Pinodytes tibialis ( Hatch, 1957 ) Peck & Cook, 2011

Peck, Stewart B. & Cook, Joyce, 2011, Systematics, distributions and bionomics of the Catopocerini (eyeless soil fungivore beetles) of North America (Coleoptera: Leiodidae: Catopocerinae) 3077, Zootaxa 3077 (1), pp. 1-118 : 43

Pinodytes tibialis ( Hatch, 1957) , new combination

( Figs. 34 , 310–318 )

Catopocerus tibialis Hatch 1957: 20 .

Type material. Type male in USNM, seen. Type label data: Gold Beach , Curry County, Oregon ; 21.IV.1955, J. Capizzi.

Additional material examined. We examined 90 additional specimens (see Appendix) for a total of 91 specimens.

Distribution. Specimens ( Fig. 310 ) are known only from Curry County in southwestern Oregon.

Diagnosis. Total length 1.40–1.60 mm; greatest width 0.65–0.80 mm. Reddish brown; elongate-oval in shape ( Fig. 34 ). Head. Moderately finely, irregularly punctate; with reticulate microsculpture. Eyes absent. Antenna ( Fig. 311 ) with antennomere 2 longer than 3; antennomere 5 slightly larger than 4 and 6; antennomere 7 clearly larger than 8; antennomeres 9 and 10 lack visible sensory vesicles. Pronotum. Finely punctate, punctures separated by 3– 4 diameters; with reticulate microsculpture. Disc with three pairs of larger punctures, positioned subapically, posteromedially and subbasally. Slightly wider at middle than at base, narrowing to apex; apical margin weakly emarginate, basal margin straight; apical angles rounded, basal angles weakly obtuse. Elytra. Moderately coarsely punctate; a few striae weakly indicated adjacent to suture; punctures joined by fine strioles that create a weakly imbricate pattern basally. Joined elytra slightly wider than pronotum; sides subparallel in basal one-half, narrowing to apex. Legs. Protibia ( Fig. 312 ) narrow at base, apical one-half strongly widened in male, less so in female; two curved spines at apex of outer margin; apical one-half of inner margin with fine, dense spines. Male mesotibia ( Fig. 313 ) with basal two-thirds sinuate, angulate at apical one-third; unmodified in female; strong spines on outer margin in both sexes. Metatibia ( Fig. 314 ) elongate, narrow, straight in both sexes. Metafemur ( Fig. 314 ) slender in both sexes. Male protarsomeres ( Fig. 312 ) bearing elongate setae laterally and thin, colorless, broad phanerae ventrally. Mesotarsomeres without phanerae. Venter. Mesoventrite ( Fig. 318 ) carinate, with a small median tooth; excavation behind transverse carina. Male genitalia. Median lobe of aedeagus ( Figs. 315, 316 ) cylindrical, curved, with flattened apex narrowing to weak rounded lobe. Inverted internal sac ( Fig. 316 ) with elongate sclerotized structures. Parameres ( Figs. 315, 316 ) narrow, about three-fourths length of median lobe; each paramere with one apical and one subapical seta. Spermatheca. Tubular ( Fig. 317 ), curved.

Figures

FIGURE 310. Distributions of species in the Pinodytes cryptophagoides species group in northwestern United States. P.rothi, open circles in northwestern Oregon down Coast Mountains to northern coastal California. P. tibialis, closed circles in southwestern Oregon. P. angulatus, open triangles in northwestern Oregon. P. delnorte, closed squares in northwestern California. P. garibaldi, closed triangle in northwestern Oregon.

FIGURES 311–318. Pinodytes tibialis. 311. Male right antenna, dorsal. 312. Male right protibia and protarsus. 313. Male right mesotibia. 314. Male right metafemur and metatibia. 315. Aedeagus, lateral. 316. Aedeagus, dorsal. 317. Spermatheca. 318. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 317), 0.3 mm (Figs. 315, 316, 318), 0.4 mm (Figs. 311–314).

FIGURES 28–36. Pinodytes habitus photos. 28. P. cryptophagoides, total length 1.88 mm. 29. P. subterraneus, total length 3.48 mm. 30. P. imbricatus, total length 3.20 mm. 31. P. newelli, total length 2.30 mm. 32. P. capizzii, total length 1.64 mm. 33. P.rothi, total length 1.58 mm. 34. P. tibialis, total length 1.60 mm. 35. P. angulatus, total length 1.86 mm. 36. P. delnorte, total length 1.72 mm.