Monoplex comptus (A. Adams, 1855 )

Sacchetti, Claudia, Landau, Bernard & Ávila, Sérgio P., 2023, The Lower Pliocene marine gastropods of Santa Maria Island, Azores: Taxonomy and palaeobiogeographic implications, Zootaxa 5295 (1), pp. 1-150 : 61-62

Monoplex comptus (A. Adams, 1855)

Plate 4 D 1 -D 2

* Triton comptus A. Adams 1855: 312 .

Tritonium secans Bronn in Reiss 1862: 28, pl, 1, fig.10.

Tritonium secans Bronn — Mayer 1864: 72, pl. 7, fig. 52.

Triton ridleyi E. A. Smith 1890: 489 , pl. 30, fig. 1.

Lampusia gracile (Reeve) — Dall 1889: 227, pl. 29, fig. 2 (non Triton gracilis Reeve, 1844 = Triton pfeifferianus Reeve, 1844).

Cymatium comptum (A. Adams, 1855) — Yen 1942: 215, pl. 18, fig. 112.

Cymatium (Septa) gemmatum (Reeve, 1844) — Clench & Turner 1957: 222, pl. 110, fig. 2, pl. 113, fig. 6, pl. 125, fig. 1-2 (non Triton gemmatus Reeve, 1844 ).

Cymatium (Septa) vespaceum ( Lamarck, 1822) — Warmke & Abbott 1961: 101, pl. 18b (non Triton vespaceum Lamarck, 1822) .

Cymatium (Septa) vespaceum ( Lamarck, 1822) — Kilias 1973: 193, fig. 139 (non Triton vespaceum Lamarck, 1822 ).

Cymatium (Septa) vespaceum ( Lamarck, 1822) — Abbott 1974: 163, fig. 1754 (non Triton vespaceum Lamarck, 1822 ).

Cymatium (Septa) vespaceum ( Lamarck, 1822) — Rios 1975: 79, pl. 79, fig. 322 (non Triton vespaceum Lamarck, 1822 ).

Cymatium vespaceum ( Lamarck, 1822) — Hinton 1978: 30, fig. 9 (non Triton vespaceum Lamarck, 1822 ).

Cymatium (Septa) vespaceum ( Lamarck, 1822) — Kay 1979: 223, fig. 79E (not 79F) (non Triton vespaceum Lamarck, 1822 ).

Cymatium (Septa) vespaceum ( Lamarck, 1822) — Saunders 1980: 5, upper fig. (non Triton vespaceum Lamarck, 1822 ).

Cymatium (Septa) vespaceum ( Lamarck, 1822) — Coelho et al. 1981: 124, fig. 9 (non Triton vespaceum Lamarck, 1822 ).

Cymatium vespaceum ( Lamarck, 1822) — Garcia-Talavera 1983: 108, pl. 5, fig. 2 (non Triton vespaceum Lamarck, 1822 ).

Cymatium (Septa) comptum . (A. Adams, 1855)— Rios 1985: 75, pl. 27, fig. 329.

Cymatium (Septa) comptum (A. Adams, 1855) — Springsteen & Leobrera 1986: 113, pl. 31, fig. 3a-b.

Cymatium (Turritriton) comptum (A. Adams, 1855) — Garcia-Talavera 1987: 253, pl. 2, fig. 5.

Cymatium (Turritriton) comptum (A. Adams, 1855) — Lai 1989: 125, fig. 44.

Cymatium (Turritriton) comptum (A. Adams, 1855) — Henning & Hemmen 1993: 99, pl. 21, fig. 5.

Cymatium (Septa) comptum (A. Adams, 1855) — Rios 1994: 89, pl. 29, fig. 345.

Cymatium comptum (A. Adams, 1855) —Kubo, in Kubo & Kurozumi 1995: 75, fig. 9.

Cymatium vespaceum ( Lamarck, 1822) — Wellens & Wils 1997: 79, fig. 14 (non Triton vespaceum Lamarck, 1822 ).

Cymatium (Monoplex) comptum (A. Adams, 1855) — Beu 1998b: 86, fig. 23i, 26a-k.

Cymatium (Monoplex) comptum (A. Adams, 1855) — Beu 1999: 21, fig. 35.

Cymatium (Monoplex) comptum (A. Adams, 1855) — Okutani 2000: 287, pl. 142, fig. 11.

