Neoacanthoneura goniata Kameneva

Kameneva, Elena P., 2012, Revision of Aciuroides Hendel and Neoacanthoneura Hendel (Diptera: Ulidiidae: Pterocallini), Zootaxa 3227, pp. 1-33 : 23-24

publication ID

https://doi.org/ 10.5281/zenodo.280302

DOI

https://doi.org/10.5281/zenodo.6165886

persistent identifier

https://treatment.plazi.org/id/A72387FD-FFC0-FFAD-FF07-FF16FAC74FA1

treatment provided by

Plazi

scientific name

Neoacanthoneura goniata Kameneva
status

sp. nov.

Neoacanthoneura goniata Kameneva , new species

( Figs. 35–36 View FIGURES 27 – 40 , 69–75 View FIGURES 69 – 75 )

Type material. Holotype ♂: TRINIDAD, Simla / Arima Valley , 6–12.II.1966, S.S. & W.D.Duckworth ( USNM) . Paratypes: TRINIDAD: ♂, Simla / Arima Valley , 6–12.II.1966, S.S. & W.D.Duckworth ; ♂, W.[est] I.[ndies], Simla ,, 1–15.VII.1964, J.M.Capriles ( USNM) ; ♂, W.[est] I.[ndies], Arima Valley , 800–1200ft., 10–22.II.1964, J.G.Rozen & P.Wygodzinsky Collectors ( AMNH) ; 3♂, ♀, Arima, Blanchisseuse Rd. , 2000’, 28–31.XII.1981, Morton S. Adams ( AMNH, SIZK) ; 2♂, same but 3–9.I.1982, Morton S. Adams; ♂, ♀, same but 12–14.I.1982; ♂, same but 28–31.I.1982, Morton S. Adams ( AMNH) ; TOBAGO: ♀, Goldborough , 23.II–24.III.1994, M.J.Sommeijer ( ZMAN) .

Diagnosis. Neoacanthoneura goniata is similar to N. chiroptera and N. steyskali in having wing pattern with the discal crossband and the apical spot joined, differing by the costal vein in the male slightly produced anteriorly, forming blunt angle at crossvein dm-cu level.

Description. Male. Head length: height: width ratio 1: 1.5: 1.7, with compound eye 1.6 times as high as long and frons 1.7 times as long as wide. Head setae as in N. chiroptera ; coloration and appendages as in N. chiroptera . Thorax dark brown with partly yellowish, densely yellow microtrichose pleura, postpronotal lobes and antepronotum and bordering part of mesonotal scutum yellow. Legs yellow with black setae and setulae, tarsi mostly brown.

Wing ( Figs. 35–36 View FIGURES 27 – 40 ) with pattern as described for N. chiroptera female, except long cuneiform subbasal crossband and short brown bar posterior of pterostigma base fused along R4+5 vein. In larger ♂, costal vein conspicuously widened between R1 and R4+5 apices, forming anteriorly produced blunt angle at dm-cu level, with ventral row of setulae with 20–25 thick dentate setulae at R1 apex level, in smaller ♂ costal vein non-modified and subbasal cuneiform crossband widely or entirely fused with short bar distal of it leaving no hyaline window between them. Pterostigma narrow, as long as costal cell. Vein R1 setulose on apical half only, mostly subparallel to costa. Vein R2+3 undulate. Vein M ratio m3: m2: m4 = 1: 7.1–12: 7–11. Vein M ratio m3: m2: m4 = 1: 11–12: 10–11.Wing length 1.5–2.4 mm.

Male abdomen and epandrium as in N. chiroptera ; epandrium ( Figs. 69–71 View FIGURES 69 – 75 ) brown, moderately long, parallelsided, with brown setulae and very short surstyli divergent at 120–150º, each with 2 subapical prensisetae. Phallus ( Fig. 71 View FIGURES 69 – 75 ) moderately long, with sparse and fine, poorly recognizable seta-like scales.

Female similar to male, but wing venation less modified, similar to that in smaller ♂ (as on Fig. 36 View FIGURES 27 – 40 ), R1 setulose only along pterostigma, bare proximally of Sc apex level; pterostigma 0.9–1.2 times as long as costal cell, wing pattern as in N. brachioptera , except subbasal triangle as in N. chiroptera male (with hyaline window in it) or cuneiform (without hyaline window); vein R2+3 slightly undulate to almost straight in small specimens; vein M ratio m3: m2: m4 = 1: 8–11: 7–9; wing length 2.6–3.5 mm; thorax, abdomen and legs as described for N. apicalis , except mid and hind femora sometimes yellow; eversible membrane ( Fig. 72 View FIGURES 69 – 75 ) with 2 pairs of long taeniae 0.5 times as long as whole membrane, without spine-like scales, posterior half of membrane uniformly covered with minute multidentate scales. Aculeus 4.6 times as long as wide, with blunt cercal unit ( Fig. 73 View FIGURES 69 – 75 ). Ventral receptacle ( Fig. 74 View FIGURES 69 – 75 ), simple, with sclerotized apical “cap” and slightly folded or knobby basal “peduncle”. 3 spermathecae ( Fig. 75 View FIGURES 69 – 75 ) similar in size, spherical, brown.

Etymology. The name is from the Greek gonios, angle, reflecting the shape of the male wing.

Remarks. Four smaller male specimens (WL=1.5–2.0) collected together with typically larger ♂ (WL=2.2– 2.6) have more reduced wing pattern ( Fig. 36 View FIGURES 27 – 40 ): apical crescent spot isolated from discal crossband at its anterobasal margin, with blurred posterior angle; hyaline areas of wing mostly gray microtrichose; costal vein evenly arcuate, with ventral row of setulae at R1 apex level inconspicuously thickened; costa neither flattened, nor widened; vein R2+3 slightly undulate, and vein M ratio m3: m2: m4 = 1: 7.1: 7.7 (1: 11–12: 10–11 in larger specimens). Larger range of variability in wing measurements and pattern can also be expected in other species of the genus.

USNM

Smithsonian Institution, National Museum of Natural History

AMNH

American Museum of Natural History

SIZK

Schmaulhausen Institute of Zoology

ZMAN

Instituut voor Taxonomische Zoologie, Zoologisch Museum

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Diptera

Family

Ulidiidae

Genus

Neoacanthoneura

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