Cryptolophocolea pallida (Mitt.) L.Söderstr. 2013

Söderström, Lars, Váňa, Jiŕí, Crandall-Stotler, Barbara, Stotler, Raymond E., Hagborg, Anders & Konrat, Matt Von, 2013, Notes on Early Land Plants Today. 43. New combinations in Lophocoleaceae (Marchantiophyta), Phytotaxa 112 (1), pp. 18-32 : 22

Cryptolophocolea pallida (Mitt.) L.Söderstr. , comb. nov.

Basionym:— Lophocolea pallida Mitt. in Hooker, Bot. Antarct. Voy. II (Fl. Nov.-Zel. 2): 135, 1854 ( Mitten 1854).

Type:— NEW ZEALAND. " Northern Island, Auckland, Sinclair " ( NY) .

Ξ Chiloscyphus pallidus (Mitt.) J.J.Engel et R.M.Schust. , Nova Hedwigia 39: 420, 1984 [1985] ( Engel & Schuster 1985).

= Jungermannia multipenna Hook. f. et Taylor, London J. Bot. 4: 81, 1845, nom. illeg. (ICN Art. 53.1; hom. illeg. non Hook.f. et Taylor 1844: 381) ( Hooker & Taylor 1845), syn. fide Mitten (1854: 135). Ξ Chiloscyphus multipennus J.J.Engel et R.M.Schust. , Nova Hedwigia 39: 419, 1984 [1985] ( Engel & Schuster 1985).

Note:—When Hooker & Taylor (1844) first described Jungermannia multipenna , they inadvertently repeated the diagnosis for Jungermannia intortifolia Hooker & Taylor (1844: 374) , which is the basionym of Isotachis intortifolia (Hook.f. et Taylor) Gottsche (1864: 121; see Hooker & Taylor 1845: 81 where this error was first discussed). This makes Jungermannia multipenna 1844 (based on Campbell I. material) a superfluous name (nom. illeg. ICN Art. 52.1) since it was based on the same type as Jungermannia intortifolia . Aware of their error, the following year Hooker and Taylor published Jungermannia multipenna Hooker & Taylor (1845: 81) , based upon Auckland Is. material, but this is also illegitimate as a later homonym (ICN Art. 53.1) since Jungermannia multipenna 1844 had already been effectively published. When Gottsche et al. (1845) transferred Jungermannia multipenna to Lophocolea they cited the 1844 publication, technically making this name a synonym to Isotachis intortifolia . In Gottsche et al. (1847) they tried to correct it with a new combination citing the 1845 basionym, but as the earlier Lophocolea multipenna 1845 was effectively published, this later name also became an illegitimate homonym. In 1854 Mitten named Lophocolea pallida to include the nom. illeg. "[ Jungermannia ] Lophocolea multipenna Hook. fil et Tayl. Flor Antarct. p.155 " as a synonym, noting that the Auckland Island specimen in Hooker's herbarium is a mix of vegetative shoots of Leioscyphus decipiens and a fertile shoot of Lophocolea pallida , and that the 1845 description of Jungermannia multipenna was compiled from both taxa. When Engel & Schuster (1985) transferred Jungermannia multipenna 1845 to Chiloscyphus , it could have been considered a valid nom. nov. for all illegitimate names based on Jungermannia multipenna 1845 following ICN Art. 58.1 except for the fact that another epithet is available that has priority at that rank, namely Lophocolea pallida Mitt. When Engel (1991b) selected a lectotype for Jungermannia multipenna 1845 of Hooker plants from Auckland Is., he also reduced L. pallida to it, and in fact, all of the illustrations for Chiloscyphus multipennus used in his publication were from the type of Lophocolea pallida . Based on ICN Art. 58.1, this earlier epithet is the correct name for this taxon and is the basionym for the new combination above.

= Chiloscyphus involucratus Colenso, Trans. & Proc. New Zealand Inst. 22: 455, 1890 ( Colenso 1890) " involucrata ", syn. fide Engel (2010: 137).

= Lophocolea erectifolia Steph., J. Linn. Soc., Bot. 29: 269, 1893 [1892] ( Stephani 1892), syn. fide Hodgson (1953: 345).?= Chiloscyphus erectifolius Steph., Bull. Herb. Boissier sér. 2, 8: 147 (Spec. Hep. 3: 263), 1908 ( Stephani 1908), nom. inval. (ICN Art. 38.1(a); no description and ICN Art. 41.5; no basionym cited). Note:—The name “ Chiloscyphus erectifolius St. ” was never validly published by Stephani but appears only in a list of taxa that he associated with the genus Lophocolea .