Lednia tumana (Ricker)

Baumann, Richard W. & Kondratieff, Boris C., 2010, The Stonefly Genus Lednia In North America (Plecoptera: Nemouridae), Illiesia 6 (25), pp. 315-327 : 316-319

Lednia tumana (Ricker)

( Figs. 1 -12)

Nemoura (Lednia) tumana Ricker 1952:29 . Holotype ♂, Illinois Natural History Survey: Many Glaciers (sic), Glacier National Park , Montana.

Material examined. CANADA: Alberta: Waterton Lakes National Park, Bertha Brook, below Bertha Lake , 25 August 1976, D.B. Donald, 1♀ ( DBDC) . USA: Montana: Glacier National Park: Glacier Co., Base of Mount Clements , behind moraine, 16 Oct.1997, J. Giersch, 1♂, 11♀ ( UMBS); Clements Drain, high, 6 Oct . 1998, J. Giersch, 4 larvae ( UMBS) ; Clements Drain , low, 15 July 1998, J. Giersch, 3 larvae ( UMBS) ; Small glacial stream, 5,500’ , Many Glacier, 24 August 1946, W.E. Ricker, 1♂ (Holotype), 1♀ (Allotype) ( INHS) ; same locality, 24 August 1946, W.E. Ricker, 1♂ (Paratype) ( CNCI) ; Reynolds Creek , headwaters, above boardwalk bridge, Logan Pass, 6 Sept. 1997, J. Giersch, 1♂, 2♀ ( UMBS) ; Reynolds , high, Logan Pass, 10 Sept. 1997, J. Giersch, 3♂, 9♀ ( UMBS) ; Reynolds Spring , Logan Pass, 14 Sept. 1997, J. Giersch, 5♂, 7♀ ( BYUC, UMBS) ; Reynolds , high, Logan Pass, 17 July 1998, J. Giersch, 4 larvae ( UMBS) ; Below Reynolds Pass , 28 July 1998, J. Giersch, 47 larvae ( UMBS) ; Reynolds Spring , Logan Pass, 29 July 1998, J. Giersch, 100 larvae ( UMBS) ; South Face Reynolds , 4 August 1998, J. Giersch, ( UMBS) ; Siyeh Bottom , 7 July 1998, J. Giersch, 2 larvae ( UMBS) . Flathead Co., Hidden Lake overlook pond, Logan Pass, 18 Sept. 1997 J. Giersch, 1♀ ( UMBS) ; Logan Creek , Logan Pass, 6 August 1966, A.R. Gaufin, 1♂, 3♀ ( BYUC) ; Logan Creek , Logan Pass, 5 Sept. 1972, R.A. Haick & D.S. Potter, 1♂, 1♀ ( USNM) ; Logan Creek , Logan Pass, 22 July 1979, K.W. Stewart, R.W. Baumann & B.P. Stark, 8 larvae ( BYUC) ; Logan Creek , Logan Pass, 11 August 1979, J. Fraley, 4♂, 1♀, 1 larva ( UNTC) ; Logan Creek , springhead, 20 July 1998, J. Giersch, 117 larvae ( UMBS) ; Logan Creek , springhead, 3 August 1998, J. Giersch, 29 larvae ( UMBS) ; Logan Creek , springhead, 29 Sept. 1998, J. Giersch, 211 larvae ( UMBS) ; Logan Creek , near water tank, Logan Pass, 20 July 1998, J. Giersch, 2 larvae ( UMBS) ; Logan Creek , near water tank, Logan Pass, 3 August 1998, J. Giersch, 1♀ ( UMBS) ; Logan Creek , near water tank, Logan Pass, 29 Sept. 1998, J. Giersch, 1 larva ( UMBS) ; Mineral Creek , headwaters at East Fork, below Highline Trail, at stream reemergence, 28 August 1997, J. Giersch, 1♂, 5♀, 13 larvae ( BYUC, UMBS) .

Additional material ( L. tumana ?): Wyoming: Teton Co. headwaters, South Fork Darby Creek, Wind Cave , southeast of Driggs , 14 August 2008, R.G. Call, 1♀ ( BYUC) ; Same locality, 11 July 2009, R.G. Call, 1♀ ( BYUC) ; Same locality, 26 July 2010, R.G. Call, 6 larvae ( BYUC) .

Male. Macropterous, wings hyaline with darker veins near cord. Length of body 4.5-5.5 mm, color dark brown, anterior abdominal segments lightly sclerotized. Epiproct bilaterally symmetrical, with dorsal groove that extends from tip to near base ( Figs. 2 , 6, 7); ventral sclerite flat, darkly sclerotized, bearing few stout ventral spines apically, apex narrowly pointed and extending beyond apex of dorsal sclerite, tip with dorsal groove terminating in an arrowhead-like point ( Fig. 7 ); dorsal sclerite with broad, bare base, consisting of two large triangular lobes laterally, narrowing gradually toward apex, anterior portion bearing paired patches of large, equally sized, backward directed spines on dorsolateral margins ( Figs. 1-2 , 5-6, 8-9). Hypoproct sclerotized, broad at base, becoming gradually narrower towards apex, vesicle absent ( Fig. 11 ). Paraprocts with two lobes, inner lobe long, thin and darkly sclerotized, tip sharply pointed; outer lobe short, broadly rounded, lightly sclertotized bearing numerous hairs ( Fig. 11 ). Tergum ten with median anterior groove, bordered by paired mace-like prongs, which approach tip of epiproct ( Figs. 1, 2 , 8, 10).

Female. General color of body and wings similar to male. Length 5.5-7.0 mm. Sternum seven with broadly rounded, lightly sclerotized lobe posteriormedially. Subgenital plate covering most of sternum eight, darkly sclerotized, bearing sparse long hairs, base broad, tapering gradually to narrowly rounded apex that approaches junction of sternum nine, sometimes with small slit at tip, lateral posterior margins of sternum eight bearing nearly rounded, sclerotized lobes on either side of subgenital plate apex ( Figs. 4 , 12).

Larva. Gills absent. Length: mature male 4.5-5.5 mm, mature female 5.5-6.5 mm. Dorsal surface of body with few small spines, except at anterior margins of thoracic and anterior abdominal segments. Legs bearing numerous small spines, 3-5 larger spines present on dorsal margins of femora. Female larvae exhibiting large, truncate, plate-like lobe on dorsomedian margin of sternum eight. Cerci with whorls of spines on posterior margins of segments and sparse, small intercalary spinules present on all segments. For a more detailed description see Baumann and Stewart (1980).

Figures

Figs. 1-3. Lednia tumana male genitalia: 1. Dorsal. 2. epiproct, dorsal. 3. lateral. Fig. 4. Lednia tumana female terminalia, ventral.

Figs.5-11. Lednia tumana male genitalia: 5. terminalia, dorsal. 6. epiproct, dorsal. 7. epiproct apex, dorsal. 8. terminalia, lateral. 9. epiproct, lateral. 10. prong on tergum 10. 11. terminalia, ventral. 12. female terminalia, ventral.