Omus californicus californicus Eschscholtz, 1829

Bousquet, Yves, 2012, Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico, ZooKeys 245, pp. 1-1722 : 201-206

Omus californicus californicus Eschscholtz, 1829

Omus californicus Eschscholtz, 1829: 3. Type locality: "Cabo de los Reyes [Marin County], Californien" (original citation). Syntype(s) location unknown (possibly in ZMMU).

Omus xanti LeConte, 1859a: 69. Type locality: "Fort Tejon [Kern County, California]" (original citation). Holotype [by monotypy] (♂) in MCZ [# 23592]. Synonymy established by Horn (1903a: 196). Etymology. The species name was proposed for John Xantus de Vesey [1825-1894], a Hungarian exile who worked in the United States as a bookseller, druggist, teacher, and hospital steward. He became over the years a gifted collector of natural history specimens. He eventually returned to his native Hungary and served as Director of the Zoological Garden of Budapest and as curator of ethnography at the Hungarian National Museum.

Omus laevis G.H. Horn, 1867a: 394. Type locality: "high Sierras near the head waters of King’s and Tulé rivers [California]" (original citation), restricted to "southeast Fresno County and East Tulare County" by Ward (1982: 59). Lectotype (♂), designated by Ward (1982: 59), in MCZ [# 35314]. Synonymy established by Bousquet and Larochelle (1993: 52) based on Leffler (1979a: 209) unpublished thesis.

Omus lecontei G.H. Horn, 1872b: 143. Type locality: "near Monterey [Monterey County], California" (original citation). Lectotype (♂), designated by Ward (1982: 59), in MCZ [# 58]. Synonymy established by Bousquet and Larochelle (1993: 52) based on Leffler (1979a: 209) unpublished thesis. Note. This form is considered a valid subspecies of Omus californicus Eschscholtz by Knisley and Haines (2010).

Omus sequoiarum Crotch, 1874b: 73. Type locality: "Calaveras in the Sierra Nevada" (original citation). Syntype(s) [20 originally cited] in MCZ [# 57] and AMNH [# 444]. Synonymy established by Bousquet and Larochelle (1993: 52) based on Leffler (1979a: 209) unpublished thesis.

Omus edwardsii Crotch, 1874b: 73. Type locality: "Lake Tahoe [Placer County, California]" (original citation). Syntype(s) [6 originally cited] in MCZ [# 56] and AMNH [# 445] (Grossbeck 1912: 360). Synonymy established by Bousquet and Larochelle (1993: 52) based on Leffler (1979a: 209) unpublished thesis. Etymology. The specific name honors Henry Edwards [1830-1891], a stage actor by profession and enthusiastic collector of Lepidoptera and other insects. Born in England, Edwards travelled to Australia, South and Central America, Mexico, and from 1865 to 1877 resided in San Francisco before moving to eastern United States. In 1891, his widow put Edwards’ collection, which consisted of about 300,000 specimens of all orders, for sale. The American Museum of Natural History acquired the collection as well as his correspondence and notes.

Omus hornii LeConte, 1875a: 157. Type locality: "Yosemite, California" (original citation). Holotype [by monotypy] (♀) in MCZ [# 46]. Synonymy established, under the name Omus californicus sequoiarum Crotch, by Horn (1930: 78).

Omus hornianus W. Horn, 1892a: 91. Type locality: «California?» (original citation). Holotype [by monotypy] (♂) in DEI ( Döbler 1973: 384). Synonymy established by Horn (1902b: 388).

Omus montanus Casey, 1897: 290. Type locality: "Placer Co[unty], California" (original citation). Two syntypes in USNM [# 45842]. Synonymy established by, under the name Omus californicus edwardsii Crotch, by Horn (1905: 54).

Omus lugubris Casey, 1897: 290. Type locality: «California» (original citation). One syntype [2 ♂ originally cited] in USNM [# 45859]. Synonymy established, under the name Omus edwardsii Crotch, by Leng (1902: 102).

Omus punctifrons Casey, 1897: 291. Type locality: «California» (original citation). Holotype [by monotypy] (♀) in USNM [# 45861]. Synonymy established, under the name Omus sequoiarum Crotch, by Leng (1902: 102).

Omus confluens Casey, 1897: 291. Type locality: «California» (original citation). Holotype [by monotypy] (♀) in USNM [# 45863]. Synonymy established, under the name Omus sequoiarum Crotch, by Leng (1902: 102).

