Tropidolaemus subannulatus ( Gray, 1842 )
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publication ID |
https://doi.org/ 10.11646/zootaxa.1644.1.1 |
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publication LSID |
lsid:zoobank.org:pub:D12224EE-5F87-4631-8618-2841D7CBC069 |
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persistent identifier |
https://treatment.plazi.org/id/002D87BA-FFCE-FFF7-FF58-C2FFFBD8FCC9 |
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treatment provided by |
Felipe |
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scientific name |
Tropidolaemus subannulatus ( Gray, 1842 ) |
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Tropidolaemus subannulatus ( Gray, 1842)
( Figs. 12–25 View FIGURE 12 View FIGURE 13 View FIGURE 14 View FIGURE 15 View FIGURE 16 View FIGURE 17 View FIGURE 18 View FIGURE 19 View FIGURE 20 View FIGURE 21 View FIGURE 24 View FIGURE 25 )
Diagnosis. — A species of the genus Tropidolaemus , characterized by (1) the internasals separated by 2 (rarely 1) scales, never in contact; (2) a background colour in the shades of green, namely green, blue or green and blue ground in females, green (blue in some Negros populations) in males and juveniles; (3) crossbands around the body blue and white, red and white, blue, blue and red or white in adult females, white parts can be light blue in life, white spots or white and red spots in males and juveniles; (4) a variable postocular stripe in adult females and a white and red one in juveniles and males; (5) in adult females the belly is uniform or blotched with blue or red, never banded, in males and juveniles the belly is uniform or with red dots; (6) 128– 148 VEN in males and 127–147 in females, SC: 45–53 in males and 40–54 in females; (7) 21–23 MSR in males and 21–29 in females, keeling variable in both sexes; (9) 4–7 scales on the snout at males and 5 – 8 in females; (10) 3 rd SL nearly always (87 / 90 occurrences) separated from the subocular by 1 scale or 2 scales, without sexual dimorphism; (10) tail long in males, with a ratio TaL/TL between 0.146 and 0.182, moderate to long in females, 0.139 –0.183; (11) occipital scales distinctly keeled in males.
Description and variation. —The maximal TL among our sample is 963 mm (SVL 820 mm, TaL 143 mm) for a female from “Siau, Sangi Islands”, now Sangihe Islands (BMNH not registered). Our largest male is 463 mm long (SVL 384 mm, TaL 79 mm; FMNH 222869 from Borneo).
We refrain from giving a more detailed description here, as the variation among this species or complex of species will be discussed in the next and forthcoming paper of the series. A splitting into several taxa seems to be likely.
Range. — Brunei. See Das (2007). – Federation of Malaysia. States of Sabah and Sarawak on Borneo ( Stuebing & Inger, 1999; Malkmus et al., 2002). – Indonesia. Known from the islands of Belitung, Borneo (Kalimantan), Buton, Kalimantan, Sangihe Archipelago, and Sulawesi. – Philippines. Recorded from the islands of Balabac, Basilan, Bohol, Dinagat, Jolo, Leyte, Luzon, Mindanao, Negros, Palawan, Panay, Samar, Sibutu, and Tumindao ( Alcala, 1986).
FIGURE 22.
Tropidolaemus subannu-
latus. Living adult
female from Negros
Island, Philippines.
Photograph by Mario
Lutz.
FIGURE 23.
Tropidolaemus
subannulatus. Living
adult female from
Palawan Island,
Philippines. Photograph
by Mario Lutz.
