw Ka Thaung Cave in Myanmar (Figs 1 View Figure 1 , 2B View Figure 2 ). This limestone is surrounded by paddy fields which are temporarily flooded during the monsoon season (Fig. 2B View Figure 2 ).