BUCCINIDAE INCERTAE SEDIS

Kantor, Yuri I., Fedosov, Alexander E., Kosyan, Alisa R., Puillandre, Nicolas, Sorokin, Pavel A., Kano, Yasunori, Clark, Roger & Bouchet, Philippe, 2022, Molecular phylogeny and revised classification of the Buccinoidea (Neogastropoda), Zoological Journal of the Linnean Society 194, pp. 789-857 : 824-825

BUCCINIDAE INCERTAE SEDIS

As circumscribed herein, the family Buccinidae includes mostly boreal and, to a lesser extent, temperate to subtropical taxa of Northern Hemisphere Buccinoidea . Notwithstanding, there remain numerous, sometimes common, boreal genera that the lack of suitable material prevented us from including in the molecular analysis. Among them are Helicofusus Dall, 1916 , Latisipho Dall, 1916 and Troschelia Mörch, 1876 . Their taxonomic position remain uncertain, since shell and radula characters that were the basis for earlier placement appear to be weakly informative due to their high homoplasy.

Besides, representatives of two nominal subfamilies described within Buccinidae are not present in our analysis and, therefore, their status remains unconfirmed.

SUBFAMILY LIOMESINAE P.FISCHER, 1884

( FIG. 12Q)

= Buccinopsidae G.O. Sars, 1878 [invalid: type genus (Buccinopsis Jeffreys, 1867) a junior homonym]

Ty p e g e n u s: L i o m e s u s S t i m p s o n, 1 8 6 5 (T S † B u c c i n u m d a l e i J. d e C. S o w e r b y, 1 8 2 5; O D). Synonym: Buccinopsis Jeffreys, 1867 (TS † Buccinum dalei ; M).

Remarks: The radula of Liomesus ovum (Turton, 1825) , was illustrated by G.O. Sars (1878: tab. X, fig. 15) as Buccinopsis eburnea (M.Sars, 1851) . It is characterized by acuspate subrectangular central tooth and unicuspidate lateral ones. This morphology is different from other buccinoidean radulae, and has led Habe & Sato (1973) to suggest close relationships to Beringius , which was included by them, together with Liomesus , in the subfamily Liomesusinae . Bouchet & Warén (1985: fig. 461) provided a line drawing of a very different radula of Liomesus ovum , from an egg capsule, with muticuspidate central and lateral teeth. Because of this ambiguity, and the lack of molecular and anatomical data, we cannot propose any plausible hypothesis on the position of Liomesus .

SUBFAMILY TRUNCARIINAE COSSMANN, 1901

( FIG. 12P)

Type genus: Truncaria A.Adams & Reeve, 1850 (TS Buccinum filosum A.Adams & Reeve, 1850 ; M).

Remarks: The anatomy and radula of the type (and only included) genus remains unstudied ( Olsson, 1971) and molecular data are lacking as well. The peculiar shell shape ( Fig. 12P) does not provide any clue on the possible relationships of Truncaria , and even its position within Buccinidae remains to be confirmed.