Cymatium (Turritriton) comptum (A. Adams, 1855) — Redfern 2001: 62, pl. 31, fig. 263A, B, pl. 109, fig. 263C.

Cymatium comptum amphiatlanticum (A. Adams, 1855) — Garcia-Talavera & de Vera 2003: 202, fig. 1-5.

Cymatium (Monoplex) comptum (A. Adams, 1855) — Zhang & Ma 2004: 147, text-fig. 91.

Cymatium (Monoplex) comptum (A. Adams, 1855) —Beu & Segers in Poppe 2008: 644, pl. 267, fig. 1, 3, 4, 7, 8.

Monoplex comptus (A. Adams, 1855) — Beu 2010: 149, pl. 36, figs. 10, 14.

Cymatium comptum (A. Adams, 1855) —Hernández et al. 2011: 172, fig. 53L.

Santa Maria material examined. Maximum height 25.0 mm, width 14 mm. Two fragments from Pinheiros, Santa Maria Island, Azores, Touril Complex, Lower Pliocene .

Discussion. Tritonium secans Bronn, 1862 was based on four fragments, two from Pinheiros, one from Feteirinhas (= Ponta das Salinas) and one from Cré. No further material is available, but the figure undoubtedly represents a Monoplex species, which in our opinion is M. comptum (A. Adams, 1855) . The specimen illustrated by Bronn in Reiss (1862: pl, 1, fig.10; refigured pl. 4, fig. 4 D 1) and by Mayer (1864: pl. 7, fig. 52; refigured pl. 4, fig. 4 D 2) is undoubtedly the same specimen, but the illustrator has highlighted the primary spiral cords much more strongly in Mayer’s illustration. For full discussion and comparison with related congeners, see Beu (2010: 149). This species and Aspa marginata ( Gmelin, 1791) are the two amphiatlantic species present in the Azorean Pliocene assemblages.

Distribution. Lower Pliocene: eastern Atlantic, Santa Maria Island, Azores (Bronn in Reiss 1862; Mayer 1864). Lower Pleistocene: eastern Atlantic, Moín formation, Costa Rica. Pleistocene or Holocene: Atlantic, Panama; Red Sea, Egypt (NHMW coll.). Present-day: western Atlantic, Bermuda, Carolinian biogeographic province, Gulf of Mexico, Caribbean to Brasil; eastern Atlantic, Canary Islands, Indian Ocean, and Red Sea (all references from Beu 2010).

Figures

PLATE 1. A. Danilia pterostomus (Bronn, 1861), height 5.0 mm, width 4.0 mm, Forno da Cré (figure taken from Bronn in Hartung, 1861, pl. 19, fig. 2). B1-B2. Emarginula octaviana Coen, 1939, DBUA-F 487-1, height 1.8 mm, diameter 3.0 mm, Ponta do Castelo. C. Bolma cf. rugosa (Linné, 1767), DBUA-F 471-3, diameter 4.1 mm, Ponta do Castelo. D1-D5. Turbo hartungi Bronn 1861, DBUA-F 1295-1, height 9.0 mm, width 8.0 mm, Malbusca (east cave). E1-E3. Nerita cf. funata Dujardin, 1837, DBUA-F 917-1, height 4.0 mm, width 6.0 mm, Ponta do Castelo. F1-F2. Thericium crenulosum (Bronn, 1862), height 10.0 mm, width 6.0 mm, Ponta do Norte lighthouse, LNEG collection (unnumbered). G1-G2. Thericium hartungi (Mayer, 1862), height 29.0 mm, width 13.0 mm, Pinheiros (figure taken from Bronn in Reiss, 1862, pl. 1, fig. 11). H1-H2. Thericium incultum (Mayer, 1862), DBUA-F 1034-2, height 23.0 mm, width 10.0 mm, Macela viewpoint. I1-I2. Thericium incultum (Mayer, 1862), DBUA-F 1027-1, height 29.0 mm, width 11.0 mm, Malbusca II. J1-J2, K1-K2. Bittium miradouroense nov. sp., Holotype DBUAF 1034-5, height 6.8 mm, width 1.9 mm (J); Paratype DBUA-F 1034-6(1), height 5.3 mm, width 1.9 mm (K), Macela viewpoint. L1-L2, M1-M2. Bittium nov. sp., DBUA-F 488-2, height 4.5 mm, width 1.8 mm (L), height 4.2 mm, width 1.7 mm (M), Ponta do Castelo.

