Perkins, Philip D., 2009, Revisions of the genera Parhydraena Orchymont, Protozantaena Perkins, Decarthrocerus Orchymont, and Parhydraenopsis nomen novum, aquatic and humicolous beetles from Africa and Madagascar, and comparative morphology of the tribe Parhydraenini (Coleoptera: Hydraenidae)

Perkins, Philip D., 2009, Revisions of the genera Parhydraena Orchymont, Protozantaena Perkins, Decarthrocerus Orchymont, and Parhydraenopsis nomen novum, aquatic and humicolous beetles from Africa and Madagascar, and comparative morphology of the tribe Parhydraenini (Coleoptera: Hydraenidae), Zootaxa 2038, pp. 1-119

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Parhydraenini key    11
Parhydraena key    12-13
Protozantaena key    13
Parhydraenopsis key    13
Parhydraena     14
Decarthrocerus key    14
Parhydraena brevipalpis     15-16
Parhydraena divisa   new species  16-17
Parhydraena maureenae   new species  17-19
Parhydraena maculicollis   new species  19-20
Parhydraena parva   new species  20-21
Parhydraena seriata     21-23
Parhydraena ancylis   new species  23-25
Parhydraena asperita   new species  25-26
Parhydraena lancicula     26-28
Parhydraena namaqua   new species  28-29
Parhydraena ora   new species  29-30
Parhydraena semicostata   new species  30-31
Parhydraena sebastiani   new species  31-32
Parhydraena brunovacca   new species  33-35
Parhydraena mpumalanga   new species  35-36
Parhydraena brahma   new species  36-39
Parhydraena toro   new species  39-40
Protozantaena     41
Protozantaena labrata     41-42
Protozantaena malagasica   new species  43-44
Protozantaena palpalis   new species  44-45
Protozantaena ankaratra   new species  45-47
Protozantaena grebennikovi   new species  47-48
Parhydraenopsis     48-49
Parhydraenopsis cooperi     49-50
Parhydraenopsis alta   new species  50-51
Parhydraenopsis simiensis   new species  51-52
Decarthrocerus     52-53
Decarthrocerus jeanneli     53-54
Decarthrocerus bambusicus   new species  54-55
Decarthrocerus mbizi   new species  56
Decarthrocerus mahalicus   new species  57-58