Omus sculptilis Casey, 1897: 292. Type locality: "coast regions north of San Francisco, California" (original citation). Holotype [by monotypy] (♂) in USNM [# 45824]. Synonymy established by Horn (1902b: 388).

Omus elongatus Casey, 1897: 293. Type locality: "near San Francisco, California" (original citation). One syntype in USNM [# 45852]. Synonymy established, under the name Omus lecontei Horn, by Horn (1903a: 188, 198).

Omus californicus fuchsi W. Horn, 1903a: 188. Type locality: «Küste von Californien, südlich von San Francisco gefangen: wahrscheinlich in oder in der Nähe von Monterey Co. [= coast of California, south of San Francisco, probably in or close to Monterey County]" (original citation). One syntype in CMNH. Synonymy established, under the name Omus californicus lecontei Horn, by Boyd (1982: 2).

Omus mimus Casey, 1909: 256. Type locality: "probably northern California" (original citation). One syntype in USNM [# 45819]. Synonymy established with doubt by Horn (1910a: 125).

Omus dunni Casey, 1909: 258. Type locality: "vicinity of San Francisco [California]" (original citation). Two syntypes in USNM [# 45853]. Synonymy established, under the name Omus californicus lecontei Horn, by Horn (1910a: 126).

Omus dunni regularis Casey, 1909: 258. Type locality: "Carmel, Monterey Co[unty] [California]" (original citation). Two syntypes in USNM [# 45854]. Synonymy established, under the name Omus californicus lecontei Horn, by Horn (1910a: 126).

Omus dunni maritimus Casey, 1909: 259. Type locality: "Monterey Co[unty] [California]" (original citation). One syntype in USNM [# 45855]. Synonymy established, under the name Omus californicus lecontei Horn, by Horn (1910a: 126).

Omus cribripennis Casey, 1909: 261. Type locality: "Placerville, El Dorado Co[unty] [California]" (original citation). One syntype in USNM [# 45846]. Synonymy established by Bousquet and Larochelle (1993: 53) based on Leffler (1979a: 210) unpublished thesis.

Omus edwardsi lobatus Casey, 1909: 261. Type locality: "Placer Co[unty] [California]" (original citation). One syntype in USNM [# 45840]. Synonymy established, under the name Omus californicus edwardsii Crotch, by Horn (1910a: 126).

Omus montanus lucidicollis Casey, 1909: 262. Type locality: "Placer Co[unty] [California]" (original citation). Two syntypes in USNM [# 45841]. Synonymy established, under the name Omus californicus edwardsii Crotch, by Horn (1910a: 126).

Omus montanus brunnescens Casey, 1909: 262. Type locality: "Placer Co[unty] [California]" (original citation). Two syntypes in USNM [# 45843]. Synonymy established, under the name Omus californicus edwardsii Crotch, by Horn (1910a: 126).

Omus punctifrons degener Casey, 1909: 263. Type locality: "Sierra Co[unty] [California]" (original citation). One syntype in USNM [# 45865]. Synonymy established, under the name Omus californicus punctifrons Casey, by Horn (1910a: 125).

Omus fraterculus Casey, 1909: 263. Type locality: "Placer Co[unty] [California]" (original citation). Syntype(s) in USNM [# 45862]. Synonymy established, under the name Omus californicus punctifrons Casey, by Horn (1930: 78).

Omus collaris Casey, 1909: 265. Type locality: "Wawona, Mariposa Co[unty] [California]" (original citation). One syntype in USNM [# 45881]. Synonymy established, under the name Omus hornii LeConte, by Horn (1910a: 126).

Omus compositus Casey, 1909: 265. Type locality: "Wawona, Mariposa Co[unty] [California]" (original citation). One syntype in USNM [# 45882]. Synonymy established, under the name Omus californicus hornii LeConte, by Horn (1910a: 126).

Omus tularensis Casey, 1909: 265. Type locality: "Davenport (6,400 feet), Soldiers’ Camp (5,800 feet), and Colony Mill (5,415 feet), Tulare Co[unty] [California]" (original citation). Six syntypes in USNM [# 45885]. Synonymy established, under the name Omus californicus laevis LeConte, by Horn (1910a: 126).

Omus tularensis gracilior Casey, 1909: 266. Type locality: "Tulare Co[unty] [California]" (original citation). Holotype [by monotypy] (♂) in USNM [# 45886]. Synonymy established, under the name Omus californicus laevis LeConte, by Horn (1910a: 126).