Material (64 specimens).— Belitung Island. ZMA 17709, “Billiton”. — Borneo. Federation of Malaysia . FMNH 128265 View Materials , FMNH 129467 View Materials , Pengkalang Lobang, Fourth Division, State of Sarawak . - FMNH 128433 View Materials , Tangap, Fourth Division, State of Sarawak ; FMNH 131848 View Materials , FMNH 131860 View Materials , FMNH 131865 View Materials , Niah, Fourth Division, State of Sarawak ; FMNH 131852 View Materials , Lower Niah, Fourth Division, State of Sarawak ; FMNH 131857 View Materials , Sungei Subis, Fourth Division, State of Sarawak ; FMNH 131861 View Materials , Niah, Sakaloh, Fourth Division, State of Sarawak ; FMNH 131862 View Materials , Niah, Pengkalan Lobang, Fourth Division, State of Sarawak ; FMNH 138663-64 View Materials , Sungei Mengiong, Nanga Tekalit Camp, Third Division, Kapit District, State of Sarawak ; FMNH 148833 View Materials , Labang Camp on Sungei Seran, Bintulu District, Fourth Division , State of Sarawak ; FMNH 222869 View Materials , Nanga Tekalit, Kapit District, Seventh Division, State of Sarawak . - MNHN 1880.0432 View Materials , MNHN 1991.3167 View Materials - 68 View Materials , Sandakan, State of Sabah ; MNHN 1906.0014 View Materials , No Sang, State of Sabah . - ZMH R 07438, “Mandalan River, Borneo”, now Sungei Medalam, Limbang (Fifth) Division (04°12'N, 114°57'E), State of Sarawak. - Indonesia GoogleMaps . MNHN 1891.0082 View Materials , “Sebroeang”, now Nangah Sebroeang, near Semitau, Province of Kalimantan Barat ; MNHN 2006.0419 View Materials , Pontianak, Province of Kalimantan Barat. - No locality . MNHN 1880.0435 View Materials , MNHN 1957.0817 View Materials , “ Borneo ” ; MNHN 1998.0488 View Materials , MNHN 2002.0447 View Materials - 49 View Materials , MNHN 2006.0420 View Materials , “Borneo” (from the pet trade). — Sulawesi : BMNH 1980.1718 , Ranu River, Morowali Nature Reserve ; BMNH 1980.0936 , Torro, Kulawi, Kabupaten Donggala, Central Sulawesi ; BMNH 49.3 .2.38, “ Celebes ” ; BMNH 96.12 .9.79, “Bone Valley, N. Celebes ”, now in Province of North Sulawesi . - MNHN 1999.9032 View Materials , MNHN 2002.0408 View Materials , “Sulawesi” (from the pet trade) ; MNHN 2002.0446 View Materials , an island off South Sulawesi (from the pet trade) . - ZMA 17711, “Posso”, now Poso, Province of Central Sulawesi. — Philippines. Luzon Is. 95.1.11.10- 12, “ Albay, S. E. Luzon ”, now Albay Province , Luzon Is.; MNHN 1880.0232 View Materials , “ Province Albay ( Luzon Philippines ) , now Albay Province, Luzon Is.; MNHN 1876.0032 View Materials , “Dans la Laguna (Luzon)”. - Mindanao Is . BMNH 77.10.9.71-72, “ Butuan, N. Mindanao ”, now Butuan City , Province of Agusan del Norte ; CAS 8679 View Materials , “San Ramon, Zamboanga Prov. ”, now in Province of Zamboanga del Sur ; FMNH 15046 View Materials , “ Agusan River, Mindanao Is” ; FMNH 22580 View Materials , Bunawan, Agusan del Sur Province ; FMNH 53564-67 View Materials , Tagum, Madaum , Davao Province , near sea level. - Negros Is. CAS-SUR 18792-93, Mabaja River area just West of Mayaposi Hill (Alt: 1800'), Negros Oriental Province. - Palawan Is. BMNH 79.11 .16.12 , CAS 15816, Puerta Princesa ; BMNH 94.6.30.64-65, “ Palawan ” ; CAS 28531, about 1 1/ 2 km. W.S.W. of Iwahig, Site 488 (300 ft), Palawan Prov. - Sulu Archipelago. MNHN 1880.0383 View Materials , “Archipel de Sulu entre Bornéo et Mindanao ”. - No locality . BMNH 1946.1.19.32-33, MNHN 1884.0046 View Materials , MNHN 1884.0181 View Materials - 0182 View Materials “Philippines”. — Unknown origin . MNHN 1878.0365 View Materials - 66 View Materials , both labelled as from “ Mansinam, Nouvelle Guinée ” .
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