Toechorychus morelus Tedesco, 2013

Tedesco, Anazélia M. & Aguiar, Alexandre P., 2013, <strong> Phylogeny and Revision of <em> Toechorychus </ em> Townes (Hymenoptera, Ichneumonidae, Cryptinae), with descriptions of thirty-five new species </ strong>, Zootaxa 3633 (1), pp. 1-138 : 53-54

publication ID	https://doi.org/ 10.11646/zootaxa.3633.1.1
publication LSID	lsid:zoobank.org:pub:60F0956C-A1F5-44CA-9D14-8C04343AD01F
DOI	https://doi.org/10.5281/zenodo.5261835
persistent identifier	https://treatment.plazi.org/id/03B987AE-FF94-FFF6-0AE1-FEAEFB0EF95C
treatment provided by	Felipe
scientific name	Toechorychus morelus Tedesco
status	sp. nov.

Treatment

Toechorychus morelus Tedesco , sp. nov.

( Figs 18 View FIGURES 16–19 , 47 View FIGURES 45–54. 45 , 71 View FIGURES 70–84 , 103 View FIGURES 97–104. 97–98 , 142 View FIGURES 139–144 , 188 View FIGURES 187–199 )

Description. Holotype FEMALE. Fore wing length 6.18 mm.

Head. Mandible 1.55 as long as basal width, densely pilose; ventral margin slightly projected as crest; dorsal tooth distinctly longer than ventral tooth. Clypeus 2.09 as wide as high, subrectangular, minutely strigulate, apically smooth; apex 1.43 as long as base, truncate; apical margin sharp, medially straight. Supraclypeal area medially strigate, laterally finely punctate, densely pilose, medially slightly prominent; between antennal foramens without V-shaped carina; radicle foveolate. Antenna with 21 flagellomeres; white band starting at flagellomere 5, reaching flagellomere 10; flagellum slender; subapical flagellomeres slightly flattened. Supra-antennal area with stout, median, arched, transverse carina, medially without longitudinal elevation, near antennal sockets not striate; dorsal half medially rugulose. Paraocular area finely punctulate. Vertex rugulose around ocelli, without sulcus; gena and vertex behind ocelli minutely strigulate or with scarce punctures, sparsely pilose; gena in lateral view at level of dorsal portion of occipital carina narrow, ventrally wide; occipital carina stout, dorsally V-shaped, ventrally slightly projected as crest, reaching hypostomal carina at mandible base or nearly so; hypostomal carina slightly projected as crest; malar space 0.94 as long as basal width of mandible.

Mesosoma . Mesosoma in dorsal view 1.86 as long as wide; in lateral view, mesosoma middle width 0.16 mm. Pronotum centrally corrugated, latero-ventrally markedly strigate behind collar, margin near mesopleuron entirely corrugated; pronotal swelling finely rugulose; collar dorso-laterally rounded, distinctly swollen, anteriorly rugulose; epomia stout, restricted to space between posterior margin of collar and pronotal swelling. Mesoscutum sparsely pilose, lobes dorsally punctulate, laterally markedly corrugated, central lobe with sparse coarse punctures, central lobe without longitudinal sulcus or carina; notauli deeply impressed, posteriorly abruptly curved, U-shaped. Scutoscutellar groove markedly corrugated; scutellum 0.96 as long as wide, smooth; scutellar carina incomplete, restricted approximately to anterior 0.3; postscutellum 0.34 as long as wide; hind margin of metanotum with two lateral teeth. Subalar ridge somewhat elongate; mesopleuron entirely markedly strigate, ventrally strigulate; mesopleural groove corrugated; epicnemial carina stout, dorsally curving backwards; dorsal end of epicnemial carina reaching 0.2 of posterior margin of pronotum, behind epicnemial carina with rounded confused-rugulose area; sternaulus medially shallow, interrupted, reaching base of mid coxa, markedly sinuate, moderately impressed; faintly corrugated; scrobe deeply impressed, forming pit. Mesothoracic venter strigulate; median portion of posterior transverse carina of mesothoracic venter short and straight, regularly shaped, without projections. Transverse furrow at base of propodeum laterally 0.22 as long as anterior portion of propodeum, moderately deep and narrow, medially wide, uniformly corrugated; propodeum 1.16 as long as wide medially; anterior margin of propodeum with two faint lateral teeth; anterior portion of propodeum finely strigulate or with sparse coarse punctures, medially with two posteriorly convergent faint longitudinal carinae; lateral longitudinal carina of propodeum indistinct; posterior portion of propodeum confused-rugose, medially without distinctly stout longitudinal carina; behind posterior transverse carina with two divergent longitudinal carinae; anterior transverse carina of propodeum complete, stout, entirely slightly arched forwards; posterior transverse carina represented by crests, present laterally as carina; propodeal spiracle 1.50 as long as wide; pleural carina of propodeum irregular and faint, obsolescent posteriorly, fused with sculpture of lower division of metapleuron and propodeum; lower division of metapleuron strigulate, sparsely pilose, juxtacoxal carina short and stout.