Omus lugubris sierricola Casey, 1913: 3. Type locality: «California» (original citation). Holotype [by monotypy] (♂) in USNM [# 45860]. Synonymy established, under the name Omus californicus sequoiarum Crotch, by Horn (1915: 432).

Omus californicus intermedio-pronotalis W. Horn, 1913: 346. Type locality: "Plumas Co[unty] orientalis, Calif[ornia]" (original citation). Two syntypes in MCZ [# 25598] and IRSN. Synonymy established, under the name Omus californicus edwardsii Crotch, by Horn (1930: 79).

Omus californicus nunenmacheri W. Horn, 1913: 347. Type locality: "Lassen Co[unty] centralis, Calif[ornia]" (original citation). Syntype(s) [6 originally cited] in MHNP and MCZ [# 25597]. Synonymy established by Bousquet and Larochelle (1993: 53) based on Leffler’s (1979a: 210) unpublished thesis. Etymology. The subspecific name was proposed for Frederick William Nunenmacher [1870-1946], a successful collector of beetles in western United States and a specialist of Coccinellidae . His coccinellid collection went to the California Academy of Sciences and his general collection to the Field Museum of Natural History.

Omus californicus vermiculatus Casey, 1914: 5. Type locality: "probably near San Francisco, California" (original citation). Three syntypes [3 originally cited] in USNM [# 45823]. Synonymy established by Horn (1915: 443).

Omus pronotalis Casey, 1914: 10. Unjustified emendation of Omus intermediopronotalis Horn, 1913.

Omus sequoiarum longitarsis Casey, 1914: 12. Type locality: "Big Trees, Calaveras Co[unty], California" (original citation). One syntype in USNM [# 45858]. Synonymy established, under the name Omus californicus sequoiarum Crotch, by Horn (1915: 443).

Omus horni temperatus Casey, 1914: 15. Type locality: "Giant Forest, Tulare Co[unty], California" (original citation). One syntype in USNM [# 45866]. Synonymy established, under the name Omus californicus laevis LeConte, by Horn (1915: 443).

Omus tularensis opacellus Casey, 1914: 16. Type locality: "Tulare Co[unty], California" (original citation). One syntype in USNM [# 45889]. Synonymy established, under the name Omus californicus laevis LeConte, by Horn (1915: 443).

Omus levis W. Horn, 1915: 443. Unjustified emendation of Omus laevis Horn, 1867.

Omus shastanicus Casey, 1916: 11. Type locality: "Shasta Co[unty], California" (original citation). Two syntypes in USNM [# 45808]. Synonymy established by Bousquet and Larochelle (1993: 53) based on Leffler (1979a: 211) unpublished thesis.

Omus shastanicus cephalicus Casey, 1916: 11. Type locality: "Shasta Co[unty], California" (original citation). One syntype in USNM [# 45802]. Synonymy established, under the name Omus californicus shastanicus Casey, by Horn (1926: 55).

Omus shastanicus tenuiculus Casey, 1916: 12. Type locality: "Shasta Co[unty], California" (original citation). One syntype in USNM [# 45815]. Synonymy established, under the name Omus californicus shastanicus Casey, by Horn (1926: 55).

Omus semilucens Casey, 1916: 12. Type locality: "San Francisco Co[unty], California" (original citation). One syntype in USNM [# 45834]. Synonymy established by Horn (1926: 54).

Omus semilucens diminuens Casey, 1916: 13. Type locality: "Leona Heights, Alameda Co[unty], California" (original citation). One syntype in USNM [# 45833]. Synonymy established by Horn (1926: 55).

Omus californicus latipennis Casey, 1916: 13. Type locality: "Leona Heights, Alameda Co[unty], California" (original citation). One syntype in USNM [# 45822]. Synonymy established by Horn (1926: 55).

Omus sculptilis opacipennis Casey, 1916: 13. Type locality: "S[ain]t Helena, Napa Co[unty], California" (original citation). One syntype in USNM [# 45817]. Synonymy established by Horn (1926: 55).

Omus lacertus Casey, 1916: 15. Type locality: "Carmel, Monterey Co[unty], California" (original citation). One syntype in USNM [# 45856]. Synonymy established, under the name Omus californicus lecontei Horn, by Horn (1926: 59).

Omus laticollis Casey, 1916: 16. Type locality: "Tuolumne Co[unty], California" (original citation). Two syntypes in USNM [# 45864]. Synonymy established, under the name Omus californicus fuchsi Horn, by Horn (1930: 78).

