Camchaya gracilis (Gagnep.) S. Bunwong & H. Rob., Proc. Biol. Soc. Washington 122(3): 361. 2009.

Bunwong, Sukhonthip, Chantaranothai, Pranom & Keeley, Sterling C., 2014, Revisions and key to the Vernonieae (Compositae) of Thailand, PhytoKeys 37, pp. 25-101 : 50

Camchaya gracilis (Gagnep.) S. Bunwong & H. Rob., Proc. Biol. Soc. Washington 122(3): 361. 2009.

Iodocephalus gracilis Thorel ex Gagnep., Notul. Syst. (Paris) 4: 17. 1920.

Type.

Laos, Bassac, Thorel 2396 (holotype: P!). Fig. 6A .

Description.

Annual herbs, 50-100 cm tall. Stems erect, terete, inconspicuously ribbed; villose with uniseriate hairs, T-shaped hairs and glands. Leaves alternate, elliptic-oblong, 3-6 by 0.3-1 cm, margin serrate or entire, apex acute, base attenuate, subcoriaceous; upper surface scabrous lacking glands, lower surface scabrous with whip-shaped hairs and capitate glands, lateral veins 8-10-paired; petioles up to 3 mm long. Capitulescences terminal and axillary, corymbose. Capitula campanulate, 8-10 mm long, pedunculate. Receptacle flat, 2-3 mm in diam., glabrous. Involucres campanulate, in 4-5 series, 7-8 mm long, 6-7 mm in diam. Phyllaries imbricate, light green or purple, margin entire without spines, outer surface arachnoid, glandular; the outer ones ovate, apex acute to acuminate; the inner ones ovate-lanceolate, apex acute to acuminate. Florets 50-70; corollas funnelform, purple, puberulous, glands capitate; corolla tubes 3-5 mm long; corolla lobes 2-3 mm long. Anthers ca. 2 mm long, apical appendage acute, base rounded. Styles purple, 5-7 mm long, branches 1.5-2 mm long. Achenes obovate, ca. 2 mm long, puberulous with twin hairs and capitate glands, 4-5-ribbed, carpopodium absent. Pappus 1-2 mm long in 1 series, frequently deciduous or lacking.

Distribution.

Thailand: Ubon Ratchathani. Laos.

Specimens examined.

Thailand, Ubon Ratchathani, route number 2222, 15°18.59'N, 105°23.15'E, route number 2222, 28 Oct 2007, S. Bunwong 346 (KKU, US). Route number 2112 to Khong Chiam, 16 Sep 2004, R. Pooma, K. Phattanahirankanok & S. Sirimongkol 4737 (AAU).

Diagnostic characters.

Camchaya gracilis is characterized by phyllaries without marginal spines, achenes with 4-5 ribs and leaves that are narrowly elliptic-oblong. This species is included in Camchaya as it shares 6-porate pollen found in no other genera.

Ecology.

Dipterocarp forest, alt. 150 m; flowering October to December.

Vernacular name.

Ao Ra Nid (อรนิช).

Figures

Figure 6. Morphology of Vernonieae in Thailand 2. A Camchaya gracilis B Camchaya loloana C Camchaya loloana var. mukdahanensis D Camchaya pentagona E-F Camchaya spinulifera G Camchaya tenuiflora H-I Camchaya thailandica.