Wings. Fore wing: vein 2+3Rs slightly concave; ramellus absent; vein 1M+Rs entirely slightly sinuous; vein 1cu-a posteriorly slightly curved, arising far from vein 1M+Rs base, angle with vein M+Cu about 90°; vein 2Cu 0.69 as long as vein 2cu-a; vein 4Rs 0.85 as long as vein 4M, slightly sinuous, almost straight; cell 1+2Rs 0.9 as high as pterostigma, pentagonal; vein 2M distinctly longer than vein 3M; vein 3M distinct, 0.50 as long as vein 2M. Hind wing: vein 1Cu 1.43 as long as vein cu-a; vein 2-1A reaching 0.76 of distance to posterior margin.

Legs. Tibia with dense short bristles; hind coxa globose.

Metasoma. T1 0.59 as long as hind femur, with basolateral tooth; anteriorly smooth, posteriorly coriarious; spiracle at anterior 0.54, prominent; postpetiole dorso-anteriorly flat; T2–8 minutely coriarious; centrally almost glabrous, laterally pilose; T2 0.92 as long as wide at apex; apex of T2 1.93 as wide as base. Dorsal valve of ovipositor with nodus indistinct; apex of lower valve with series of teeth; notch present.

Color. Head, mesosoma and metasoma black and yellow (234,216,097). Head: yellow; mandible teeth dorsally, M-shaped mark at ventral half of supraclypeal area, supra-antennal area medially, interocellar area, vertex centrally, and occiput, black; scape ventrally yellow, dorsally dark brown; pedicel and flagellum brown, except for f6–9 dorsally, completely white, and f5 and f10, partially whitish. Mesosoma : black, except for the following bright yellow marks: propleuron, collar laterally, pronotal swelling, longitudinal lanceolate mark at posterior 0.9 of central lobe of mesoscutum, axillary carina, scutellum, postscutellum, both upper and lower division of metapleuron, hind margin of metanotum, two lateral longitudinal marks at propodeum, mesothoracic venter, and two large marks at mesopleuron: anteriorly extending from subalar ridge to entire dorsal margin of sternaulus except around hypoepimeron and just behind epicnemial carina, and at entire length of hypoepimeron, not connected. Legs: mostly orange; all coxae bright yellow with dark brown or black spots; fore coxa anteriorly with apical black spot, posteriorly brownish (114,067,036); mid coxa dorsally with apical dark brown spot; hind coxa with two longitudinal black marks, not basally connected, dorsal one anchor–shaped, reaching apex of coxa, ventral one at basal 0.7; all t5 dark brown. Metasoma: black and yellow; T1 dorsally black, laterally and at posterior 0.2, yellow, with black rounded spot opposite to spiracle; T2–8 black with posterior 0.4 bright yellow; T2 basally with lateral narrow stripe anteriorly to spiracle; S1 black; S2–8 bright yellow, laterally black.

Variation. White band at antenna starting at flagellomere 3. Malar space only ventrally black. Clypeus entirely bright yellow. Orbital band complete, not interrupted at ventral gena. Longitudinal lanceolate mark at central lobe of mesoscutum narrow. Black marks at mesopleuron at hypoepimeron and confused-rugulose area connected. Black mark at confused rugulose area connected with black mark over sternaulus. Longitudinal marks ventrally to sternaulus and laterally to suture of mesothoracic venter large, occupying 0.8 of mesothoracic venter area. Hind coxa ventrally black only at basal 0.8.

Comments. Similar to T. linaresius sp. nov., from which it is isolated by having central lobe of mesoscutum without sulcus or carina (vs. with longitudinal sulcus in T. linaresius sp. nov.); mesopleuron with one black mark extending anteriorly around hypoepimeron and at confused-rugulose area just behind dorsal end of epicnemial carina (vs. with two black marks not connected, dorsally around hypoepimeron, and ventrally at confused-rugulose area; Fig. 19 View FIGURES 16–19 ); sternaulus with wide brown mark (vs. brown mark at sternaulus very narrow, centrally interrupted); pleural carina of propodeum irregular and faint, obsolescent posteriorly, fused with sculpture of propodeum and lower division of metapleuron (vs. complete, moderately stout, as in Fig. 41 View FIGURES 41–44. 41 ); hind leg, except coxa and t5, yellowish (vs. blackish at first trochanter, 0.2 apical of tibia, and 0.5 basal of t1); hind wing vein 1Cu about 1.5 as long as vein cu-a (vs. almost 2 as long as vein cu-a); T1 with faint but distinct basolateral tooth (vs. without basolateral tooth, as in Fig. 54 View FIGURES 45–54. 45 ).

Male. Unknown.

Distribution. Mexico and Colombia. Recorded from four localities ( Fig. 215 View FIGURES 208–215 ). The type locality is Cali, Valle del Cauca (3°25’14”N 76°31’20”W). These records comprise a range of 15°31’ in latitude.

Biology. Unknown.

Material examined. 4 females. Holotype ♀, COLOMBIA: Valle del Cauca, Cali , X.1971, W.Eberhard leg., Toechorychus, Townes det. ( AEIC) . Complete , in good shape . Paratypes: MEXICO: 1 ♀ from Chiapas, San Cristóbal de las Casas , 7087’, 12.VI.1969, B.V.Peterson ( CNCI) . 1 ♀ from Morelos, Cuernavaca , June H.H.S., Godman-Salvin leg. 1904.–1. ( BMNH) . COLOMBIA: 1 ♀ from Valle del Cauca, Peñas Brancas , 1750 m, 10 Km W Cali, very wet premontane forest, 31.I.1975, Malaise trap, R.C.Wilkerson leg. ( FSCA) .