Omus temperatus difficilis Casey, 1916: 17. Type locality: "Mariposa Co[unty], California" (original citation). One syntype in USNM [# 45867]. Synonymy established, under the name Omus californicus hornii LeConte, by Horn (1926: 57).

Omus temperatus mariposae Casey, 1916: 17. Type locality: "Mariposa Co[unty], California" (original citation). One syntype in USNM [# 45869]. Synonymy established, under the name Omus californicus hornii LeConte, by Horn (1926: 57).

Omus temperatus sparsellus Casey, 1916: 18. Type locality: "Wawona, Mariposa Co[unty], California" (original citation). One syntype in USNM [# 45873]. Synonymy established, under the name Omus californicus hornii LeConte, by Horn (1926: 57).

Omus subsericeus Casey, 1916: 18. Type locality: "Kaweah [Tulare County], California" (original citation). One syntype in USNM [# 45874]. Synonymy established, under the name Omus californicus laevis Horn, by Horn (1926: 57).

Omus collaris antennalis Casey, 1916: 19. Type locality: "Mariposa Co[unty], California" (original citation). Two syntypes in USNM [# 45875]. Synonymy established, under the name Omus californicus laevis Horn, by Horn (1926: 57).

Omus collaris trapezicollis Casey, 1916: 19. Type locality: "Mariposa Co[unty], California" (original citation). One syntype in USNM [# 45872]. Synonymy established, under the name Omus californicus laevis Horn, by Horn (1926: 57).

Omus collaris erraticus Casey, 1916: 20. Type locality: "Tuolumne Co[unty], California" (original citation). One syntype [2 originally cited] in USNM [# 45868]. Synonymy established, under the name Omus californicus laevis Horn, by Horn (1926: 57).

Omus horni brevis Casey, 1916: 21. Type locality: "Tuolumne Co[unty], California" (original citation). One syntype in USNM [# 45883]. Synonymy established, under the name Omus californicus hornii LeConte, by Horn (1926: 57).

Omus horni propinquus Casey, 1916: 21. Type locality: "Mariposa Co[unty], California" (original citation). One syntype in USNM [# 45871]. Synonymy established, under the name Omus californicus hornii LeConte, by Horn (1926: 57).

Omus horni asperatus Casey, 1916: 22. Type locality: "Tuolumne Co[unty], California" (original citation). One syntype in USNM [# 45879]. Synonymy established, under the name Omus californicus hornii LeConte, by Horn (1926: 57).

Omus horni granosus Casey, 1916: 22. Type locality: "Mariposa Co[unty], California" (original citation). One syntype in USNM [# 45880]. Synonymy established, under the name Omus californicus hornii LeConte, by Horn (1926: 57).

Omus horni sinuosus Casey, 1916: 23. Type locality: "Wawona, Mariposa Co[unty], California" (original citation). One syntype in USNM [# 45877]. Synonymy established, under the name Omus californicus hornii LeConte, by Horn (1926: 57).

Omus horni punctatus Casey, 1916: 23. Type locality: "Tuolumne Co[unty], California" (original citation). Four syntypes [4 originally cited] in USNM [# 45878]. Synonymy established, under the name Omus californicus hornii LeConte, by Horn (1926: 57). Note. Hennessey (1990: 467) reported the presence of a syntype of Omus punctatus Casey in SIM.

Omus horni farctus Casey, 1916: 23. Type locality: "Mariposa Co[unty], California" (original citation). One syntype in USNM [# 45884]. Synonymy established, under the name Omus californicus hornii LeConte, by Horn (1926: 57).

Omus marginalis Casey, 1916: 24. Type locality: "Tuolumne Co[unty], California" (original citation). One syntype in USNM [# 45870]. Synonymy established, under the name Omus californicus hornii LeConte, by Horn (1926: 57).

Omus tularensis remissus Casey, 1916: 25. Type locality: "Colony Mill (5400 ft.), Tulare Co[unty], California" (original citation). One syntype in USNM [# 45887]. Synonymy established, under the name Omus californicus laevis Horn, by Horn (1926: 57).

Omus laevis peropacus Casey, 1916: 25. Type locality: "Tulare Co[unty], California" (original citation). One syntype [3 originally cited] in USNM [# 45888]. Synonymy established, under the name Omus californicus laevis Horn, by Horn (1926: 57).