PLATE 2. A1-A2. Bittium nanum (Mayer, 1864), DBUA-F 1296-2, height 4.9 mm, width 2.0 mm, Malbusca. B. Tenagodus (Tenagodus) obtusus (Schumacher, 1817), DBUA-F 1283-2, height 12.0 mm, width 11.0 mm, Ponta do Castelo. C. Epitonium sp., DBUA-F 931-1, height 4.0 mm, width 2.8 mm, Pedra-que-pica. D1-D3. Janthina krejcii, SMF10135a, height 25.0 mm, diameter 32.0 mm, Macela viewpoint (pictures from Beu, 2017, fig. 26). E1-E3. Janthina typica (Bronn, 1861), DBUA-F-428, height 19.0 mm, diameter 21.0 mm, Ponta do Castelo, (pictures from Beu, 2017, fig. 25). F.Cheilea equestris (Linnaeus, 1758), DBUA-F 1283-1, diameter 30.0 mm, Ponta do Castelo. G1-G3. Hipponix sulcatus (Borson, 1820), DBUA-F 937-1, height 5.3 mm, diameter 14.0 mm, Ponta do Castelo. H1-H2. Euspira atlantica (Mayer, 1862), height 16.0 mm, diameter 19.0 mm, Ponta do Norte lighthouse, LNEG collection (unnumbered). I. Triphorid s.l., DBUA-F 471-5, height 3.2 mm, width 1.3 mm, Ponta do Castelo. J1-J2. Cerithiopsis cf. tubercularis (Montagu, 1803), DBUA-F 1023-1, height 3.0 mm, width 1.2 mm, Ponta do Castelo. K. Petaloconchus intortus (Lamarck, 1818), DBUA-F 946-2, height 13.0 mm, width 6.0 mm, Ponta dos Cedros. L. Thylacodes arenarius (Linnaeus, 1758), DBUA-F 869-1, height 33.0 mm, width 24.0 mm, Ponta dos Cedros. M1-M3. Alvania cancellata (da Costa, 1778), DBUA-F 1296-3, height 4.0 mm, width 2.2 mm, Malbusca.

PLATE 4. A. Tethystrombus coronatus (Defrance, 1827), CIADP-F 32, height 119.1 mm, width 107.2 mm, Figueiral. B. Talisman scrobilator (Linnaeus, 1758), Ponta dos Cedros, size not known, LNEG collection (unnumbered, specimen not seen, part of a photograph database). C. Aspa marginata (Gmelin, 1791), DBUA-F 1428-1, height 28.0 mm, width 20.0 mm, Ponta dos Cedros. D1-D2. Monoplex comptus (A. Adams, 1855) height 25.0 mm, width 14.0 mm, Pinheiros (figure D1 taken from Bronn in Reiss, 1862, pl, 1, fig.10 and figure D2 from Mayer, 1864, pl. 7, fig. 52). E1-E3. Distorsio mcgintyi Emerson & Puffer, 1953, DBUA-F 1189-1, height 31.0 mm, width 18.0 mm, Pedra-que-pica. F. Xenophora sp., height 70.0 mm, diameter 15.0 mm, unknown locality, LNEG collection (unnumbered). G1-G3. Gibberula vignali Dautzenberg & Fischer, 1896, DBUA-F 1044-1, height 2.8 mm, width 1.6 mm, Pedra-que-pica. H1-H2. Cancellaria parcestriata Bronn, 1862, height 10.0 mm, width 6.5 mm, Pinheiros (figure H1 taken from Bronn in Reiss (1862), pl. 1, fig. 7 and figure H2 from Mayer (1864), pl. 7, fig. 51). I1-I2. Costoanachis semicostata (Sacco, 1890), DBUA-F 1034-4, height 6.9 mm, width 3.0 mm, Macela viewpoint. J. Costoanachis sp., DBUA-F 1059, height 2.7 mm, width 1.6 mm, Baía do Raposo. K. Angustifusus longiroster (Brocchi, 1814), height 36.0 mm, width 18.1 mm, unknown locality, LNEG collection (unnumbered). L. Fusinus nov. sp.?, height 51.0 mm, width 24.0 mm, Ponta do Norte lighthouse, LNEG collection (unnumbered).