Omus cupreonitens Blaisdell and Reynolds, 1917: 49. Type locality: "shore of Humboldt Bay near Arcata, Humboldt County, California" (original citation). Holotype (♂) in USNM [# 21355]. Synonymy established by Bousquet and Larochelle (1993: 53) based on Leffler (1979a: 212) unpublished thesis.

Omus [cupreonitens] reynoldsi Casey, 1924: 5. Type locality: "Arcata, Humboldt Co[unty], California" (original citation). Holotype [by monotypy] (♂) in USNM [# 45839]. Synonymy established, under the name Omus cupreonitens Blaisdell and Reynolds, by Blaisdell (1925: 80).

Omus [sculptilis] densicollis Casey, 1924: 5. Type locality: "Mendocino Co[unty], California" (original citation). One syntype in USNM [# 45825]. Synonymy established by Horn (1926: 55).

Omus [sculptilis] argutus Casey, 1924: 6. Type locality: "Alameda Co[unty], California" (original citation). One syntype in USNM [# 45826]. Synonymy established by Horn (1930: 78).

Omus [mimus] debiliceps Casey, 1924: 6. Type locality: "San Francisco [San Francisco County], California" (original citation). One syntype in USNM [# 45818]. Synonymy established by Horn (1926: 54).

Omus [mimus] insulsus Casey, 1924: 6. Type locality: "S[an]ta Cruz Co[unty], California" (original citation). Two syntypes in USNM [# 45828]. Synonymy established by Horn (1926: 55).

Omus [mimus] modicus Casey, 1924: 7. Type locality: "Mendocino Co[unty], California" (original citation). Two syntypes [2 ♂ originally cited] in USNM [# 45827]. Synonymy established by Horn (1926: 55).

Omus [mimus] stolidus Casey, 1924: 7. Type locality: "Mendocino Co[unty], California" (original citation). Two syntypes in USNM [# 45829]. Synonymy established by Horn (1926: 54).

Omus [mimus] subparallelus Casey, 1924: 7. Type locality: "S[an]ta Cruz Co[unty], California" (original citation). Holotype [by monotypy] (♂) in USNM [# 45830]. Synonymy established by Horn (1926: 55).

Omus ventricosus Casey, 1924: 8. Type locality: "Mendocino Co[unty], California" (original citation). One syntype in USNM [# 45831]. Synonymy established by Horn (1926: 54).

Omus leachi Casey, 1924: 8. Type locality: "Trinity Co[unty], California" (original citation). Four syntypes [4 originally cited] in USNM [# 45838]. Synonymy established by Horn (1926: 55).

Omus pullatus Casey, 1924: 9. Type locality: "Sonoma Co[unty], California" (original citation). One syntype in USNM [# 45820]. Synonymy established by Horn (1926: 55).

Omus [vermiculatus] pollens Casey, 1924: 9. Type locality: "Marin Co[unty], California" (original citation). One syntype in USNM [# 45821]. Synonymy established by Horn (1926: 55).

Omus [californicus] turbulentus Casey, 1924: 10. Type locality: "Sonoma Co[unty], California" (original citation). One syntype in USNM [# 45835]. Synonymy established by Horn (1926: 55).

Omus [californicus] aethiops Casey, 1924: 10. Type locality: "Shasta Co[unty], California" (original citation). One syntype in USNM [# 45836]. Synonymy established by Horn (1926: 55).

Omus [californicus] sparsus Casey, 1924: 10. Type locality: "S[an]ta Cruz, California" (original citation). One syntype in USNM [# 45837]. Synonymy established by Horn (1926: 55).

Omus [horni] callosus Casey, 1924: 12. Type locality: "Tuolumne Co[unty], California" (original citation). One syntype in USNM [# 45876]. Synonymy established, under the name Omus californicus hornii LeConte, by Horn (1926: 57).

Omus vanlooi Nunenmacher, 1940: 144. Type locality: "Butte County, California" (original citation). Holotype (♂) in CAS [# 8164]. Synonymy established by Bousquet and Larochelle (1993: 53) based on Leffler (1979a: 212) unpublished thesis.

Distribution.

This subspecies, also known as the "California Night-stalking Tiger Beetle", ranges from southwestern Oregon to southern California along the coast and through the Sierra Nevada, at elevation below 900 m near the range of Omus californicus intermedius (Leffler 1979a: 218; Fig. 18).

Records.

USA: CA, OR